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Taxonomy of the family Colubridae
Bibliography of the genus Ptyas
Biodiversity of the family Colubridae








Genus
Ptyas

Giant Rat Snakes (Asian Rat Snakes) & Oriental Green Snakes

Riesen-Rattennattern* (Rattenschlangen) & Orientalische Grünnattern*

Kæmperottesnoge* (Orientalske Rottesnoge) & Orientalske Smaragdsnoge*

1843 Ptyas Fitzinger (type species: Coluber blumenbachii Merrem 1820)
1861 Eurypholis Hallowell [not Eurypholis Pictet 1850 (Pisces)] (type species: Eurypholis semicarinatus Hallowell 1861; syn. Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016)
1861 Zaocys Cope (type species: Coluber dhumnades Cantor 1842; syn. Lazell, Keirans & Samuelson 1991; Lazell 1998)
1864 Zapyrus Günther (type species: Coryphodon fuscus Günther 1864; syn. Wallach, Williams & Boundy 2014)
1865 Megablabes Günther (type species: Megablabes olivaceus = Herpetodryas dipsas Schlegel 1837; syn. Wallach, Williams & Boundy 2014)
1878 Hemidryas Peters & Doria (type species: Herpetodryas dipsas Schlegel 1837; syn. Wallach, Williams & Boundy 2014)
1888 Cyclophiops Boulenger (type species: Cyclophiops doriae Boulenger 1888; syn. Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016)
1895 Entechinus Cope (type species: Cyclophis major Günther 1858; syn. [of Cyclophiops] Wallach, Williams & Boundy 2014)
Contents: 13 species, of which 4 (30.8%) are endemic.
Other common names:
Ptyas (pre-2016 use): Giant Rat Snakes (Asian Rat Snakes)
Cyclophiops: Oriental Green Snakes
Remarks: Status of Zaocys is uncertain (see discussion in David & Das 2004). Most recent publications have synonymized Zaocys with Ptyas (e.g., David & Vogel 1996; Lazell 1998; Whitaker & Captain 2004; David, Cox, Pauwels, Chanhome & Thirakhupt 2004; Das 2010; Wallach, Williams & Boundy 2014), but Zaocys may prove valid (David, Cox, Pauwels, Chanhome & Thirakhupt 2004), and some authors still treat it as such (e.g., Zaher, Grazziotin, Cadle, Murphy, Moura-Leite & Bonatto 2009). Lazell, Keirans & Samuelson (1991) and Lazell (1998) included both Ptyas and Zaocys in a greatly expanded Coluber, however, most subsequent works have retained Ptyas as a valid genus (e.g., Pinou & Dowling 2000; Wallach, Williams & Boundy 2014). David & Das (2004) noted that the gender of Ptyas is feminine.
Distribution: Asia, Malay Archipelago.
Reported from: Afghanistan, Bangladesh, Bhutan, Brunei, Cambodia, China (Anhui, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hebei, Henan, Hong Kong [incl. Shek Kwu Chau Island], Hubei, Hunan, Jiangsu, Jiangxi, Macau, Shaanxi, Shanxi, Sichuan, Tibet, Yunnan, Zhejiang), India (Andaman & Nicobar Islands [Andaman Islands], Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Chhattisgarh, Delhi, Goa, Gujarat, Haryana, Himachal Pradesh, Jammu & Kashmir, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Punjab, Rajasthan, Sikkim, Tamil Nadu, Tripura, Uttar Pradesh, Uttarakhand, West Bengal), Indonesia (Bali, Bangka, Belitung, Java, Kalimantan, Madura, Natuna Islands, Nias, Penida, Sulawesi, Sumatra), Japan (Ryukyu Islands [Okinawa Islands, Sakishima Islands (incl. Yaeyama Islands), Satsunan Islands (incl. Tokara Islands)]), Laos, Malaysia (Sabah, Sarawak, West Malaysia [incl. Langkawi Archipelago, Seribuat Archipelago (incl. Tioman)]), Myanmar, Nepal, Pakistan, Philippines (Catanduanes, Leyte, Luzon, Negros, Palawan, Panay, Polillo), Singapore, Sri Lanka, Taiwan, Thailand, Turkmenistan, Vietnam (incl. Cat Ba Island).

Ptyas carinata

Malayan Keeled Rat Snake, Black Giant Rat Snake, Carinated Rat Snake

Gekielte Rattennatter

Kølskællet Rottesnog*

1858 Coryphodon carinatus Günther
Zaocys carinatus Günther 1864
Ptyas carinatus Wall 1926
1891 Zaocys tenasserimensis Sclater (Smith 1943)
Ptyas tenasserimensis Wall 1923

Distribution: Brunei, Cambodia, China (Yunnan), Indonesia (Java, Kalimantan, Sumatra), Malaysia (Sabah, Sarawak, West Malaysia [incl. Langkawi Archipelago, Seribuat Archipelago (incl. Tioman)]), Myanmar, Philippines (Palawan), Singapore, Thailand, Vietnam.


Ptyas dhumnades

Black-striped Rat Snake, Big-eyed Rat Snake, Dhamin Rat Snake

Schwarzstrefige Rattennatter*

Sortstribet Rottesnog*

1842 Coluber dhumnades Cantor
Zaocys dhumnades Cope 1860
Coryphodon dhumnades Müller 1878
Ptyas dhumnades Welch 1988
1854 Ablabes vittatus Duméril, Bibron & Duméril (Zhao & Adler 1993)
1925 Zaocys oshimai Stejneger (Zhao & Adler 1993)
Zaocys dhumnades oshimai Pope 1935
Zaocys nigromarginatus oshimai Wang & Wang 1956
1928 Zaocys dhumnades montanus Pope (Zhao & Adler 1993)
Ptyas dhumnades montanus Welch 1988

Introduced to: China (Beijing).
Distribution: China (Anhui, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hebei, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Shanxi, Sichuan, Zhejiang), Laos, Taiwan, Vietnam.


Ptyas dipsas

Sulawesi Black Rat Snake

Indonesische Schwarze Rattennatter*

Indonesisk Sort Rottesnog*

1837 Herpetodryas dipsas Schlegel
Ahaetulla dipsas Peters & Doria 1878
Zamenis dipsas Boulenger 1893
Ptyas dipsas Haas 1950
Coluber dipsas Lazell, Keirans & Samuelson 1991

Remarks: Although reported from Halmahera, presence of dipsas in this island is considered doubtful (Koch 2012).
Distribution: Indonesia (Sulawesi).


Ptyas doriae

Burmese Green Snake*, Green Grass Snake, (Doria's Green Snake)

Burmesische Grünnatter*

Burmesisk Smaragdsnog*

1888 Cyclophiops doriae Boulenger
Ablabes doriae Boulenger 1890
Liopeltis doriae Wall 1924
Eurypholis doriae Pope 1935
Opheodrys doriae Smith 1943
Entechinus doriae Cundall 1981
Ptyas doriae Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016
1900 Ablabes hamptoni Boulenger (Meetei, Das, Campbell, Raha & Bag 2018)
Opheodrys hamptoni Smith 1943
Entechinus hamptoni Cundall 1981
Cyclophiops hamptoni Toriba 1989
Ptyas hamptoni Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016

Other common names:
hamptoni: (Hampton's Green Snake)
Remarks: Mahendra (1984) regarded hamptoni as a synonym of doriae. The synonymy was supported by Meetei, Das, Campbell, Raha & Bag (2018).
Distribution: China (Yunnan), India (Manipur), Myanmar.


Ptyas fusca

White-bellied Rat Snake, (Brown Rat Snake)

Weißbauch-Rattennatter*

Hvidbuget Rottesnog*

1858 Coryphodon fuscus Günther
Zaocys fuscus Günther 1864
Ptyas fuscus Wall 1923
2001 Ptyas zoazys Nutphand [nomen nudum] (David, Cox, Pauwels, Chanhome & Thirakhupt 2004)

Distribution: Brunei, Indonesia (Bangka, Belitung, Kalimantan, Natuna Islands, Nias, Sumatra), Malaysia (Sabah, Sarawak, West Malaysia [incl. Seribuat Archipelago (incl. Tioman)]), Singapore, Thailand.


Ptyas herminae

Sakishima Green Snake, Japanese Green Snake

Sakishima-Grünnatter*

Sakishima-smaragdsnog*

1895 Ablabes herminae Boettger
Liopeltis herminae Stejneger 1907
Opheodrys herminae Schmidt & Necker 1936
Eurypholis herminae Pope 1935
Entechinus herminae Cundall 1981
Cyclophiops herminae Toriba 1989
Ptyas herminae Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016

Distribution: Japan (Ryukyu Islands: Sakishima Islands: Yaeyama Islands).


Ptyas korros

Indochinese Rat Snake, (Chinese Rat Snake, Schlegel's Rat Snake, Javan Rat Snake)

Indomalaiische Rattennatter

Indokinesisk Rottesnog*

1837 Coluber korros Schlegel
Ptyas korros Cope 1860
Coryphodon korros Jan 1867
Zamenis korros Boulenger 1890
1837 Coluber cancellatus Oppel in Schlegel (Stejneger 1907)
1910 Liopeltis libertatis Barbour (Smith 1943)
Ablabes libertatis Brongersma in Damerman 1929
1930 Ptyas korros chinensis Mell [1929] (Zhao & Adler 1993)
1931 Ptyas korros indicus Mell [1929] (Zhao & Adler 1993)

Distribution: Bangladesh, Bhutan, Cambodia, China (Anhui, Fujian, Guangdong [incl. Nan Ao Island], Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Jiangxi, Macau, Yunnan, Zhejiang), India (Arunachal Pradesh, Assam, Meghalaya, Nagaland, West Bengal), Indonesia (Bali, Java, Kalimantan, Madura, Penida, Sumatra), Laos, Malaysia (Sarawak, West Malaysia), Myanmar, Singapore, Taiwan, Thailand, Vietnam (incl. Cat Ba Island).


Ptyas luzonensis

Smooth-scaled Mountain Rat Snake, Smooth-scaled Rat Snake, (Philippine Rat Snake)

Luzon-Rattennatter*

Luzon-rottesnog*

1873 Zaocys luzonensis Günther
Ptyas luzonensis Welch 1988

Distribution: Philippines (Catanduanes, Leyte, Luzon, Negros, Panay, Polillo).


Ptyas major

Greater Green Snake, Chinese Green Snake, Oriental Smooth Green Snake

Große Grünnatter*

Stor Smaragdsnog*

1858 Cyclophis major Günther
Ablabes major Boettger 1894
Entechinus major Cope 1895
Liopeltis major Stejneger 1907
Eurypholis major Pope 1935
Opheodrys major Schmidt & Necker 1936
Cyclophiops major Ota 1991
Ptyas major Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016
1860 Herpetodryas chloris Hallowell (Smith 1943)
1930 Coluber delacouri Smith (Smith 1943)
1931 Liopeltis major bicarinata Maki

Distribution: China (Anhui, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hong Kong, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang), Laos, Taiwan, Vietnam.


Ptyas mucosa

Common Oriental Rat Snake*, Common Indian Rat Snake, Common Rat Snake, Dhaman, Greater Rat Snake, Rope Snake, (Oriental Rat Snake)

Gebänderte Rattennatter

Almindelig Orientalsk Rottesnog*

1758 Coluber mucosus Linnaeus
Natrix mucosa Laurenti 1768
Ptyas mucosus Cope 1860
Zamenis mucosus Boulenger 1890
Zaocys mucosus Wall 1914
1820 Coluber blumenbachii Merrem (Smith 1943)
1839 Coluber dhumna Cantor (Smith 1943)
1860 Leptophis trifrenatus Hallowell (Smith 1943)
1867 Coryphodon blumenbachi carinata Jan (Stejneger 1907)
1918 Ptyas mucosus pallidus Dábreu (Iskandar & Colijn 2001)
1949 Ptyas mucosus nigriceps Terentjev & Chernov
1955 Ptyas mucosus maximus Deraniyagala

Other common names:
nigricens: Dark Brown Rat Snake
Remarks: Present in the past, but not confirmed recently from Singapore (Baker & Lim 2008).
Extinct in: Singapore.
Distribution: Afghanistan, Bangladesh, Cambodia, China (Anhui, Chongqing, Fujian, Guangdong [incl. Nan Ao Island], Guangxi, Guizhou, Hainan, Hong Kong [incl. Shek Kwu Chau Island], Hubei, Hunan, Jiangxi, Macau, Sichuan, Tibet, Yunnan, Zhejiang), India (Andaman & Nicobar Islands [Andaman Islands], Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Chhattisgarh, Delhi, Goa, Gujarat, Haryana, Himachal Pradesh, Jammu & Kashmir, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Mizoram, Odisha, Punjab, Rajasthan, Tamil Nadu, Tripura, Uttar Pradesh, Uttarakhand, West Bengal), Indonesia (Bali, Bangka, Java, Sumatra), Laos, Malaysia (West Malaysia), Myanmar, Nepal, Pakistan, Singapore, Sri Lanka, Taiwan, Thailand, Turkmenistan, Vietnam.


Ptyas multicincta

Many-banded Green Snake, Northern Chinese Green Snake

Vielbändige Grünnatter*

Mangebåndet Smaragdsnog*

1907 Ablabes multicinctus Roux
Eurypholis multicinctus Pope 1935
Liopeltis multicinctus Angel & Bourret 1933
Liopeltis major multicinctus Bourret 1936
Opheodrys multicinctus Schmidt & Necker 1936
Eurypholis multicinctus Pope 1936
Entechinus multicinctus Cundall 1981
Cyclophiops multicinctus Zhao & Adler 1993
Ptyas multicincta Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016
1920 Ablabes retrofasciatus Angel (Smith 1943)
1921 Zamenis moi Smith (Smith 1943)
1929 Ablabes multicinctus bicolor Angel (Smith 1943)
Liopeltis major bicolor Bourret 1936
1931 Liopeltis sinii Fan (Iskandar & Colijn 2001)

Distribution: China (Guangdong, Guangxi, Hainan, Yunnan), Laos, Thailand, Vietnam.


Ptyas nigromarginata

Oriental Green Rat Snake, Black-bordered Rat Snake

Grüne Gekielte Rattennatter

Sortrandet Rottesnog*

1854 Coluber nigromarginatus Blyth
Zaocys nigromarginatus Günther 1864
Ptyas nigromarginatus Wall 1923
Zaocys dhumnades nigromarginatus Bourret 1936

Remarks: Records from Taiwan refer to oshimai, regarded as a subspecies of nigromarginatus by Wang & Wang (1956), but as a synonym of dhumnades by Zhao & Adler (1993). Presence in Nepal needs confirmation (Kästle, Rai & Schleich 2013).
Distribution: Bangladesh, Bhutan, China (Chongqing, Gansu, Guizhou, Sichuan, Tibet, Yunnan), India (Arunachal Pradesh, Meghalaya, Nagaland, Sikkim, West Bengal), Myanmar, Thailand, Vietnam.


Ptyas semicarinata

Ryukyu Green Snake

Ryukyu-Grünnatter*

Ryukyu-smaragdsnog*

1861 Eurypholis semicarinatus Hallowell
Ablabes semicarinatus Boulenger 1894
Liopeltis semicarinata Stejneger 1907
Opheodrys semicarinatus Schmidt & Necker 1936
Entechinus semicarinatus Cundall 1981
Cyclophiops semicarinatus Ota, Shiroma & Hikida 1995
Ptyas semicarinata Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016
1868 Cyclophis nebulosus Günther (Stejneger 1907)

Distribution: Japan (Ryukyu Islands [Okinawa Islands, Satsunan Islands (Amami Islands, Tokara Islands)]).