Reptiles of Mali    Bibliography of the reptiles of Mali Taxonomic history of the reptiles of Mali Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Cynisca (West African Round-headed Worm Lizards)   Cynisca leucura White-tailed Worm Lizard Mane & Trape 2015

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   EX Crocodylus suchus West African Crocodile Boehme, Meinig & Roedel 1996 [as niloticus] Fuchs, Mertens & Wermuth 1974 Joger & Lambert 1996 [as niloticus] King & Burke 1989 [as niloticus] Lescure 2014 Trape, Trape & Chirio 2012 Wermuth & Mertens 1977 Genus Mecistops (African Slender-snouted Crocodiles)   EX Mecistops cataphractus West African Slender-snouted Crocodile Shirley, Carr, Nestler, Vliet & Brochu 2018 Genus Osteolaemus (Dwarf Crocodiles)   Osteolaemus tetraspis West African Dwarf Crocodile Trape, Trape & Chirio 2012

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama agama Common Rainbow Agama Böhme & Heath 2018 Böhme, Meinig & Rödel 1996 Geniez, Padial & Crochet 2011 Joger & Lambert 1996 Joger 1981 Leache, Grummer, Miller, Krishnan, Fujita, Böhme, Schmitz, LeBreton, Ineich, Chirio, Ofori-boateng, Eniang, Greenbaum, Rödel & Wagner 2016 [as A. picticauda] Mediannikov, Trape & Trape 2012 Papenfuss 1969 Schatti 1986 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Agama boensis Boe Agama Trape, Trape & Chirio 2012   Agama boueti Mali Agama Berre 1989 Böhme & Heath 2018 Geniez, Padial & Crochet 2011 Joger & Lambert 1996 Joger 1979 Mediannikov, Trape & Trape 2012 Trape, Trape & Chirio 2012   Agama boulengeri West African Desert Agama Berre 1989 Joger & Lambert 1996 Joger 1979 Joger 1981 Mediannikov, Trape & Trape 2012 Trape 2011 Trape, Trape & Chirio 2012 Wagner, Ineich, Leache, Wilms, Trape, Böhme & Schmitz 2009   Agama cristata African Crested Agama Mediannikov, Trape & Trape 2012 Trape 2011 Trape, Trape & Chirio 2012 Wermuth 1967   Agama gracilimembris Lesser Ground Agama Mediannikov, Trape & Trape 2012 Trape 2012 Trape, Trape & Chirio 2012   Agama sankaranica Sankaranian Agama Joger & Lambert 1996 Joger 1979 Joger 1981 Mediannikov, Trape & Trape 2012 Papenfuss 1969 Trape, Trape & Chirio 2012   Agama tassiliensis Tassili Agama Berre 1989 [as agama] Geniez, Padial & Crochet 2011 Joger & Lambert 1996 [as impalearis] Mediannikov, Trape & Trape 2012 [as Agama sp.] Papenfuss 1969 [as impalearis] Schleich, Kastle & Kabisch 1996 [as impalearis] Sindaco, Venchi & Grieco 2013 Trape, Trape & Chirio 2012   Agama weidholzi Senegal Agama Joger & Lambert 1996 Joger 1979 Joger 1981 Mediannikov, Trape & Trape 2012 Trape, Trape & Chirio 2012 Wagner & Bauer 2011 Wagner, Ineich, Leache, Wilms, Trape, Böhme & Schmitz 2009 Genus Trapelus (Afro-Asian Ground Agamas)   Trapelus mutabilis North African Desert Agama Berre 1989 Joger & Lambert 1996 Papenfuss 1969 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012 [as mutabilis complex] Wagner, Melville, Wilms & Schmitz 2011 Genus Uromastyx (Western Spiny-tailed Agamas)   Uromastyx dispar South Saharan Spiny-tailed Agama Joger & Lambert 1996 Trape, Trape & Chirio 2012 Wilms & Boehme 2001 Wilms 2001 Wilms, Boehme, Wagner, Lutzmann & Schmitz 2009   Uromastyx geyri Central Saharan Spiny-tailed Agama Berre 1989 Joger & Lambert 1996 Papenfuss 1969 Schleich, Kastle & Kabisch 1996 Trape, Trape & Chirio 2012 Wilms & Boehme 2001 Wilms 2001 Wilms, Boehme, Wagner, Lutzmann & Schmitz 2009  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo africanus Sahel Chameleon Boehme, Meinig & Roedel 1996 Glaw 2015 Joger & Lambert 1996 Klaver & Boehme 1997 Papenfuss 1969 Tilbury 2010 Trape, Trape & Chirio 2012   Chamaeleo gracilis Graceful Chameleon Trape 2010 Trape, Trape & Chirio 2012   Chamaeleo senegalensis Senegal Chameleon Joger & Lambert 1996 Joger 1981 Papenfuss 1969 Schatti 1986 Trape, Trape & Chirio 2012  Family Eublepharidae (Eyelid Geckos) Genus Hemitheconyx (African Fat-tailed Geckos)   Hemitheconyx caudicinctus West African Fat-tailed Gecko Joger & Lambert 1996 Loveridge 1947 Papenfuss 1969 Schatti 1986 Seufer, Kaverkin & Kirschner (eds.) 2005 Trape, Trape & Chirio 2012  Family Gekkonidae (Typical Geckos) Genus Hemidactylus (Half-toed Geckos)   Hemidactylus angulatus West African House Gecko Boehme, Meinig & Roedel 1996 [as brooki] Chirio & LeBreton 2007 Joger & Lambert 1996 [as "brooki complex"] Joger 1981 [as "cf. brookii"] Papenfuss 1969 Schatti 1986 [as "cf. brookii"] Trape, Trape & Chirio 2012   Hemidactylus mabouia Cosmopolitan House Gecko Joger & Lambert 1996 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus gutturalis Chevron-throated Dwarf Gecko Chirio & LeBreton 2007 Trape 2012 Trape, Trape & Chirio 2012 Genus Stenodactylus (Afro-Asian Thin-toed Geckos)   Stenodactylus petrii North African Thin-toed Gecko Joger & Lambert 1996 Papenfuss 1969 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Stenodactylus sthenodactylus Elegant Thin-toed Gecko Baha el Din 2006 Joger & Lambert 1996 Papenfuss 1969 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012 Genus Tropiocolotes (Afro-Asian Desert Dwarf Geckos)   Tropiocolotes tripolitanus Tripoli Dwarf Gecko Boehme & Heath 2018 Joger & Lambert 1996 Minton, Anderson & Anderson 1970 Papenfuss 1969 Ribeiro Junior, Tamar, Maza, Flecks, Wagner, Shacham, Calvo, Geniez, Crochet, Koch & Meiri 2022 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Szczerbak & Golubev 1996 Trape, Trape & Chirio 2012  Family Lacertidae (Typical Lizards) Genus Acanthodactylus (Fringe-toed Lizards)   Acanthodactylus boskianus Afro-Arabian Fringe-toed Lizard Joger & Lambert 1996 Papenfuss 1969 Salvador 1982 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Acanthodactylus dumerilii Lesser Fringe-toed Lizard Crochet, Geniez & Ineich 2003 [as senegalensis] Joger & Lambert 1996 Salvador 1982 Trape, Trape & Chirio 2012 [as senegalensis]   Acanthodactylus guineensis Guinea Fringe-toed Lizard Boehme, Meinig & Roedel 1996 Joger & Lambert 1996 Meinig & Boehme 2002 Trape, Trape & Chirio 2012   Acanthodactylus longipes Long-footed Fringe-toed Lizard Arnold 1983 Crochet, Geniez & Ineich 2003 Joger & Lambert 1996 Salvador 1982 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Acanthodactylus scutellatus Nidua Fringe-toed Lizard Arnold 1983 Crochet, Geniez & Ineich 2003 Joger & Lambert 1996 Papenfuss 1969 Salvador 1982 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012 Genus Heliobolus (Bushveld Lizards)   Heliobolus nitida Red-tailed Bushveld Lizard Trape, Trape & Chirio 2012 Genus Latastia (African Long-tailed Lizards)   Latastia longicaudata Common Long-tailed Lizard Boehme & Heath 2018 Joger & Lambert 1996 Papenfuss 1969 Trape, Trape & Chirio 2012 Genus Mesalina (Afro-Asian Desert Lizards)   Mesalina pasteuri Sahara Desert Lizard Joger & Lambert 1996 Sindaco & Jeremchenko 2008 Trape 2012 Trape, Trape & Chirio 2012 Yousefkhani, Marin & Rastegar-Pouyani 2015   Mesalina rubropunctata Red-spotted Desert Lizard Berre 1989 Bischoff 1991 Joger & Lambert 1996 Papenfuss 1969 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Szczerbak 1975 Trape, Trape & Chirio 2012 Genus Philochortus (Shield-backed Ground Lizards)   Philochortus zolii Saharan Shield-backed Ground Lizard Joger & Lambert 1996 [as spinalis] Papenfuss 1969 [as spinalis] Trape, Trape & Chirio 2012  Family Phyllodactylidae (Leaf-toed Geckos) Genus Ptyodactylus (Fan-toed Geckos)   Ptyodactylus togoensis Sahelian Fan-toed Gecko Boehme, Meinig & Roedel 1996 [as P. hasselquistii] Chirio & LeBreton 2007 [as P. ragazzii] Heimes 1987 [as P. ragazzii] Joger & Lambert 1996 [as P. ragazzii] Metallinou, Cervenka, Crochet, Kratochvil, Wilms, Geniez, Shobrak, Brito & Carranza 2015 Papenfuss 1969 [as P. hasselquistii] Sindaco & Jeremchenko 2008 [as P. ragazzii] Trape, Trape & Chirio 2012 [as P. ragazzii] Genus Tarentola (Wall Geckos)   Tarentola annularis Egyptian White-spotted Gecko Berre 1989 Boehme, Meinig & Roedel 1996 Joger & Lambert 1996 Joger 1984 Papenfuss 1969 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Tarentola ephippiata African Wall Gecko Berre 1989 Boehme & Heath 2018 [as "cf. ephippiata"] Joger & Lambert 1996 Joger 1984 Papenfuss 1969 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Tarentola hoggarensis Hoggar Wall Gecko Berre 1989 Joger 1984 Trape, Trape & Chirio 2012   Tarentola parvicarinata Small-keeled Gecko Berre 1989 Boehme, Meinig & Roedel 1996 Joger & Lambert 1996 Joger 1981 Joger 1984 Schatti 1986 Trape, Trape & Chirio 2012  Family Scincidae (Skinks) Genus Chalcides (Cylindrical Skinks)   Chalcides delislei Central Saharan Three-fingered Skink Boehme & Heath 2018 Joger & Lambert 1996 Papenfuss 1969 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Chalcides ocellatus Mediterranean Ocellated Skink Boehme & Heath 2018 Greenbaum 2005 Greenbaum, Campbell & Raxworthy 2006 Trape, Trape & Chirio 2012   Chalcides pulchellus West African Cylindrical Skink Carranza, Arnold, Geniez, Roca & Mateo 2008 Greenbaum 2005 Joger & Lambert 1996 Schatti 1986 Trape, Trape & Chirio 2012 Genus Panaspis (African Snake-eyed Skinks)   Panaspis togoensis Togo Snake-eyed Skink Schmitz, Ineich & Chirio 2005 Trape & Ineich 2012 Trape, Trape & Chirio 2012 Genus Scincopus (Tiger Skinks)   Scincopus fasciatus Banded Tiger Skink Joger & Lambert 1996 Lanza & Corsi 1981 Papenfuss 1969 Trape, Trape & Chirio 2012 Genus Scincus (Sand Fish)   Scincus albifasciatus White-banded Sand Fish Arnold & Leviton 1977 Geniez, Mateo, Geniez & Pether 2004 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis affinis Lesser Brown-flanked Skink Trape, Trape & Chirio 2012   Trachylepis keroanensis Kerouane Skink Penner, Doumbia, Kouame, Chirio, Sandberger-Loua, Boehme & Barej 2017   Trachylepis perrotetii Orange-flanked Skink Boehme, Meinig & Roedel 1996 Joger & Lambert 1996 Joger 1981 Papenfuss 1969 Schatti 1986 Trape, Trape & Chirio 2012   Trachylepis quinquetaeniata African Five-lined Skink Broadley & Bauer 1998 Joger & Lambert 1996 Joger 1979 Joger 1981 Koppetsch 2020 Papenfuss 1969 Trape, Trape & Chirio 2012  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus exanthematicus Western Savanna Monitor Bayless 2002 Bennett in Pianka, King & King (eds.) 2004 Boehme, Meinig & Roedel 1996 Joger & Lambert 1996 Papenfuss 1969 Trape, Trape & Chirio 2012   Varanus griseus Desert Monitor Bayless 2002 Joger & Lambert 1996 Papenfuss 1969 Trape, Trape & Chirio 2012   Varanus niloticus Nile Monitor Bayless 2002 Boehme & Ziegler 1997 Papenfuss 1969 Trape, Trape & Chirio 2012

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx muelleri West African Sand Boa Berre 1989 Hughes & Barry 1969 Joger & Lambert 1996 Papenfuss 1969 Stimson 1969 Trape & Mane 2017 Villiers 1975  Family Colubridae (Typical Snakes or Colubrids) Genus Afronatrix (West African Brown Water Snake)   Afronatrix anoscopus West African Brown Water Snake Chirio & LeBreton 2007 Trape & Mane 2017 Wallach, Williams & Boundy 2014 Genus Bamanophis (West African Whip Snake)   Bamanophis dorri West African Whip Snake Joger & Lambert 1996 Joger 1981 Papenfuss 1969 Schatti & Trape 2008 Trape & Mane 2017 Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hippocrepis West African Herald Snake Chippaux & Jackson 2019 Trape & Mane 2017   Crotaphopeltis hotamboeia Common Herald Snake Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017 Genus Dasypeltis (African Egg Eaters)   Dasypeltis confusa Diamondback Egg Eater Trape & Mane 2017 Trape, Mediannikov & Trape 2012   Dasypeltis fasciata Central African Egg Eater Chirio & LeBreton 2007 Joger & Lambert 1996 Schatti 1986   Dasypeltis gansi Savanna Egg Eater Bates, Trape & Spawls 2013 Trape & Mane 2006 Trape & Mane 2017 Trape, Mediannikov & Trape 2012   Dasypeltis latericia Senegalese Egg Eater Chippaux & Jackson 2019 Trape & Mane 2006 Trape & Mane 2017 Trape, Mediannikov & Trape 2012   Dasypeltis sahelensis Sahel Egg Eater Sindaco, Venchi & Grieco 2013 Trape & Mane 2006 Trape & Mane 2017 Trape, Mediannikov & Trape 2012 Genus Dispholidus (Boomslangs)   Dispholidus sp. (Boomslang) David & Ineich 1999 [as D. typus] Joger & Lambert 1996 [as D. t. typus] Papenfuss 1969 [as D. typus] Spawls & Branch 1995 [as D. typus] Trape & Mane 2017 [as D. typus] Villiers 1965 [as D. typus] Genus Grayia (Equatorial African Water Snakes)   Grayia smythii Equatorial Water Snake Chirio & LeBreton 2007 Trape & Mane 2017 Genus Meizodon (African Smooth Snakes)   Meizodon coronatus West African Crowned Smooth Snake Joger & Lambert 1996 Papenfuss 1969 Schatti 1985 Trape & Mane 2017   Meizodon regularis East African Crowned Smooth Snake Schatti 1985 Genus Natriciteres (African Marsh Snakes)   Natriciteres olivacea Olive Marsh Snake Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017 Genus Philothamnus (African Green Snakes)   Philothamnus irregularis Irregular Green Snake Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017 Villiers 1965   Philothamnus semivariegatus Variegated Bush Snake Joger & Lambert 1996 Papenfuss 1969   Philothamnus smithii Smith's Bush Snake Trape & Mane 2017 Genus Spalerosophis (Diadem Snakes)   Spalerosophis diadema Common Diadem Snake Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus tripolitanus North African Cat Snake Berre 1989 [as dhara] Crochet, Rasmussen, Wilms, Geniez, Trape & Boehme 2008 Joger & Lambert 1996 [as obtusus] Kramer & Schnurrenberger 1963 Papenfuss 1969 Sindaco, Venchi & Grieco 2013 Trape & Mane 2017 Wilms, Wagner & Spawls 2021   Telescopus variegatus Variable Cat Snake Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Elapsoidea (African Garter Snakes)   Elapsoidea semiannulata Half-banded Garter Snake David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Spawls & Branch 1995 Trape & Mane 2017 Genus Naja (Typical Cobras)   Naja haje Egyptian Cobra Sindaco, Venchi & Grieco 2013 Trape & Mane 2017 Trape, Chirio, Broadley & Wuster 2009   Naja katiensis Brown Spitting Cobra Broadley 1968 Chippaux & Jackson 2019 David & Ineich 1999 Joger & Lambert 1996 Schatti 1986 Spawls & Branch 1995 Trape & Mane 2017 Wallach, Williams & Boundy 2014 Wuster, Crookes, Ineich, Mane, Pook, Trape & Broadley 2007   Naja nigricollis Black-necked Spitting Cobra Joger & Lambert 1996 Papenfuss 1969 Spawls & Branch 1995 Trape & Mane 2017 Villiers 1965 Wallach, Williams & Boundy 2014   Naja savannula West African Banded Cobra David & Ineich 1999 [as N. melanoleuca] Joger & Lambert 1996 [as N. melanoleuca] Schatti 1986 [as N. melanoleuca] Spawls & Branch 1995 [as N. melanoleuca] Trape & Mane 2017 [as "cf. melanoleuca" [=savanna form]] Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018   Naja senegalensis Senegalese Cobra Shi, Vogel, Ding, Rao, Liu, Zhang, Wu & Chen 2022 Trape & Mane 2017 Trape, Chirio, Broadley & Wuster 2009  Family Lamprophiidae (House Snakes, etc.) Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas unicolor Equatorial Purple-glossed Snake Trape & Mane 2017 Genus Atractaspis (Stiletto Snakes)   Atractaspis aterrima Western Forest Stiletto Snake David & Ineich 1999 Joger & Lambert 1996 Schatti 1986 Spawls & Branch 1995 Trape & Mane 2017 Wallach, Williams & Boundy 2014   Atractaspis dahomeyensis Dahomey Stiletto Snake David & Ineich 1999 Joger & Lambert 1996 Schatti 1986 Spawls & Branch 1995 Trape & Mane 2006 Trape & Mane 2017 Wallach, Williams & Boundy 2014   Atractaspis micropholis Sahelian Stiletto Snake David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017 Trape, Mane & Ineich 2006 Wallach, Williams & Boundy 2014   Atractaspis watsoni Sokoto Stiletto Snake Chirio & LeBreton 2007 Trape & Mane 2006 Trape & Mane 2017 Trape, Mane & Ineich 2006 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon fuliginosus Common African House Snake Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017   Boaedon lineatus Lined House Snake Joger & Lambert 1996 Schatti 1986 Trape & Mane 2006 Trape & Mane 2017 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Hughes 2004 Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017   Dromophis praeornatus Ornate Olympic Snake Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017 Villiers 1965 Genus Gonionotophis (Lesser File Snakes)   Gonionotophis grantii Savanna Dwarf File Snake Chirio & LeBreton 2007 Trape & Mane 2017 Genus Limaformosa (Venus File Snakes)   Limaformosa crossi Savanna File Snake Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017 Villiers 1965 Genus Lycophidion (African Wolf Snakes)   Lycophidion albomaculatum White-spotted Wolf Snake Chippaux & Jackson 2019 Condamin 1994 Joger & Lambert 1996 Trape & Mane 2006 Trape & Mane 2017   Lycophidion irroratum Western Forest Wolf Snake Trape & Mane 2017   Lycophidion semicinctum Semi-annulated Wolf Snake Chippaux & Jackson 2019 Joger & Lambert 1996 Trape & Mane 2006 Trape & Mane 2017 Villiers 1965 Genus Malpolon (Montpellier Snakes)   Malpolon moilensis Moila Snake Berre 1989 Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017 Genus Polemon (African Snake Eaters)   Polemon neuwiedi West African Snake Eater Trape & Mane 2006 Trape & Mane 2017 Genus Prosymna (Shovel-snout Snakes)   Prosymna collaris Collared Shovel-snout Chirio & LeBreton 2007 Chirio, Ineich, Schmitz & Trape 2011 Trape & Mane 2006 Trape & Mane 2017   Prosymna greigerti Sudan Savanna Shovel-snout Chirio & LeBreton 2007 Chirio, Ineich, Schmitz & Trape 2011 Papenfuss 1969 [as P. laurenti] Trape & Mane 2006 Trape & Mane 2017 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis afroccidentalis West African Sand Snake Berre 1989 [as P. sibilans] Brito, Rebelo, Crochet & Geniez 2008 [as P. sibilans] Papenfuss 1969 [as P. sibilans] Trape & Mane 2017 [as "aff. sibilans"] Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019 Villiers 1965 [as P. sibilans]   Psammophis elegans Elegant Sand Snake Joger & Lambert 1996 Joger 1981 Papenfuss 1969 Trape & Mane 2017   Psammophis phillipsii West African Olive Grass Snake Boehme, Meinig & Roedel 1996 [as "cf. phillipsii"] Joger & Lambert 1996 Joger 1981 Schatti 1986 Trape & Mane 2017 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis schokari Arabian Sand Snake Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017 Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis oxyrhynchus Western Beaked Snake Chirio & Ineich 1991 Joger & Lambert 1996 Joger 1981 Papenfuss 1969 Trape & Mane 2017  Family Leptotyphlopidae (Thread Snakes) Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis adleri Chadian Thread Snake Trape & Mane 2017   Myriopholis albiventer White-bellied Thread Snake Trape & Mane 2006 Trape & Mane 2017   Myriopholis algeriensis Algerian Thread Snake Hahn & Wallach 1998 Sindaco, Venchi & Grieco 2013 Trape & Mane 2017   Myriopholis boueti Mali Thread Snake Adalsteinsson, Branch, Trape, Vitt & Hedges 2009 Hahn & Wallach 1998 Hahn 1980 Ineich & Prudent 2014 Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017   Myriopholis rouxestevae Senegal Thread Snake Trape & Mane 2006 Genus Rhinoguinea (West African Long-tailed Thread Snake)   * Rhinoguinea magna West African Long-tailed Thread Snake Trape & Mane 2017 Trape 2014 Genus Rhinoleptus (West African Sharp-nosed Thread Snake)   Rhinoleptus koniagui West African Sharp-nosed Thread Snake Adalsteinsson, Branch, Trape, Vitt & Hedges 2009 Jallow & Trape 2013 Trape & Mane 2006 Trape & Mane 2017 Trape 2014 Genus Tricheilostoma (West African Forest Thread Snakes)   Tricheilostoma bicolor Bicoloured Thread Snake Hahn & Wallach 1998 Hahn 1980 [as Leptotyphlops brevicaudus] Joger & Lambert 1996 McDiarmid, Campbell & Toure 1999 [as brevicaudus] Papenfuss 1969 [as brevicauda] Trape & Mane 2006 Trape & Mane 2017 Villiers 1965  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python regius Ball Python Joger & Lambert 1996 McDiarmid, Campbell & Toure 1999 Trape & Mane 2017   Python sebae Northern Rock Python Boehme, Meinig & Roedel 1996 Broadley 1984 Joger & Lambert 1996 Papenfuss 1969 Spawls & Branch 1995 Trape & Mane 2017 Villiers 1965  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops lineolatus Lineolate Blind Snake Trape & Mane 2017   Afrotyphlops punctatus Spotted Blind Snake Hahn 1980 Joger & Lambert 1996 Papenfuss 1969 Trape & Mane 2017 Wallach, Williams & Boundy 2014  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Boehme, Meinig & Roedel 1996 David & Ineich 1999 Joger & Lambert 1996 McDiarmid, Campbell & Toure 1999 Papenfuss 1969 Schatti 1986 Spawls & Branch 1995 Trape & Mane 2017 Genus Causus (Night Adders)   Causus maculatus West African Night Adder David & Ineich 1999 Joger & Lambert 1996 Schatti 1986 Spawls & Branch 1995 Trape & Mane 2017 Genus Cerastes (Horned Vipers)   Cerastes cerastes Sahara Horned Viper Berre 1989 Geniez 2018 Joger & Lambert 1996 McDiarmid, Campbell & Toure 1999 Papenfuss 1969 Sindaco, Venchi & Grieco 2013 Spawls & Branch 1995 Trape & Mane 2017   Cerastes vipera Sahara Sand Viper Berre 1989 Geniez 2018 Joger & Lambert 1996 McDiarmid, Campbell & Toure 1999 Papenfuss 1969 Spawls & Branch 1995 Trape & Mane 2017 Genus Echis (Carpet Vipers)   Echis jogeri Mali Carpet Viper David & Ineich 1999 Joger & Lambert 1996 McDiarmid, Campbell & Toure 1999 Trape & Mane 2017 Trape 2018   Echis leucogaster White-bellied Carpet Viper Berre 1989 Boehme & Heath 2018 David & Ineich 1999 Joger & Lambert 1996 McDiarmid, Campbell & Toure 1999 Pook, Joger, Stumpel & Wuster 2009 Sindaco, Venchi & Grieco 2013 Spawls & Branch 1995 Trape & Mane 2017   Echis ocellatus West African Carpet Viper David & Ineich 1999 Hughes 1976 Joger & Lambert 1996 Joger 1981 McDiarmid, Campbell & Toure 1999 Pook, Joger, Stumpel & Wuster 2009 Schatti 1986 Spawls & Branch 1995 Trape & Mane 2006 Trape & Mane 2017 Trape 2018

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa olivacea Sahelian Helmeted Turtle Boehme, Meinig & Roedel 1996 [as subrufa] Boycott & Bourquin 2008 [as subrufa] Fritz & Havas 2006 [as subrufa] Joger & Lambert 1996 [as subrufa] King & Burke 1989 [as subrufa] McCord, Joseph-Ouni & Bour 2004 [as subrufa] Papenfuss 1969 [as subrufa] Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Schatti 1986 [as subrufa] Trape, Trape & Chirio 2012 Genus Pelusios (African Mud Turtles)   Pelusios adansonii Sahelian Mud Turtle Bour 2008 Joger & Lambert 1996 King & Burke 1989 McCord, Joseph-Ouni & Bour 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Wermuth & Mertens 1977  Family Testudinidae (Tortoises) Genus Centrochelys (African Spurred Tortoise)   Centrochelys sulcata African Spurred Tortoise Berre 1989 Fritz & Havas 2006 Gasperetti, Stimson, Miller, Ross & Gasperetti 1993 Joger & Lambert 1996 King & Burke 1989 Papenfuss 1969 Petrozzi, Hema, Demaya, Benansio, Eniang, Diagne, Segniagbeto & Luiselli 2020 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Kinixys (Hinge-back Tortoises)   Kinixys nogueyi Western Savanna Hinge-back Tortoise Chirio & LeBreton 2007 Joger & Lambert 1996 King & Burke 1989 [as belliana] Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 [as belliana]  Family Trionychidae (Softshell Turtles) Genus Cyclanorbis (SubSaharan Flapshell Turtles)   Cyclanorbis senegalensis Senegal Flapshell Turtle Joger & Lambert 1996 Joger 1981 King & Burke 1989 McCord & Joseph-Ouni 2003 McGovern, Diagne, Diagne, Luiselli & Meylan 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Trape, Trape & Chirio 2012 Genus Trionyx (African Softshell Turtle)   Trionyx triunguis African Softshell Turtle Joger & Lambert 1996 Trape, Trape & Chirio 2012