Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk).
Latest update:
October 17th, 2024.
 
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Genus
Holcosus
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Northwestern Ameivas
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Nordwestliche Ameiven
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Nordvestlige Ameivaer
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1862 |  |
Holcosus Cope (type species: Ameiva septemlineata Dumeril 1851) |
Contents:
19 species, of which 9 (47.4%) are endemic.
Endemism: 0%  100%
Remarks:
Previously regarded as a synonym of
Ameiva (e.g., Peters & Donoso-Barros 1970).
Revalidated by Harvey, Ugueto & Gutberlet (2012).
This generic taxonomic arrangement was considered unwarranted by Giugliano, Nogueira, Valdujo, Collevatti & Colli (2013), but the validity of Holcosus was
supported by Tucker, Colli, Giugliano, Hedges, Hendry, Lemmon, Lemmon, Sites & Pyron (2016).
Introduced to:
Nicaragua
(Islas del Maiz:
Isla Pequeña del Maiz).
Distribution:
Mexico, Central America, NW. South America.
Reported from:
Belize,
Colombia
(incl.
Gorgona Island),
Costa Rica,
Ecuador,
El Salvador,
Guatemala,
Honduras,
Mexico
(Campeche,
Chiapas,
Colima,
Guerrero,
Hidalgo,
Jalisco,
Michoacan,
Morelos,
Nayarit,
Oaxaca,
Puebla,
Queretaro,
Quintana Roo,
San Luis Potosi,
Tabasco,
Tamaulipas,
Veracruz,
Yucatan),
Nicaragua
(incl.
Islas del Maiz:
Isla Grande del Maiz),
Panama.
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Holcosus amphigrammus
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Eastern Mexican Ameiva
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Ostmexicanische Ameive
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Østmexicansk Ameiva
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1945 | |
Ameiva undulata amphigramma Smith & Laufe |
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Holcosus undulatus amphigrammus Harvey, Ugueto & Gutberlet 2012 |
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Holcosus amphigrammus Meza-Lázaro & Nieto-Montes de Oca 2015 |
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1946 | |
Ameiva undulata podarga Smith & Laufe (Meza-Lázaro & Nieto-Montes de Oca 2015) |
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Holcosus undulatus podargus Harvey, Ugueto & Gutberlet 2012 |
Remarks:
Previously regarded as a subspecies of undulatus.
Raised to species status by Meza-Lázaro & Nieto-Montes de Oca (2015).
Current presence in Nuevo Leon needs confirmation, since the species has not been found in the state since 1934 (Lemos-Espinal, Wylie & Smith 2017).
Distribution:
Mexico
(Hidalgo,
Oaxaca,
Puebla,
Queretaro,
San Luis Potosi,
Tabasco,
Tamaulipas,
Veracruz).
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Holcosus anomalus
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Cauca Ameiva, (Echternacht's Ameiva)
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Cauca-Ameive
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Cauca-ameiva
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1977 | |
Ameiva anomala Echternacht |
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Holcosus anomalus Harvey, Ugueto & Gutberlet 2012 |
Distribution:
Colombia.
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Holcosus bridgesii
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Colombian Ameiva
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Kolumbianische Ameive
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Colombiansk Ameiva
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1869 | |
Holcosus bridgesii Cope |
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Ameiva bridgesii Barbour & Noble 1915 |
Distribution:
Colombia
(incl.
Gorgona Island),
Ecuador.
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Holcosus bridgesii
© Henrik Bringsøe
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Holcosus chaitzami
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Alta Verapaz Ameiva, (Chaitzam's Ameiva)
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Alta-Verapaz-Ameive
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Alta Verapaz-ameiva
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1924 | |
Ameiva chaitzami Stuart |
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Holcosus chaitzami Harvey, Ugueto & Gutberlet 2012 |
Remarks:
Previously included thomasi.
Distribution:
Guatemala,
Mexico
(Chiapas).
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Holcosus festivus
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Central American Ameiva, Central American Whiptail, Middle American Ameiva, Tiger Ameiva, Festive Ameiva
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Zentralamerikanische Ameive
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Centralamerikansk Ameiva
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1856 | |
Cnemidophorus festivus Lichtenstein |
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Ameiva festivus Bocourt 1874 |
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Holcosus festivus Harvey, Ugueto & Gutberlet 2012 |
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1862 | |
Ameiva eutropia Cope (Peters & Donoso-Barros 1970; Echternacht 1974) |
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1873 | |
Ameiva edwardsii Bocourt (Peters & Donoso-Barros 1970) |
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Ameiva festiva edwardsii Stuart 1943 |
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Holcosus festivus edwardsii Harvey, Ugueto & Gutberlet 2012 |
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1956 | |
Ameiva festiva occidentalis Taylor (Peters & Donoso-Barros 1970) |
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Holcosus festivus occidentalis Harvey, Ugueto & Gutberlet 2012 |
Other common names:
edwardsii: Edwards' Middle American Ameiva
Distribution:
Belize,
Colombia,
Costa Rica,
Guatemala,
Honduras,
Mexico
(Chiapas,
Tabasco),
Nicaragua,
Panama.
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Holcosus festivus
© Henrik Bringsøe
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Holcosus gaigeae
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Yucatan Ameiva
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Yucatan-Ameive
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Yucatan-ameiva
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1946 | |
Ameiva undulata gaigeae Smith & Laufe |
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Holcosus undulatus gaigeae Harvey, Ugueto & Gutberlet 2012 |
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Holcosus gaigeae Meza-Lázaro & Nieto-Montes de Oca 2015 |
Remarks:
Previously regarded as a subspecies of undulatus.
Raised to species status by Meza-Lázaro & Nieto-Montes de Oca (2015).
Not mentioned for Yucatan by González-Sánchez, Johnson, García-Padilla, Mata-Silva, DeSantis & Wilson (2017), although reported in several works
(e.g., Smith & Taylor 1950; Duellman 1965; Dundee, White & Rico-Gray 1986; Meza-Lázaro & Nieto-Montes de Oca 2015).
Distribution:
Belize,
Guatemala,
Mexico
(Campeche,
Quintana Roo,
Yucatan).
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Holcosus hartwegi
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Northern Chiapan Ameiva
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Nördlicher Chiapas-Ameive
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Nordlig Chiapas-ameiva
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1946 | |
Ameiva undulata hartwegi Smith |
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Holcosus undulatus hartwegi Harvey, Ugueto & Gutberlet 2012 |
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Holcosus hartwegi Meza-Lázaro & Nieto-Montes de Oca 2015 |
Remarks:
Previously regarded as a subspecies of undulatus.
Raised to species status by Meza-Lázaro & Nieto-Montes de Oca (2015).
Distribution:
Guatemala,
Honduras,
Mexico
(Chiapas,
Campeche,
Quintana Roo,
Yucatan).
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Holcosus leptophrys
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Delicate Ameiva
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Costa-Rica-Ameive
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Costa Rica-ameiva
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1893 | |
Amiva leptophrys Cope |
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Holcosus leptophrys Harvey, Ugueto & Gutberlet 2012 |
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1915 | |
Ameiva ruthveni Barbour & Noble (Peters & Donoso-Barros 1970) |
Distribution:
Colombia,
Costa Rica,
Panama.
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Holcosus miadis
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Islas del Maiz Ameiva
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Islas-del-Maiz-Ameive
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Islas del Maiz-ameiva
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1929 | |
Ameiva festiva miadis Barbour & Loveridge |
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Ameiva undulata miadis Peters & Donoso-Barros 1970 |
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Holcosus undulatus miadis Harvey, Ugueto & Gutberlet 2012 |
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Holcosus miadis Meza-Lázaro & Nieto-Montes de Oca 2015 |
Remarks:
Previously regarded as a subspecies of undulatus.
Raised to species status by Meza-Lázaro & Nieto-Montes de Oca (2015).
Introduced to:
Nicaragua
(Islas del Maiz:
Isla Pequeña del Maiz).
Distribution:
Nicaragua
(Islas del Maiz:
Isla Grande del Maiz).
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Holcosus niceforoi
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Sasaima Ameiva
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Sasaima-Ameive
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Sasaima-ameiva
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1943 | |
Ameiva festiva niceforoi Dunn |
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Ameiva niceforoi Echternacht 1970 |
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Holcosus niceforoi Harvey, Ugueto & Gutberlet 2012 |
Distribution:
Colombia.
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Holcosus orcesi
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Azuay Ameiva
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Azuay-Ameive
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Azuay-ameiva
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1964 | |
Ameiva orcesi Peters |
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Holcosus orcesi Harvey, Ugueto & Gutberlet 2012 |
Distribution:
Ecuador.
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Holcosus parvus
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Guatemalan Rainbow Ameiva
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Guatemalischer Regenbogen-Ameive
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Guatemalansk Regnbue-ameiva
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1915 | |
Ameiva undulata parva Barbour & Noble |
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Holcosus undulatus parvus Harvey, Ugueto & Gutberlet 2012 |
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Holcosus parvus Meza-Lázaro & Nieto-Montes de Oca 2015 |
Remarks:
Previously regarded as a subspecies of undulatus.
Raised to species status by Meza-Lázaro & Nieto-Montes de Oca (2015).
Distribution:
Costa Rica,
El Salvador,
Guatemala,
Mexico
(Chiapas,
Oaxaca).
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Holcosus pulcher
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Nicaraguan Rainbow Ameiva
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Nicaraguanischer Regenbogen-Ameive
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Nicaraguansk Regnbue-ameiva
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1861 | |
Ameiva pulchra Hallowell |
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Ameiva undulata pulchra Stuart 1942 |
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Holcosus undulatus pulcher Harvey, Ugueto & Gutberlet 2012 |
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Holcosus pulcher Meza-Lázaro & Nieto-Montes de Oca 2015 |
Remarks:
Status uncertain, but tentatively regarded as a valid species herein, following Meza-Lázaro & Nieto-Montes de Oca (2015), who concluded that pulcher is unlikely to be conspecific with undulatus.
Distribution:
Nicaragua.
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Holcosus quadrilineatus
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Four-lined Ameiva
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Vierstreifen-Ameive
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Firstribet Ameiva
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1861 | |
Cnemidophorus quadrilineatus Hallowell |
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Ameiva quadrilineata Cope 1862 |
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Holcosus quadrilineatus Harvey, Ugueto & Gutberlet 2012 |
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1876 | |
Amiva gabbiana Cope (Peters & Donoso-Barros 1970) |
Distribution:
Costa Rica,
Nicaragua,
Panama.
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Holcosus septemlineatus
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Seven-lined Ameiva
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Siebenstreifige Ameive
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Syvstribet Ameiva
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1851 | |
Ameiva septemlineata Dumeril |
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Holcosus septemlineatus Harvey, Ugueto & Gutberlet 2012 |
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1859 | |
Ameiva sex-scutata Günther (Peters & Donoso-Barros 1970) |
Distribution:
Colombia,
Ecuador.
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Holcosus sinister
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Western Mexican Ameiva
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Westmexicanische Ameive
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Vestmexicansk Ameiva
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1946 | |
Ameiva undulata sinistra Smith & Laufe |
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Holcosus undulatus sinistrus Harvey, Ugueto & Gutberlet 2012 |
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Holcosus sinister Meza-Lázaro & Nieto-Montes de Oca 2015 |
Remarks:
Previously regarded as a subspecies of undulatus.
Raised to species status by Meza-Lázaro & Nieto-Montes de Oca (2015).
Includes older, northwestern records of dextra (see undulatus), but these records may represent a distinct species (Meza-Lázaro & Nieto-Montes de Oca 2015).
Records from Morelos (e.g., Smith & Laufe 1946; Smith & Taylor 1950; Davis & Smith 1953), are erroneous (Echternacht 1971; Castro-Franco & Bustos-Zagal 1994), however,
Lemos-Espinal & Smith (2020) still listed the species for the state.
Distribution:
Mexico
(Colima,
Guerrero,
Jalisco,
Michoacan,
Nayarit,
Puebla).
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Holcosus stuarti
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Palenque Ameiva
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Palenque-Ameive
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Palenque-ameiva
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1940 | |
Ameiva undulata stuarti Smith |
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Holcosus undulatus stuarti Harvey, Ugueto & Gutberlet 2012 |
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Holcosus stuarti Meza-Lázaro & Nieto-Montes de Oca 2015 |
Remarks:
Previously regarded as a subspecies of undulatus.
Raised to species status by Meza-Lázaro & Nieto-Montes de Oca (2015).
Distribution:
Mexico
(Campeche,
Chiapas,
Tabasco).
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Holcosus thomasi
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Rio Grijalva Ameiva
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Rio-Grijalva-Ameive
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Rio Grijalva-ameiva
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1946 | |
Ameiva undulata thomasi Smith & Laufe |
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Holcosus chaitzami thomasi Harvey, Ugueto & Gutberlet 2012 |
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Holcosus thomasi Meza-Lázaro & Nieto-Montes de Oca 2015 |
Remarks:
Previously regarded as a subspecies of undulata (e.g., Peters & Donoso-Barros 1970) or a synonym of chaitzami (e.g., Echternacht 1970). Raised to species status by Meza-Lázaro & Nieto-Montes de Oca (2015).
Distribution:
Guatemala,
Mexico
(Chiapas).
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Holcosus undulatus
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Mexican Rainbow Ameiva, Metallic Ameiva
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Mexicanische Regenbogen-Ameive
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Mexicansk Regnbue-ameiva
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1834 | |
Cnemidophorus undulatus Wiegmann |
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Ameiva undulata Gray 1845 |
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Holcosus undulatus Harvey, Ugueto & Gutberlet 2012 |
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1946 | |
Ameiva undulata dextra Smith & Laufe (Meza-Lázaro & Nieto-Montes de Oca 2015, see remarks) |
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Holcosus undulatus dextrus Harvey, Ugueto & Gutberlet 2012 |
Remarks:
Previously included amphigrammus, gaigeae, hartwegi, miadis, parvus, pulcher, sinister, stuarti, and thomasi,
which were elevated to species status by Meza-Lázaro & Nieto-Montes de Oca (2015).
This arrangement was questioned by McCranie, Sunyer & Martínez-Fonseca (2019), who considered it poorly supported.
Northwestern records of dextra refer to sinister, but may represent a distinct species (Meza-Lázaro & Nieto-Montes de Oca 2015).
Distribution:
Mexico
(Guerrero,
Oaxaca).
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