Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk).
Latest update:
October 9th, 2024.
 
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Genus
Ptyas
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Giant Rat Snakes (Asian Rat Snakes) and Oriental Green Snakes
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Riesen-Rattennattern (Rattenschlangen) und Orientalische Grünnattern
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Kæmperottesnoge (Orientalske Rottesnoge) og Orientalske Smaragdsnoge
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1843 |  |
Ptyas Fitzinger (type species: Coluber blumenbachii Merrem 1820) |
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1861 | |
Eurypholis Hallowell [not Eurypholis Pictet 1850 (Pisces)] (type species: Eurypholis semicarinatus Hallowell 1861; syn. Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016) |
|
1861 | |
Zaocys Cope (type species: Coluber dhumnades Cantor 1842; syn. Lazell, Keirans & Samuelson 1991; Lazell 1998) |
|
1864 | |
Zapyrus Günther (type species: Coryphodon fuscus Günther 1864; syn. Wallach, Williams & Boundy 2014) |
|
1865 | |
Megablabes Günther (type species: Megablabes olivaceus = Herpetodryas dipsas Schlegel 1837; syn. Wallach, Williams & Boundy 2014) |
|
1878 | |
Hemidryas Peters & Doria (type species: Herpetodryas dipsas Schlegel 1837; syn. Wallach, Williams & Boundy 2014) |
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1888 | |
Cyclophiops Boulenger (type species: Cyclophiops doriae Boulenger 1888; syn. Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016) |
|
1895 | |
Entechinus Cope (type species: Cyclophis major Günther 1858; syn. [of Cyclophiops] Wallach, Williams & Boundy 2014) |
Contents:
14 species, of which 5 (35.7%) are endemic.
Endemism: 0%  100%
Other common names:
Ptyas (pre-2016 use): Giant Rat Snakes (Asian Rat Snakes)
Cyclophiops: Oriental Green Snakes
Remarks:
Status of Zaocys is uncertain (see discussion in David & Das 2004).
Most recent publications have synonymized Zaocys with Ptyas
(e.g., David & Vogel 1996; Lazell 1998; Whitaker & Captain 2004; David, Cox, Pauwels, Chanhome & Thirakhupt 2004; Das 2010; Wallach, Williams & Boundy 2014),
but Zaocys may prove valid (David, Cox, Pauwels, Chanhome & Thirakhupt 2004), and some authors still treat it as such (e.g., Zaher, Grazziotin, Cadle, Murphy, Moura-Leite & Bonatto 2009).
Lazell, Keirans & Samuelson (1991) and Lazell (1998) included both Ptyas and Zaocys in a greatly expanded
Coluber, however, most subsequent works have retained Ptyas as a valid genus
(e.g., Pinou & Dowling 2000; Wallach, Williams & Boundy 2014).
David & Das (2004) noted that the grammatical gender of Ptyas is feminine.
Distribution:
Asia, Malay Archipelago.
Reported from:
Afghanistan,
Bangladesh,
Bhutan,
Brunei,
Cambodia,
China
(Anhui,
Chongqing,
Fujian,
Gansu,
Guangdong,
Guangxi,
Guizhou,
Hainan,
Hebei,
Henan,
Hong Kong
[incl.
Shek Kwu Chau Island],
Hubei,
Hunan,
Jiangsu,
Jiangxi,
Macau,
Shaanxi,
Shanxi,
Sichuan,
Tibet,
Yunnan,
Zhejiang),
India
(Andaman & Nicobar Islands
[Andaman Islands],
Andhra Pradesh,
Arunachal Pradesh,
Assam,
Bihar,
Chhattisgarh,
Delhi,
Goa,
Gujarat,
Haryana,
Himachal Pradesh,
Jammu & Kashmir,
Jharkhand,
Karnataka,
Kerala,
Madhya Pradesh,
Maharashtra,
Manipur,
Meghalaya,
Mizoram,
Nagaland,
Odisha,
Punjab,
Rajasthan,
Sikkim,
Tamil Nadu,
Telangana,
Tripura,
Uttar Pradesh,
Uttarakhand,
West Bengal),
Indonesia
(Bali,
Bangka,
Belitung,
Java,
Kalimantan,
Madura,
Natuna Islands,
Nias,
Penida,
Sulawesi,
Sumatra),
Japan
(Ryukyu Islands
[Okinawa Islands,
Sakishima Islands
(incl.
Yaeyama Islands),
Satsunan Islands
(incl.
Tokara Islands)]),
Laos,
Malaysia
(Sabah,
Sarawak,
West Malaysia
[incl.
Langkawi Archipelago,
Seribuat Archipelago
(incl.
Tioman)]),
Myanmar,
Nepal,
Pakistan,
Philippines
(Catanduanes,
Leyte,
Luzon,
Negros,
Palawan,
Panay,
Polillo),
Singapore,
Sri Lanka,
Taiwan,
Thailand,
Turkmenistan,
Vietnam
(incl.
Cat Ba Island).
| |
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Ptyas bachmaensis
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Bach Ma Rat Snake
|
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Bach-Ma-Rattennatter
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Bach Ma-rottesnog
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2024 | |
Ptyas bachmaensis Nguyen, Vo, Orlov, Phan, Le, Nguyen, Tran, Murphy & Che |
Remarks:
Previously regarded as a population of nigromarginata.
Distribution:
Vietnam.
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Ptyas carinata
|
Malayan Keeled Rat Snake, Black Giant Rat Snake, Carinated Rat Snake
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Gekielte Rattennatter
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Kølskællet Rottesnog
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1864 | |
Zaocys carinatus Günther [not Coryphodon carinatus Günther 1858; see Ptyas dhumnades] |
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| |
Ptyas carinatus Wall 1923 |
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1891 | |
Zaocys tenasserimensis Sclater (syn. Smith 1943) |
|
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Ptyas tenasserimensis Wall 1923 |
Distribution:
Brunei,
Cambodia,
China
(Yunnan),
Indonesia
(Java,
Kalimantan,
Sumatra),
Laos,
Malaysia
(Sabah,
Sarawak,
West Malaysia
[incl.
Langkawi Archipelago,
Seribuat Archipelago
(incl.
Tioman)]),
Myanmar,
Philippines
(Palawan),
Singapore,
Thailand,
Vietnam.
|
Ptyas carinata
© Rune Midtgaard
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Ptyas dhumnades
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Black-striped Rat Snake, Big-eyed Rat Snake, Dhamin Rat Snake
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Schwarzstrefige Rattennatter
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Sortstribet Rottesnog
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1842 | |
Coluber dhumnades Cantor |
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| |
Zaocys dhumnades Cope 1860 |
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Coryphodon dhumnades Müller 1878 |
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Ptyas dhumnades Welch 1988 |
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1854 | |
Ablabes vittatus Duméril, Bibron & Duméril (syn. Zhao & Adler 1993; Dubois & David 2020) |
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1858 | |
Coryphodon carinatus Günther [substitute name for Coluber dhumnades Cantor 1842] (syn. Dubois & David 2020) |
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1925 | |
Zaocys oshimai Stejneger (syn. Zhao & Adler 1993; Dubois & David 2020) |
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Zaocys dhumnades oshimai Pope 1935 |
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Zaocys nigromarginatus oshimai Wang & Wang 1956 |
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1928 | |
Zaocys dhumnades montanus Pope (syn. Zhao & Adler 1993; Dubois & David 2020) |
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Zaocys nigromarginatus montanus Mell 1929 |
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Ptyas dhumnades montanus Welch 1988 |
Introduced to:
China
(Beijing).
Distribution:
China
(Anhui,
Chongqing,
Fujian,
Gansu,
Guangdong,
Guangxi,
Guizhou,
Hebei,
Henan,
Hubei,
Hunan,
Jiangsu,
Jiangxi,
Shaanxi,
Shanxi,
Sichuan,
Zhejiang),
Laos,
Taiwan,
Vietnam.
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Ptyas dipsas
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Sulawesi Black Rat Snake
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Indonesische Schwarze Rattennatter
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Indonesisk Sort Rottesnog
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1837 | |
Herpetodryas dipsas Schlegel |
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Ahaetulla dipsas Peters & Doria 1878 |
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Zamenis dipsas Boulenger 1893 |
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Ptyas dipsas Haas 1950 |
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Coluber dipsas Lazell, Keirans & Samuelson 1991 |
Remarks:
Although reported from Halmahera, presence of dipsas in this island is considered doubtful (Koch 2012).
Distribution:
Indonesia
(Sulawesi).
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Ptyas doriae
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Burmese Green Snake, Green Grass Snake, (Doria's Green Snake)
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Burmesische Grünnatter
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Burmesisk Smaragdsnog
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1888 | |
Cyclophiops doriae Boulenger |
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Ablabes doriae Boulenger 1890 |
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Liopeltis doriae Wall 1924 |
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Eurypholis doriae Pope 1935 |
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Opheodrys doriae Smith 1943 |
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Entechinus doriae Cundall 1981 |
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Ptyas doriae Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016 |
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1900 | |
Ablabes hamptoni Boulenger (syn. Meetei, Das, Campbell, Raha & Bag 2018) |
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Opheodrys hamptoni Smith 1943 |
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Entechinus hamptoni Cundall 1981 |
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Cyclophiops hamptoni Toriba 1989 |
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| |
Ptyas hamptoni Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016 |
Other common names:
hamptoni: (Hampton's Green Snake)
Remarks:
Mahendra (1984) regarded hamptoni as a synonym of doriae.
The synonymy was supported by Meetei, Das, Campbell, Raha & Bag (2018).
Distribution:
China
(Yunnan),
India
(Manipur),
Myanmar.
|
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Ptyas fusca
|
White-bellied Rat Snake, (Brown Rat Snake)
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Weissbauch-Rattennatter
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Hvidbuget Rottesnog
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1858 | |
Coryphodon fuscus Günther |
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Zaocys fuscus Günther 1864 |
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Ptyas fuscus Wall 1923 |
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2001 | |
Ptyas zoazys Nutphand [nomen nudum] (syn. David, Cox, Pauwels, Chanhome & Thirakhupt 2004) |
Distribution:
Brunei,
Indonesia
(Bangka,
Belitung,
Kalimantan,
Natuna Islands,
Nias,
Sumatra),
Malaysia
(Sabah,
Sarawak,
West Malaysia
[incl.
Seribuat Archipelago
(incl.
Tioman)]),
Singapore,
Thailand.
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Ptyas herminae
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Sakishima Green Snake, Japanese Green Snake
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Sakishima-Grünnatter
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Sakishima-smaragdsnog
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1895 | |
Ablabes herminae Boettger |
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Liopeltis herminae Stejneger 1907 |
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Opheodrys herminae Schmidt & Necker 1936 |
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Eurypholis herminae Pope 1935 |
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Entechinus herminae Cundall 1981 |
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Cyclophiops herminae Toriba 1989 |
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Ptyas herminae Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016 |
Distribution:
Japan
(Ryukyu Islands:
Sakishima Islands:
Yaeyama Islands).
|
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Ptyas korros
|
Indochinese Rat Snake, (Chinese Rat Snake, Schlegel's Rat Snake, Javan Rat Snake)
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Indomalaiische Rattennatter
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Indokinesisk Rottesnog
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1837 | |
Coluber korros Schlegel |
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Ptyas korros Cope 1860 |
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Coryphodon korros Jan 1867 |
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Zamenis korros Boulenger 1890 |
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1837 | |
Coluber cancellatus Oppel in Schlegel (Stejneger 1907) |
|
1910 | |
Liopeltis libertatis Barbour (Smith 1943) |
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Ablabes libertatis Brongersma in Damerman 1929 |
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1930 | |
Ptyas korros chinensis Mell [1929] (Zhao & Adler 1993) |
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1931 | |
Ptyas korros indicus Mell [1929] (Zhao & Adler 1993) |
Distribution:
Bangladesh,
Bhutan,
Cambodia,
China
(Anhui,
Fujian,
Guangdong
[incl.
Nan Ao Island],
Guangxi,
Guizhou,
Hainan,
Hong Kong,
Hunan,
Jiangxi,
Macau,
Yunnan,
Zhejiang),
India
(Arunachal Pradesh,
Assam,
Meghalaya,
Mizoram,
Nagaland,
West Bengal),
Indonesia
(Bali,
Java,
Kalimantan,
Madura,
Penida,
Sumatra),
Laos,
Malaysia
(Sarawak,
West Malaysia),
Myanmar,
Singapore,
Taiwan,
Thailand,
Vietnam
(incl.
Cat Ba Island).
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Ptyas korros
© Henrik Bringsøe
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Ptyas luzonensis
|
Smooth-scaled Mountain Rat Snake, Smooth-scaled Rat Snake, (Philippine Rat Snake)
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Luzon-Rattennatter
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Luzon-rottesnog
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1873 | |
Zaocys luzonensis Günther |
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Ptyas luzonensis Welch 1988 |
Distribution:
Philippines
(Catanduanes,
Leyte,
Luzon,
Negros,
Panay,
Polillo).
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Ptyas major
|
Greater Green Snake, Chinese Green Snake, Oriental Smooth Green Snake
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Grosse Grünnatter
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Stor Smaragdsnog
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1858 | |
Cyclophis major Günther |
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Ablabes major Boettger 1894 |
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Entechinus major Cope 1895 |
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Liopeltis major Stejneger 1907 |
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Eurypholis major Pope 1935 |
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Opheodrys major Schmidt & Necker 1936 |
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Cyclophiops major Ota 1991 |
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Ptyas major Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016 |
|
1860 | |
Herpetodryas chloris Hallowell (Smith 1943) |
|
1930 | |
Coluber delacouri Smith (Smith 1943) |
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1931 | |
Liopeltis major bicarinata Maki |
Distribution:
China
(Anhui,
Chongqing,
Fujian,
Gansu,
Guangdong,
Guangxi,
Guizhou,
Hainan,
Henan,
Hong Kong,
Hubei,
Hunan,
Jiangsu,
Jiangxi,
Shaanxi,
Sichuan,
Yunnan,
Zhejiang),
Taiwan,
Vietnam.
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Ptyas mucosa
|
Common Oriental Rat Snake, Common Indian Rat Snake, Common Rat Snake, Dhaman, Greater Rat Snake, Rope Snake, (Oriental Rat Snake)
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Gebänderte Rattennatter
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Almindelig Orientalsk Rottesnog
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1758 | |
Coluber mucosus Linnaeus |
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Natrix mucosa Laurenti 1768 |
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Ptyas mucosus Cope 1860 |
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Zamenis mucosus Boulenger 1890 |
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Zaocys mucosus Wall 1914 |
|
1820 | |
Coluber blumenbachii Merrem (Smith 1943) |
|
1839 | |
Coluber dhumna Cantor (Smith 1943) |
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1860 | |
Leptophis trifrenatus Hallowell (Smith 1943) |
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1867 | |
Coryphodon blumenbachi carinata Jan (Stejneger 1907) |
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1918 | |
Ptyas mucosus pallidus Dábreu (Iskandar & Colijn 2001) |
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1949 | |
Ptyas mucosus nigriceps Terentjev & Chernov |
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1955 | |
Ptyas mucosus maximus Deraniyagala |
Other common names:
nigricens: Dark Brown Rat Snake
Remarks:
Present in the past, but not confirmed recently from Singapore (Baker & Lim 2008).
Extinct in:
Singapore.
Distribution:
Afghanistan,
Bangladesh,
Cambodia,
China
(Anhui,
Chongqing,
Fujian,
Guangdong
[incl.
Nan Ao Island],
Guangxi,
Guizhou,
Hainan,
Hong Kong
[incl.
Shek Kwu Chau Island],
Hubei,
Hunan,
Jiangxi,
Macau,
Sichuan,
Tibet,
Yunnan,
Zhejiang),
India
(Andaman & Nicobar Islands
[Andaman Islands],
Andhra Pradesh,
Arunachal Pradesh,
Assam,
Bihar,
Chhattisgarh,
Delhi,
Goa,
Gujarat,
Haryana,
Himachal Pradesh,
Jammu & Kashmir,
Jharkhand,
Karnataka,
Kerala,
Madhya Pradesh,
Maharashtra,
Mizoram,
Odisha,
Punjab,
Rajasthan,
Tamil Nadu,
Telangana,
Tripura,
Uttar Pradesh,
Uttarakhand,
West Bengal),
Indonesia
(Bali,
Bangka,
Java,
Sumatra),
Laos,
Malaysia
(West Malaysia),
Myanmar,
Nepal,
Pakistan,
Singapore,
Sri Lanka,
Taiwan,
Thailand,
Turkmenistan,
Vietnam.
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Ptyas mucosa
© Lars Skov
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Ptyas multicincta
|
Many-banded Green Snake, Northern Chinese Green Snake
|
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Vielbändige Grünnatter
|
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Mangebåndet Smaragdsnog
|
|
1907 | |
Ablabes multicinctus Roux |
|
| |
Eurypholis multicinctus Pope 1935 |
|
| |
Liopeltis multicinctus Angel & Bourret 1933 |
|
| |
Liopeltis major multicinctus Bourret 1936 |
|
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Opheodrys multicinctus Schmidt & Necker 1936 |
|
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Eurypholis multicinctus Pope 1936 |
|
| |
Entechinus multicinctus Cundall 1981 |
|
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Cyclophiops multicinctus Zhao & Adler 1993 |
|
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Ptyas multicincta Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016 |
|
1920 | |
Ablabes retrofasciatus Angel (syn. Smith 1943) |
|
1921 | |
Zamenis moi Smith (syn. Smith 1943) |
|
1929 | |
Ablabes multicinctus bicolor Angel (syn. Smith 1943) |
|
| |
Liopeltis major bicolor Bourret 1936 |
|
1931 | |
Liopeltis sinii Fan (syn. Iskandar & Colijn 2001) |
Distribution:
China
(Guangdong,
Guangxi,
Hainan,
Yunnan),
Laos,
Thailand,
Vietnam.
|
|
|
Ptyas nigromarginata
|
Oriental Green Rat Snake, Black-bordered Rat Snake
|
|
Grüne Gekielte Rattennatter
|
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Sortrandet Rottesnog
|
|
1854 | |
Coluber nigromarginatus Blyth |
|
| |
Zaocys nigromarginatus Günther 1864 |
|
| |
Ptyas nigromarginatus Wall 1923 |
|
| |
Zaocys dhumnades nigromarginatus Bourret 1936 |
Remarks:
Previously included the population now assigned to bachmaensis.
Records from Taiwan refer to oshimai, regarded as a subspecies of nigromarginatus by Wang & Wang (1956), but as a synonym of dhumnades by Zhao & Adler (1993).
Presence in Nepal needs confirmation (Kästle, Rai & Schleich 2013).
Distribution:
Bangladesh,
Bhutan,
China
(Chongqing,
Gansu,
Guizhou,
Sichuan,
Tibet,
Yunnan),
India
(Arunachal Pradesh,
Meghalaya,
Nagaland,
Sikkim,
West Bengal),
Laos,
Myanmar,
Thailand,
Vietnam.
|
|
|
Ptyas semicarinata
|
Ryukyu Green Snake
|
|
Ryukyu-Grünnatter
|
|
Ryukyu-smaragdsnog
|
|
1861 | |
Eurypholis semicarinatus Hallowell |
|
| |
Ablabes semicarinatus Boulenger 1894 |
|
| |
Liopeltis semicarinata Stejneger 1907 |
|
| |
Opheodrys semicarinatus Schmidt & Necker 1936 |
|
| |
Entechinus semicarinatus Cundall 1981 |
|
| |
Cyclophiops semicarinatus Ota, Shiroma & Hikida 1995 |
|
| |
Ptyas semicarinata Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016 |
|
1868 | |
Cyclophis nebulosus Günther (syn. Stejneger 1907) |
Distribution:
Japan
(Ryukyu Islands
[Okinawa Islands,
Satsunan Islands
(Amami Islands,
Tokara Islands)]).
|
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