Reptiles of Samar [Philippines] Related checklists:  Philippines Bibliography of the reptiles of Samar Taxonomic history of the reptiles of Samar Introduced species

Home - Taxonomy - Geography - Biodiversity - Literature - Purchase RepFocus

Recent updates

Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: April 14th, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus mindorensis Philippine Freshwater Crocodile Trutnau & Sommerlad 2006

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama David, Pauwels, Lays & Lenglet 2006 Hallermann 2005 Genus Draco (Gliding Dragons)   Draco bimaculatus Two-spotted Gliding Dragon Gaulke 1994 Manthey 2008 McGuire & Alcala 2000 McGuire & Kiew 2001 Musters 1983 Ross & Lazell 1990 Wermuth 1967   Draco mindanensis Mindanao Gliding Dragon Gaulke 1994 McGuire & Alcala 2000 Realubit, Diesmos, Lorenzo, Layola, Diesmos, Brown, Brown, Cobb, Bergmann, Morinaga, Siler, Huron & Siler 2015   Draco ornatus White-spotted Gliding Dragon Gaulke 1994 Manthey 2008 McGuire & Alcala 2000 McGuire & Kiew 2001 Villanueva, Fernandez, Angeles, Tolentino, Obena & Buot 2021   Draco reticulatus Reticulated Gliding Dragon Manthey 2008 McGuire & Alcala 2000 McGuire & Kiew 2001 Villanueva, Fernandez, Angeles, Tolentino, Obena & Buot 2021 Genus Gonocephalus (Western Anglehead Agamas)   Gonocephalus semperi White-spotted Anglehead Agama Welton, Siler, Grismer, Diesmos, Sites & Brown 2017 Genus Hydrosaurus (Sail-tailed Agamas)   Hydrosaurus pustulatus Philippine Sail-tailed Agama Gaulke 1994  Family Gekkonidae (Typical Geckos) Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus annulatus Mindanao Banded Bent-toed Gecko Gaulke 1994 Gaulke 1996   * Cyrtodactylus sumuroi Taft Forest Bent-toed Gecko Obena, Tolentino, Villanueva, Fernandez, Angeles & Buot 2021 Villanueva, Fernandez, Angeles, Tolentino, Obena & Buot 2021 Welton, Siler, Linkem, Diesmos & Brown 2010 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Brown & Alcala 1978 Gaulke 1994 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus platyurus Flat-tailed Gecko Gaulke 1994 Genus Hemiphyllodactylus (Slender Geckos)   Hemiphyllodactylus insularis Philippine Slender Gecko Brown & Alcala 1978 [as typus] Gaulke 1994 [as typus] Genus Lepidodactylus (Scaly-toed Geckos)   Lepidodactylus aureolineatus Golden-lined Scaly-toed Gecko Alcala 1986 Brown & Alcala 1978 Gaulke 1994   Lepidodactylus planicauda Broad-tailed Scaly-toed Gecko Alcala 1986 Brown & Alcala 1978 Gaulke 1994 Genus Pseudogekko (Smooth-scaled Geckos)   Pseudogekko brevipes Orange-spotted Smooth-scaled Gecko Alcala 1986 Brown & Alcala 1978 Davis, Watters, Koehler, Whitsett, Huron, Brown, Diesmos & Siler 2015 Huron, Realubit, Cobb, Brown, Bergmann Morinaga, Diesmos, Diesmos, Brown & Siler 2016   Pseudogekko ditoy Leyte Smooth-scaled Gecko Huron, Realubit, Cobb, Brown, Bergmann Morinaga, Diesmos, Diesmos, Brown & Siler 2016   Pseudogekko pungkaypinit Southern Philippine Smooth-scaled Gecko Huron, Realubit, Cobb, Brown, Bergmann Morinaga, Diesmos, Diesmos, Brown & Siler 2016  Family Scincidae (Skinks) Genus Brachymeles (Indo-Malayan Slender Skinks)   Brachymeles orientalis Bario Dusita Slender Skink Obena, Tolentino, Villanueva, Fernandez, Angeles & Buot 2021 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012   Brachymeles samad Eastern Visayas Slender Skink Brown & Alcala 1980 [as hilong] Gaulke 1994 [as hilong] Obena, Tolentino, Villanueva, Fernandez, Angeles & Buot 2021 Siler, Jones, Diesmos, Diesmos & Brown 2012   * Brachymeles samarensis Samar Two-toed Slender Skink Alcala 1986 Brown & Alcala 1980 Gaulke 1994 Obena, Tolentino, Villanueva, Fernandez, Angeles & Buot 2021 Siler & Brown 2010 Siler, Fuiten, Jones, Alcala & Brown 2011 Siler, Jones, Diesmos, Diesmos & Brown 2012 Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Brown & Alcala 1980 Brown 1991 Gaulke 1994 Gaulke 1996 Genus Eutropis (South Asian Sun Skinks)   Eutropis caraga Misamis Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020   Eutropis indeprensa Mindoro Sun Skink Brown & Alcala 1980 Gaulke 1994 Gaulke 1996   Eutropis islamaliit Semirara Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020   Eutropis lapulapu Mactan Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020   Eutropis multicarinata Many-keeled Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020 Villanueva, Fernandez, Angeles, Tolentino, Obena & Buot 2021   Eutropis multifasciata Many-lined Sun Skink David, Pauwels, Lays & Lenglet 2006 Villanueva, Fernandez, Angeles, Tolentino, Obena & Buot 2021 Genus Lamprolepis (Malay Archipelago Tree Skinks)   Lamprolepis smaragdina Emerald Tree Skink Brown & Alcala 1980 David, Pauwels, Lays & Lenglet 2006 Gaulke 1994 Genus Lipinia (Indo-Pacific Striped Skinks)   Lipinia pulchella Yellow-striped Slender Tree Skink Brown & Alcala 1980 Gaulke 1994 Supsup, Redoblado & Guinto 2016   Lipinia quadrivittata Black-striped Slender Tree Skink Brown & Alcala 1956   Lipinia semperi Mindanao Striped Skink Alcala 1986 Genus Otosaurus (Philippine Giant Forest Skink)   Otosaurus cumingii Philippine Giant Forest Skink Brown & Rico 2009 Obena, Tolentino, Villanueva, Fernandez, Angeles & Buot 2021 Genus Parvoscincus (Philippine Dwarf Forest Skinks)   Parvoscincus steerei Guimaras Dwarf Forest Skink Gaulke 1994 Genus Pinoyscincus (Philippine Forest Skinks)   Pinoyscincus coxi Agusan Forest Skink Alcala 1986 Brown & Alcala 1980 Gaulke 1994   Pinoyscincus jagorii Samar Forest Skink Brown & Alcala 1980 Gaulke 1994 Obena, Tolentino, Villanueva, Fernandez, Angeles & Buot 2021 Ross & Lazell 1990   Pinoyscincus llanosi Leyte Forest Skink Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Gaulke 1994   Pinoyscincus mindanensis Mindanao Forest Skink Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus acutus Sharp-nosed Forest Skink Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Gaulke 1994 Ross & Lazell 1990   Sphenomorphus fasciatus Philippine Banded Forest Skink Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Gaulke 1994   Sphenomorphus variegatus Philippine Black-spotted Forest Skink Gaulke 1994  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus samarensis Samar Water Monitor Boehme 2003 [as V. cumingi] David, Pauwels, Lays & Lenglet 2006 [as V. cumingi] Gaulke 1994 [as V. cumingi] Koch, Auliya, Schmitz, Kuch & Boehme 2007 [as V. cumingi] Koch, Gaulke & Boehme 2010 Lisle 1996 [as V. cumingi] Obena, Tolentino, Villanueva, Fernandez, Angeles & Buot 2021 Welton, Travers, Siler & Brown 2014

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake Gaulke 1994 Iskandar & Colijn 2001 Leviton 1967 Leviton, Siler, Weinell & Brown 2018 Wallach, Williams & Boundy 2014 Genus Boiga (Australasian Cat Snakes)   Boiga angulata Leyte Cat Snake Gaulke 1994 Iskandar & Colijn 2001   Boiga dendrophila Eastern Mangrove Cat Snake Brongersma 1934 Gaulke 1994 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Orlov & Ryabov 2002 Genus Calamaria (Oriental Reed Snakes)   Calamaria lumbricoidea Variable Reed Snake Alcala 1986 Gaulke 1994 Inger & Marx 1965 Iskandar & Colijn 2001 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea paradisi Paradise Flying Tree Snake Gaulke 1994 Iskandar & Colijn 2001 Leviton 1964 Leviton, Siler, Weinell & Brown 2018 Mertens 1968 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus erythrura Philippine Rat Snake Gaulke 1994 Iskandar & Colijn 2001 Leviton 1979 Leviton, Siler, Weinell & Brown 2018 Schulz 1996 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis marenae Philippine Painted Bronzeback Gaulke 1994 [as pictus] Iskandar & Colijn 2001 [as pictus] Leviton 1968 [as pictus] Leviton, Siler, Weinell & Brown 2018 Vogel & Rooijen 2008   Dendrelaphis philippinensis Mindanao Stripe-tailed Bronzeback Gaulke 1994 [as terrificus] Iskandar & Colijn 2001 [as terrificus] Leviton 1968 [as terrificus] Leviton, Siler, Weinell & Brown 2018 Rooijen & Vogel 2012 Genus Dryophiops (Malayan Vine Snakes)   Dryophiops philippina Philippine Vine Snake Gaulke 1994 Iskandar & Colijn 2001 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma oxycephalum Red-tailed Green Rat Snake Wogan, Vogel, Neang, Nguyen, Demegillo, Diesmos & Gonzalez 2012 Genus Lycodon (Oriental Wolf Snakes)   Lycodon capucinus Island Wolf Snake Gaulke 1994 Iskandar & Colijn 2001 Leviton 1965 Leviton, Siler, Weinell & Brown 2018 O'Shea, Kusuma & Kaiser 2018 Vogel & Harikrishnan 2013   Lycodon dumerilii Mindanao Wolf Snake Gaulke 1994 Leviton, Siler, Weinell & Brown 2018 Villanueva, Fernandez, Angeles, Tolentino, Obena & Buot 2021   * Lycodon ferroni Samar Wolf Snake Iskandar & Colijn 2001 Lanza 1999 Leviton, Siler, Weinell & Brown 2018 Genus Oligodon (Kukri Snakes)   Oligodon ancorus Luzon Kukri Snake Gaulke 1994 Iskandar & Colijn 2001 Leviton 1963 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis auriculatus White-lined Keelback Alcala 1986 Gaulke 1994 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton 1970 Leviton, Siler, Weinell & Brown 2018   Rhabdophis lineatus Philippine Striped Keelback Gaulke 1994 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Genus Stegonotus (Indo-Australian Ground Snakes)   Stegonotus muelleri Philippine Ground Snake Alcala 1986 Barnes & Knierim 2018 Gaulke 1994 Iskandar & Colijn 2001 Kaiser, Kaiser & O'Shea 2018 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Villanueva, Fernandez, Angeles, Tolentino, Obena & Buot 2021 Genus Tropidonophis (Indo-Australian Keelbacks)   Tropidonophis dendrophiops Mindanao Keelback Gaulke 1994 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018  Family Cyclocoridae (Philippine Burrowing Snakes, etc.) Genus Cyclocorus (Triangle-spotted Snakes)   Cyclocorus nuchalis Southern Triangle-spotted Snake Alcala 1986 Gaulke 1994 Iskandar & Colijn 2001 Leviton 1965 Leviton, Siler, Weinell & Brown 2018 Genus Levitonius (Waray Dwarf Burrowing Snake)   Levitonius mirus Waray Dwarf Burrowing Snake Weinell, Paluh, Siler & Brown 2020 Genus Oxyrhabdium (Philippine Burrowing Snakes)   Oxyrhabdium modestum Philippine Bandless Burrowing Snake Alcala 1986 Gaulke & Operiano 2006 Gaulke 1994 Iskandar & Colijn 2001 Leviton 1957 Leviton 1964 Leviton, Siler, Weinell & Brown 2018  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Calliophis (South Asian Coral Snakes)   Calliophis philippinus Philippine Striped Coral Snake Alcala 1986 Brown, Smart, Leviton & Smith 2018 Gaulke 1994 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Genus Hemibungarus (Barred Coral Snakes)   Hemibungarus sp. (Barred Coral Snake) Leviton 1963 [as Calliophis calligaster, based on Peters 1861] Genus Naja (Typical Cobras)   Naja samarensis Southeastern Philippine Spitting Cobra Alcala 1986 Gaulke 1994 Iskandar & Colijn 2001 Leviton 1964 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Wuster & Thorpe 1991 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Fernandez, Herr, Weinell & Brown 2019  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis atriceps Black-headed Sea Snake Alcala 1986 [as H. fasciatus] Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Genus Laticauda (Sea Kraits)   Laticauda colubrina Yellow-lipped Sea Krait Heatwole, Busack & Cogger 2005 Iskandar & Colijn 2001   Laticauda laticaudata Blue-lipped Sea Krait Iskandar & Colijn 2001 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Gaulke 1994 Iskandar & Colijn 2001 Leviton 1963 Leviton 1983 Leviton, Siler, Weinell & Brown 2018 Villanueva, Fernandez, Angeles, Tolentino, Obena & Buot 2021  Family Pareidae (Oriental Slug Eaters) Genus Aplopeltura (Blunt-headed Slug Eater)   Aplopeltura boa Blunt-headed Slug Eater Grismer, Diesmos, Gonzalez, Jose & Inger 2012 Leviton, Siler, Weinell & Brown 2018 Villanueva, Fernandez, Angeles, Tolentino, Obena & Buot 2021  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Gaulke 1994 Gaulke 1996 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999  Family Typhlopidae (Typical Blind Snakes) Genus Malayotyphlops (Malay Archipelago Blind Snakes)   Malayotyphlops ruber Samar Red Blind Snake Alcala 1986 Gaulke 1994 Hahn 1980 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 Genus Ramphotyphlops (Western Pacific Blind Snakes)   * Ramphotyphlops marxi Northwest Samar Blind Snake Gaulke 1994 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 Wallach 1993   Ramphotyphlops olivaceus Olive Blind Snake Alcala 1986 Gaulke 1994 Hahn 1980 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999  Family Viperidae (Vipers) Genus Parias (Malay Archipelago Pitvipers)   Parias flavomaculatus Yellow-spotted Pitviper Leviton, Siler, Weinell & Brown 2018 Genus Tropidolaemus (Temple Pitvipers)   Tropidolaemus philippensis South Philippine Temple Pitviper Iskandar & Colijn 2001 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018   Tropidolaemus subannulatus North Philippine Temple Pitviper Gaulke 1994 [as wagleri] Gaulke 1996 [as wagleri] Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 [as wagleri] Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 [as wagleri] Leviton 1964 [as wagleri] Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Obena, Tolentino, Villanueva, Fernandez, Angeles & Buot 2021 Vogel, David, Lutz, Rooijen & Vidal 2007

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle David, Pauwels, Lays & Lenglet 2006 Diesmos, Brown, Alcala & Sison 2008 Gaulke 1994 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021