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© Rune Midtgaard


Taxonomy of the family Agamidae
Bibliography of the genus Gonocephalus
Biodiversity of the family Agamidae








Genus
Gonocephalus

Western Anglehead Agamas*, Angle-headed Agamas, Angle-headed Lizards, Hump-headed Agamas, Narrow-headed Tree Lizards

Westliche Winkelkopfagamen*

Vestlige Vinkelhovedagamer*

1805 Lophyrus Dumeril [not Lophyrus Poli 1791 (Mollusca); not Lophyrus Latreille 1802 (Insecta: Hymenoptera)] (type species: not designated, but presumably Iguana chamaeleontina Laurenti 1768; syn. Wermuth 1967)
1825 Gonocephalus Kaup (type species: Lophyrus tigrinus Dumeril & Bibron 1837)
1827 Goniocephalus Kaup [emendation]
1830 Gonyocephalus Wagler [emendation]
1845 Dilophyrus Gray (type species: Dilophyrus grandis Gray 1845; syn. Cogger, Cameron & Cogger 1983)
1846 Gonycephalus Agassiz [emendation]
Contents: 16 species, of which 9 (56.3%) are endemic.
Remarks: Previously included Malayodracon robinsonii and the genera Hypsilurus and Lophosaurus.
Distribution: SE. Asia, Malay Archipelago.
Reported from: Brunei, Indonesia (Bangka, Banyak Islands, Batu Islands, Java, Kalimantan, Natuna Islands [Bunguran], Nias, Siberut, Sipura, Sumatra), Laos, Malaysia (Sabah, Sarawak, West Malaysia [incl. Penang Island, Seribuat Archipelago (incl. Tioman)]), Philippines (Bohol, Calamian Islands, Camiguin Sur, Catanduanes, Cebu, Dinagat, Leyte, Luzon, Masbate, Mindanao, Mindoro, Negros, Panay, Polillo, Samar, Sibuyan, Talikud), Thailand, Vietnam.

Gonocephalus abbotti

Southern Thai Anglehead Agama*, (Abbott's Anglehead Agama)

Abbotts Winkelkopfagame

Sydthailandsk Vinkelhovedagam*

1922 Gonocephalus abbotti Cochran
Gonocephalus doriae abbotti Manthey & Denzer 1993

Remarks: Regarded as a subspecies of doriae by some authors (e.g., Manthey & Denzer 1993; Manthey 2010), but most recent works treat it as a separate species (e.g., Grismer 2008; Grismer & Aun 2008).
Distribution: Malaysia (West Malaysia), Thailand.


Gonocephalus bellii

Blue-necked Anglehead Agama, (Bell's Anglehead Agama)

Blaukehl-Winkelkopfagame

Blåhalset Vinkelhovedagam*

1837 Lophyrus bellii Dumeril & Bibron
Gonyocephalus bellii Boulenger 1885

Remarks: Records from Borneo (e.g., Wermuth 1967) refer to borneensis (Manthey & Denzer 1992; Das 2007c). Sarker & Sarker's (1985) record from Bangladesh is regarded as erroneous.
Distribution: Malaysia (West Malaysia [incl. Penang Island]), Thailand.


Gonocephalus beyschlagi

Northern Sumatra Anglehead Agama*, (Beyschlag's Anglehead Agama)

Beyschlags Winkelkopfagame

Nordsumatransk Vinkelhovedagam*

1892 Gonyocephalus beyschlagi Boettger

Distribution: Indonesia (Sumatra).


Gonocephalus borneensis

Bornean Anglehead Agama

Borneo-Winkelkopfagame

Borneo-vinkelhovedagam*

1848 Lophyrus borneensis Schlegel
Gonyocephalus borneensis Boulenger 1885
1872 Tiaris miotympanum Günther (Manthey & Denzer 1992)
Gonyocephalus miotympanum Boulenger 1885
1991 Gonocephalus denzeri Manthey (Manthey & Grossmann 1997)

Remarks: Previously regarded as a synonym of bellii (e.g., Wermuth 1967). Revalidated by Manthey & Denzer (1992). Records from the Malay Peninsula probably refer to bellii or liogaster (Denzer & Manthey 1991).
Distribution: Brunei, Indonesia (Kalimantan), Malaysia (Sabah, Sarawak).


Gonocephalus chamaeleontinus

Chameleon Anglehead Agama, (Tioman Anglehead Agama)

Chamäleon-Winkelkopfagame

Kamæleon-vinkelhovedagam*

1768 Iguana chamaeleontina Laurenti
Gonyocephalus chamaeleontinus Wagler 1830
1802 Lacerta scutata Shaw (Manthey & Denzer 1993)
1820 Agama gigantea Kuhl (Manthey & Denzer 1993)
1820 Agama tigrina Merrem (Manthey & Denzer 1993)
1826 Lophyrus gigas Boie (Manthey & Denzer 1993)
1827 Agama superciliosa Hardwicke & Gray (Manthey & Denzer 1993)
1844 Galeotes lophyrus Schlegel [part] (Manthey & Denzer 1993)
1848 Lophyrus sumatranus Schlegel (Manthey & Denzer 1993)
Gonyocephalus sumatranus Boulenger 1885

Remarks: Das (2010) erroneously reported this species from the West Malaysian mainland (Grismer 2011). Reported from Kalimantan by Iskandar (2004), but no vouchers exist (Das 2007).
Distribution: Indonesia (Bangka, Banyak Islands, Batu Islands, Java, Natuna Islands, Nias, Siberut, Sipura, Sumatra), Malaysia (West Malaysia [incl. Seribuat Archipelago (incl. Tioman)]).


Gonocephalus doriae

Sarawak Anglehead Agama*, (Marquis Doria's Anglehead Agama)

Dorias Winkelkopfagame

Sarawak-vinkelhovedagam*

1871 Gonyocephalus doriae Peters

Remarks: Some authors regard abbotti as a subspecies of doriae (e.g., Manthey & Denzer 1993).
Distribution: Indonesia (Kalimantan), Malaysia (Sabah, Sarawak), Thailand.


Gonocephalus grandis

Greater Anglehead Agama, Great Anglehead Agama, Giant Anglehead Lizard, Creek Anglehead

Große Winkelkopfagame

Stor Vinkelhovedagam*

1845 Dilophyrus grandis Gray
Gonyocephalus grandis Schlegel 1858

Distribution: Brunei, Indonesia (Kalimantan, Nias, Siberut, Sipura, Sumatra), Laos, Malaysia (Sabah, Sarawak, West Malaysia [incl. Penang Island, Seribuat Archipelago (incl. Tioman)]), Thailand, Vietnam.


Gonocephalus interruptus

Mindoro Anglehead Agama, Mindoro Angle-headed Dragon, Mindanao Anglehead Agama

Mindanao-Winkelkopfagame

Mindoro-vinkelhovedagam*

1885 Gonyocephalus interruptus Boulenger

Distribution: Philippines (Bohol, Mindanao, Mindoro, Talikud),


Gonocephalus klossi

Western Sumatra Anglehead Agama*, (Kloss's Anglehead Agama)

Kloss' Winkelkopfagame

Vestsumatransk Vinkelhovedagam*

1920 Gonyocephalus klossi Boulenger

Distribution: Indonesia (Sumatra).


Gonocephalus kuhlii

Java Anglehead Agama*, (Kuhl's Anglehead Agama)

Kuhls Winkelkopfagame

Javansk Vinkelhovedagam*

1848 Lophyrus kuhlii Schlegel
Gonyocephalus kuhlii Boulenger 1885
1844 Galeotes lophyrus Schlegel [part] (Manthey & Denzer 1993)

Remarks: Previously regarded as a synonym of chamaeleontinus (e.g., Wermuth 1967).
Distribution: Indonesia (Java, Sumatra).


Gonocephalus lacunosus

Sikulikap Anglehead Agama

Sikulikap-Winkelkopfagame

Sikulikap-vinkelhovedagam*

1991 Gonocephalus lacunosus Manthey & Denzer

Distribution: Indonesia (Sumatra).


Gonocephalus liogaster

Blue-eyed Anglehead Agama, Orange-ringed Anglehead Agama, Bornean Anglehead Agama, Comb-crested Anglehead

Blauaugen-Winkelkopfagame

Blåøjet Vinkelhovedagam*

1872 Tiaris liogaster Günther
Gonyocephalus liogaster Boulenger 1885
1887 Gonyocephalus herveyi Boulenger (Dring 1979; Manthey & Denzer 1992)

Remarks: Das (2010) regarded herveyi as a valid species.
Distribution: Brunei, Indonesia (Kalimantan, Natuna Islands [Bunguran], Sumatra), Malaysia (Sabah, Sarawak, West Malaysia).


Gonocephalus megalepis

Sumatran Large-scaled Anglehead Agama, Green Anglehead Agama

Grüne Winkelkopfagame

Storskællet Vinkelhovedagam*

1860 Lophyrus megalepis Bleeker
Gonyocephalus megalepis Boulenger 1887
1872 Tiaris tuberculatus Günther (Boulenger 1887; Manthey & Denzer 1991)

Remarks: Presence in Borneo needs confirmation (Denzer & Manthey 1991).
Distribution: Indonesia (Sumatra).


Gonocephalus mjoebergi

Mount Murud Anglehead Agama*, (Mjöberg's Anglehead Agama)

Mjöbergs Winkelkopfagame

Murud-vinkelhovedagam*

1925 Gonocephalus mjöbergi Smith
Gonocephalus mjoebergi [emendation] Wermuth 1967

Distribution: Malaysia (Sarawak).


Gonocephalus semperi

White-spotted Anglehead Agama, White-spotted Angle-headed Dragon, (Semper's Anglehead Agama)

Sempers Winkelkopfagame

Hvidplettet Vinkelhovedagam*

1867 Lophyrus semperi Peters
Gonyocephalus semperi Boulenger 1885

Distribution: Philippines (Bohol, Calamian Islands, Camiguin Sur, Cebu, Leyte, Mindanao, Mindoro, Negros, Panay).


Gonocephalus sophiae

Dark-spotted Anglehead Agama, Dark-spotted Anglehead Agama, Negros Forest Dragon, (Sophia's Anglehead Agama)

Sophias Winkelkopfagame

Mørkplettet Vinkelhovedagam*

1845 Tiaris sophiae Gray
Gonyocephalus sophiae Boulenger 1885
1845 Tiaris bellii Gray [not Lophyrus bellii Duméril & Bibron 1837] (Taylor 1922)
1868 Tiaris petersii Günther (Wermuth 1967)

Distribution: Philippines (Cebu, Luzon, Masbate, Mindanao, Negros, Panay, Polillo, Sibuyan).