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Taxonomy of the family Lamprophiidae
Bibliography of the genus Limaformosa
Biodiversity of the family Lamprophiidae








Genus
Limaformosa

Venus File Snakes*

Venus-Feilennattern*

Venusfilsnoge*

1847 Heterolepis Smith [not Heterolepis Nees 1834 (Insecta); not Heterolepis Fitzinger 1843 (Sauria)] (type species: Heterolepis capensis Smith 1847)
1858 Simocephalus Gray in Günther [substitute name for Heterolepis Smith 1847; not Simocephalus Schödler 1858 (Crustacea)]
2018 Limaformosa Broadley, Tolley, Conradie, Wishart, Trape, Burger, Kusamba, Zassi-Boulou & Greenbaum (type species: Heterolepis capensis Smith 1847)
Contents: 6 species, none of which are endemic.
Remarks: The species of this genus were previously included in Mehelya (e.g., Loveridge 1939) or, more recently, in Gonionotophis (e.g., Kelly, Branch, Broadley, Barker & Villet 2011). Priority of Simocephalus Schödler 1958 over Simocephalus Gray in Günther 1858 was discussed by Smith, Wallach, Williams, Pennak & Chiszar (1990).
Distribution: SubSaharan Africa.
Reported from: Angola, Benin, Botswana, Burkina Faso, Burundi, Cameroon, Central African Republic, Chad, Congo-Brazzaville, Congo-Kinshasa, Eritrea, Eswatini, Ethiopia, Gabon, Ghana, Guinea, Ivory Coast, Kenya, Liberia, Malawi, Mali, Mozambique, Namibia, Niger, Nigeria, Rwanda, Senegal, Sierra Leone, Somalia, South Africa (Gauteng, KwaZulu-Natal, Limpopo, Mpumalanga, North West Province), South Sudan, Sudan, Tanzania (incl. Zanzibar), Togo, Uganda, Zambia, Zimbabwe.

Limaformosa capensis

Cape File Snake, Southern Cape File Snake, Common File Snake

Kap-Feilennatter

Kapfilsnog

1847 Heterolepis capensis Smith
Simocephalus capensis Boulenger 1893
Mehelya capensis Barbour & Loveridge 1928
Gonionotophis capensis Kelly, Branch, Broadley, Barker & Villet 2011
Limaformosa capensis Broadley, Tolley, Conradie, Wishart, Trape, Burger, Kusamba, Zassi-Boulou & Greenbaum 2018
1874 Heterolepis gueinzee Peters (Loveridge 1939; Broadley 1983)

Remarks: Previously included savorgnani and chanleri. Gabonese records refer to savorgnani (Pauwels, Albert, Arrowood, Mvele, Casanova, Dodane, Morgan, Primault, Thepenier & Fenner 2017).
Distribution: Angola, Botswana, Congo-Kinshasa, Eswatini, Malawi, Mozambique, Namibia, South Africa (Gauteng, KwaZulu-Natal, Limpopo, Mpumalanga, North West Province), Tanzania (incl. Zanzibar), Zambia, Zimbabwe.


Limaformosa chanleri

Unicolor File Snake, (Chanler's File Snake)

Einfarbige Feilennatter*

Ensfarvet Filsnog*

1893 Simocephalus chanleri Stejneger
Mehelya chanleri Lönnberg 1910
Gonionotophis chanleri Lanza & Broadley 2014
Limaformosa chanleri Broadley, Tolley, Conradie, Wishart, Trape, Burger, Kusamba, Zassi-Boulou & Greenbaum 2018
1910 Simocephalus unicolor Boulenger (Lanza & Broadley 2014)
Mehelya unicolor Werner 1929
Mehelya capensis unicolor Laurent 1956
Gonionotophis unicolor Kelly, Branch, Broadley, Barker & Villet 2011

Other common names:
unicolor: Black Cape File Snake
Remarks: Previously regarded as a synonym of savorgnani (e.g., Loveridge 1939, 1957). Revalidated as a separate species (as unicolor) by Largen & Spawls (2010, without discussion) and Kelly, Branch, Broadley, Barker & Villet (2011), after which unicolor was synonymized with chanleri (Lanza & Broadley 2014).
Distribution: Burundi, Congo-Brazzaville, Congo-Kinshasa, Eritrea, Ethiopia, Kenya, Rwanda, Somalia, Tanzania (incl. Zanzibar), Uganda.


Limaformosa crossi

Savanna File Snake, (Crosse's File Snake)

Große Savannen-Feilennatter*

Stor Savannefilsnog*

1895 Simocephalus crossi Boulenger
Mehelya crossi Hughes 1983
Gonionotophis crossii Kelly, Branch, Broadley, Barker & Villet 2011
Limaformosa crossii Broadley, Tolley, Conradie, Wishart, Trape, Burger, Kusamba, Zassi-Boulou & Greenbaum 2018
1910 Simocephalus riggenbachi Sternfeld (Broadley 2007)
Mehelya riggenbachi Werner 1929

Distribution: Benin, Burkina Faso, Cameroon, Central African Republic, Chad, Ghana, Guinea, Ivory Coast, Mali, Niger, Nigeria, Senegal, Sierra Leone, Togo.


Limaformosa guirali

White-speckled Forest File Snake, Speckled Forest File Snake, (Guiral's File Snake)

Gesprenkelte Feilennatter*

Hvidspættet Filsnog*

1887 Heterolepis Guirali Mocquard
Simocephalus guirali Boulenger 1893
Mehelya guirali Lönnberg 1910
Gonionotophis guirali Kelly, Branch, Broadley, Barker & Villet 2011
Limaformosa guirali Broadley, Tolley, Conradie, Wishart, Trape, Burger, Kusamba, Zassi-Boulou & Greenbaum 2018
1908 Simocephalus baumanni Sternfeld (Loveridge 1939)
Mehelya baumanni Lönnberg 1910

Distribution: Benin, Cameroon, Gabon, Ghana, Guinea, Ivory Coast, Liberia, Nigeria, Sierra Leone, Togo.


Limaformosa savorgnani

Central African Forest File Snake, Northwestern Cape File Snake, (Savorgnan's File Snake)

Zentralafrikanische Feilennatter*

Centralafrikansk Filsnog*

1887 Heterolepis savorgnani Mocquard
Mehelya capensis savorgnani Loveridge 1939
Mehelya savorgnani Chirio & Ineich 2006
Gonionotophis savorgnani Kelly, Branch, Broadley, Barker & Villet 2011
Limaformosa savorgnani Broadley, Tolley, Conradie, Wishart, Trape, Burger, Kusamba, Zassi-Boulou & Greenbaum 2018
1901 Simocephalus phyllopholis Werner (Loveridge 1939, 1957)
Mehelya phyllopholis Lönnberg 1910
1907 Simocephalus butleri Boulenger (Loveridge 1939, 1957)
Mehelya butleri Lönnberg 1910
1910 Mehelya lamani Lönnberg (Loveridge 1939, 1957)
1913 Mehelya somaliensis Lönnberg & Andersson (Loveridge 1939, 1957)
1929 Mehelia fiechteri Scortecci (Lanza 1983)

Remarks: Previously included chanleri (and its synonym unicolor). Formerly regarded as a subspecies of capensis (e.g., Chippaux 2001, 2006), but treated here as a separate species, following Chirio & Ineich (2006) and Chirio & LeBreton (2007), although Pauwels & Vande weghe (2008) and Pauwels & Sallé (2009) retained savorgnani as a synonym of capensis, pending further studies. Lanza & Broadley (2014) provided evidence in support of the validity of savorgnani. Included for Angola, based on Lanza & Broadley (2014) and Branch (2018), although the species was not mentioned by Marques, Ceríaco, Blackburn & Bauer (2018).
Distribution: Angola, Cameroon, Central African Republic, Congo-Brazzaville, Congo-Kinshasa, Ethiopia, Gabon, Kenya, Somalia, South Sudan, Sudan, Uganda.


Limaformosa vernayi

Angolan File Snake

Angolische Feilennatter*

Angolafilsnog

1940 Mehelya vernayi Bogert
Gonionotophis vernayi Kelly, Branch, Broadley, Barker & Villet 2011
Limaformosa vernayi Broadley, Tolley, Conradie, Wishart, Trape, Burger, Kusamba, Zassi-Boulou & Greenbaum 2018

Distribution: Angola, Namibia.