Reptiles of Togo    Bibliography of the reptiles of Togo Taxonomic history of the reptiles of Togo Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Cynisca (West African Round-headed Worm Lizards)   Cynisca leucura White-tailed Worm Lizard Gans 1987 Gans 2005 Loveridge 1952 Mane & Trape 2015 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus suchus West African Crocodile Fuchs, Mertens & Wermuth 1974 King & Burke 1989 [as niloticus] Trape, Trape & Chirio 2012 Wermuth & Mertens 1977 Genus Mecistops (African Slender-snouted Crocodiles)   Mecistops cataphractus West African Slender-snouted Crocodile King & Burke 1989 Shirley, Carr, Nestler, Vliet & Brochu 2018 Trape, Trape & Chirio 2012 Genus Osteolaemus (Dwarf Crocodiles)   Osteolaemus tetraspis West African Dwarf Crocodile Chirio & LeBreton 2007 Trape, Trape & Chirio 2012

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama agama Common Rainbow Agama Hagedoorn 2005 Leache, Grummer, Miller, Krishnan, Fujita, Böhme, Schmitz, LeBreton, Ineich, Chirio, Ofori-boateng, Eniang, Greenbaum, Rödel & Wagner 2016 [as A. picticauda] Loveridge 1952 Mediannikov, Trape & Trape 2012 Segniagbeto, Trape, Afiademanyo, Rödel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Agama gracilimembris Lesser Ground Agama Segniagbeto, Trape, Afiademanyo, Rödel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Agama parafricana Lesser African Forest Agama Leache, Grummer, Miller, Krishnan, Fujita, Böhme, Schmitz, LeBreton, Ineich, Chirio, Ofori-boateng, Eniang, Greenbaum, Rödel & Wagner 2016 Mediannikov, Trape & Trape 2012 Segniagbeto, Trape, Afiademanyo, Rödel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Agama sankaranica Sankaranian Agama Chirio & LeBreton 2007 Mediannikov, Trape & Trape 2012 Segniagbeto, Trape, Afiademanyo, Rödel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo gracilis Graceful Chameleon Glaw 2015 Loveridge 1952 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Tilbury 2010 Trape, Trape & Chirio 2012   Chamaeleo necasi Togo Chameleon Glaw 2015 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Tilbury 2010 Trape, Trape & Chirio 2012 Ullenbruch, Krause & Boehme 2007   Chamaeleo senegalensis Senegal Chameleon Glaw 2015 Grandison 1956 Loveridge 1952 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Tilbury 2010 Trape, Trape & Chirio 2012  Family Eublepharidae (Eyelid Geckos) Genus Hemitheconyx (African Fat-tailed Geckos)   Hemitheconyx caudicinctus West African Fat-tailed Gecko Chirio & LeBreton 2007 Grandison 1956 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Seufer, Kaverkin & Kirschner (eds.) 2005 Trape, Trape & Chirio 2012  Family Gekkonidae (Typical Geckos) Genus Ancylodactylus (African Rock and Forest Geckos)   Ancylodactylus spinicollis Spiny-necked Rock Gecko Perret 1986 Segniagbeto 2021 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus albituberculatus White-tubercled Half-toed Gecko Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Hemidactylus angulatus West African House Gecko Chirio & LeBreton 2007 Loveridge 1947 Loveridge 1952 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Hemidactylus fasciatus African Banded Forest Gecko Joger 1981 Loveridge 1947 Loveridge 1952 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Hemidactylus kyaboboensis Kyabobo Half-toed Gecko Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Wagner, Leache & Fujita 2014   Hemidactylus mabouia Cosmopolitan House Gecko Loveridge 1947 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Hemidactylus matschiei Adele Half-toed Gecko Loveridge 1947 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012 Wermuth 1965   Hemidactylus muriceus Prickly Half-toed Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 Henle & Boehme 2003 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012 Wermuth 1965 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus conraui Cameroon Dwarf Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Lygodactylus gutturalis Chevron-throated Dwarf Gecko Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015  Family Gerrhosauridae (Plated Lizards) Genus Broadleysaurus (Rough-scaled Plated Lizard)   Broadleysaurus major Rough-scaled Plated Lizard Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012  Family Lacertidae (Typical Lizards) Genus Acanthodactylus (Fringe-toed Lizards)   Acanthodactylus boueti Benin Fringe-toed Lizard Loveridge 1952 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012 Genus Heliobolus (Bushveld Lizards)   Heliobolus nitida Red-tailed Bushveld Lizard Chirio & LeBreton 2007 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Szczerbak 1975 Trape, Trape & Chirio 2012 Genus Holaspis (Blue-tailed Gliding Lizards)   Holaspis guentheri Western Blue-tailed Gliding Lizard Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012  Family Phyllodactylidae (Leaf-toed Geckos) Genus Ptyodactylus (Fan-toed Geckos)   Ptyodactylus togoensis Sahelian Fan-toed Gecko Bauer, Tchibozo, Pauwels & Lenglet 2006 [as P. ragazzii] Heimes 1987 [as P. ragazzii] Loveridge 1947 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 [as P. ragazzii] Sindaco & Jeremchenko 2008 [as P. ragazzii] Trape, Trape & Chirio 2012 [as P. ragazzii] Genus Tarentola (Wall Geckos)   Tarentola ephippiata African Wall Gecko Joger 1984 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Sindaco & Jeremchenko 2008  Family Scincidae (Skinks) Genus Chalcides (Cylindrical Skinks)   Chalcides thierryi Togo Cylindrical Skink Greenbaum 2005 Papenfuss 1969 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012 Genus Cophoscincopus (West African Keeled Water Skinks)   Cophoscincopus greeri Guinean Keeled Water Skink Boehme, Schmitz & Ziegler 2000 Trape, Trape & Chirio 2012   Cophoscincopus simulans Ghanese Keeled Water Skink Boehme, Schmitz & Ziegler 2000 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012 Genus Mochlus (African Supple Skinks)   Mochlus fernandi Western Red-flanked Skink Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015   Mochlus guineensis Guinean Supple Skink Loveridge 1952 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012 Genus Panaspis (African Snake-eyed Skinks)   Panaspis togoensis Togo Snake-eyed Skink Joger 1981 Loveridge 1952 Perret 1973 Schmitz, Ineich & Chirio 2005 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis affinis Lesser Brown-flanked Skink Hagedoorn 2005 Hoogmoed 1974 Joger 1981 Loveridge 1952 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Trachylepis aureogularis Orange-throated Skink Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015   Trachylepis buettneri Togo Savanna Skink Hoogmoed 1974 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Trachylepis maculilabris Common Speckled-lipped Skink Grandison 1956 Loveridge 1952 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Trachylepis perrotetii Orange-flanked Skink Hagedoorn 2005 Joger 1981 Loveridge 1952 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Trachylepis polytropis Greater Forest Skink Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015   Trachylepis quinquetaeniata African Five-lined Skink Joger 1981 Koppetsch 2020 Loveridge 1952 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus exanthematicus Western Savanna Monitor Bayless 2002 Bennett in Pianka, King & King (eds.) 2004 Grandison 1956 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Varanus niloticus Nile Monitor Bayless 2002 Chirio & LeBreton 2007 Lenz in Pianka, King & King (eds.) 2004 Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 Trape, Trape & Chirio 2012   Varanus stellatus African Forest Monitor Bayless 2002 [as V. ornatus] Segniagbeto, Trape, Afiademanyo, Roedel, Ohler, Dubois, David, Meirte, Glitho, Petrozzi & Luiselli 2015 [as V. ornatus] Trape, Trape & Chirio 2012 [as V. ornatus]

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Calabaria (Calabar Boa)   Calabaria reinhardtii Calabar Boa Chirio & LeBreton 2007 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Eryx (Sand Boas)   Eryx muelleri West African Sand Boa Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 McDiarmid, Campbell & Toure 1999 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014  Family Colubridae (Typical Snakes or Colubrids) Genus Afronatrix (West African Brown Water Snake)   Afronatrix anoscopus West African Brown Water Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Bamanophis (West African Whip Snake)   Bamanophis dorri West African Whip Snake Hulselmans & Verheyen 1970 Schatti & Trape 2008 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hippocrepis West African Herald Snake Rasmussen, Chirio & Ineich 2000 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Crotaphopeltis hotamboeia Common Herald Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Dasypeltis (African Egg Eaters)   Dasypeltis confusa Diamondback Egg Eater Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Trape, Mediannikov & Trape 2012   Dasypeltis fasciata Central African Egg Eater Bates & Broadley 2018 Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Trape, Mediannikov & Trape 2012   Dasypeltis gansi Savanna Egg Eater Bates, Trape & Spawls 2013 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Trape & Mane 2006 Trape, Mediannikov & Trape 2012   Dasypeltis parascabra Guinean Egg Eater Trape & Balde 2014 Trape, Mediannikov & Trape 2012 Genus Dipsadoboa (Nocturnal Tree Snakes)   Dipsadoboa underwoodi Cameroon Nocturnal Tree Snake Rasmussen 1993 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Dipsadoboa unicolor Unicoloured Nocturnal Tree Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Dipsadoboa viridis Green Nocturnal Tree Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Dipsadoboa weileri Grey Nocturnal Tree Snake Rasmussen 1993 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 [as "cf. weileri"] Genus Dispholidus (Boomslangs)   Dispholidus sp. (Boomslang) David & Ineich 1999 [as D. typus] Hulselmans & Verheyen 1970 [as D. typus] Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 [as D. typus] Spawls & Branch 1995 [as D. typus] Trape & Balde 2014 [as D. typus] Genus Grayia (Equatorial African Water Snakes)   Grayia smythii Equatorial Water Snake Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Hapsidophrys (Emerald Snakes)   Hapsidophrys lineatus Black-lined Green Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Hapsidophrys smaragdinus Common Emerald Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Meizodon (African Smooth Snakes)   Meizodon coronatus West African Crowned Smooth Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Meizodon regularis East African Crowned Smooth Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Natriciteres (African Marsh Snakes)   Natriciteres fuliginoides Collared Marsh Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Natriciteres olivacea Olive Marsh Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Natriciteres variegata Variable Marsh Snake Hulselmans & Verheyen 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Philothamnus (African Green Snakes)   Philothamnus carinatus Barred Green Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Philothamnus heterodermus Emerald Green Snake Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Trape, Hughes, Collet & Mediannikov 2021   Philothamnus irregularis Irregular Green Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Philothamnus nitidus Cameroon Green Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Philothamnus semivariegatus Variegated Bush Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011   Philothamnus smithii Smith's Bush Snake Trape & Balde 2014 Trape, Hughes, Collet & Mediannikov 2021 Genus Rhamnophis (Dagger-tooth Tree Snakes)   Rhamnophis aethiopissa Large-eyed Green Tree Snake Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Scaphiophis (African Hook-nosed Snakes)   Scaphiophis albopunctatus Common African Hook-nosed Snake Broadley 1994 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus variegatus Variable Cat Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Thelotornis (African Vine Snakes)   Thelotornis kirtlandii Northern Vine Snake Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014 Genus Thrasops (Black Tree Snakes)   Thrasops occidentalis Western Black Tree Snake Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Trape & Mane 2006 Genus Toxicodryas (African Tree Cat Snakes)   Toxicodryas blandingii Western Black-and-Yellow Tree Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Toxicodryas pulverulenta Western Powdered Cat Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Dendroaspis (Mambas)   Dendroaspis jamesonii Central African Green Mamba Chippaux & Jackson 2019 David & Ineich 1999 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014   Dendroaspis viridis Western Green Mamba Chippaux & Jackson 2019 David & Ineich 1999 Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014 Genus Elapsoidea (African Garter Snakes)   Elapsoidea semiannulata Half-banded Garter Snake Broadley 1971 Broadley 1998 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014 Genus Naja (Typical Cobras)   Naja guineensis Black Forest Cobra Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018   Naja katiensis Brown Spitting Cobra David & Ineich 1999 Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Naja nigricollis Black-necked Spitting Cobra Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Wuster, Crookes, Ineich, Mane, Pook, Trape & Broadley 2007   Naja savannula West African Banded Cobra Wuster, Chirio, Trape, Ineich, Jackson, Greenbaum, Barron, Kusamba, Nagy, Storey, Hall, Wuster, Barlow & Broadley 2018   Naja senegalensis Senegalese Cobra Spawls & Branch 1995 [as N. haje] Genus Pseudohaje (Tree Cobras)   Pseudohaje nigra Black Tree Cobra David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014 Villiers 1975  Family Lamprophiidae (House Snakes, etc.) Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas unicolor Equatorial Purple-glossed Snake Broadley 1971 Hulselmans & Verheyen 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Aparallactus (African Centipede Eaters)   Aparallactus lunulatus Reticulated Centipede Eater Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Villiers 1975 Wallach, Williams & Boundy 2014   Aparallactus modestus Western Forest Centipede Eater Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Atractaspis (Stiletto Snakes)   Atractaspis aterrima Western Forest Stiletto Snake David & Ineich 1999 Hulselmans & Verheyen 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Atractaspis dahomeyensis Dahomey Stiletto Snake David & Ineich 1999 Hulselmans, Roo & Vree 1970 Laurent 1950 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Atractaspis irregularis Variable Stiletto Snake David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Laurent 1950 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon fuliginosus Common African House Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Trape & Mediannikov 2016   Boaedon lineatus Lined House Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Trape & Mediannikov 2016   Boaedon olivaceus Olive House Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Boaedon virgatus West African House Snake Hulselmans & Verheyen 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Chamaelycus (African Banded Snakes)   Chamaelycus fasciatus Ituri Banded Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Dromophis (Olympic Snakes)   Dromophis lineatus Lined Olympic Snake Hughes 2004 Hulselmans & Verheyen 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Dromophis praeornatus Ornate Olympic Snake Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Gonionotophis (Lesser File Snakes)   Gonionotophis grantii Savanna Dwarf File Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Gonionotophis klingi Western Forest Dwarf File Snake Chippaux & Jackson 2019 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Villiers 1975 Genus Hemirhagerrhis (Bark Snakes)   Hemirhagerrhis nototaenia Southeastern Bark Snake Broadley & Hughes 2000 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Hormonotus (Yellow Forest Snake)   Hormonotus modestus Yellow Forest Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Kladirostratus (Striped Beaked Snakes)   Kladirostratus togoensis Western Striped Beaked Snake Chippaux & Jackson 2019 Chirio & Ineich 1991 Keates, Conradie, Greenbaum & Edwards 2019 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Limaformosa (Venus File Snakes)   Limaformosa crossi Savanna File Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Villiers 1975   Limaformosa guirali White-speckled Forest File Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Lycophidion (African Wolf Snakes)   Lycophidion irroratum Western Forest Wolf Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Laurent 1968 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Lycophidion laterale Masked Wolf Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Lycophidion nigromaculatum Black-spotted Wolf Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Lycophidion semicinctum Semi-annulated Wolf Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Mehelya (Greater File Snakes)   Mehelya poensis Western Forest File Snake Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Mehelya stenophthalmus Small-eyed File Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Polemon (African Snake Eaters)   Polemon acanthias Five-striped Snake Eater Hulselmans & Verheyen 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Polemon neuwiedi West African Snake Eater Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Villiers 1975 Genus Prosymna (Shovel-snout Snakes)   Prosymna collaris Collared Shovel-snout Chippaux & Jackson 2019 Chirio, Ineich, Schmitz & Trape 2011 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 [as P. greigerti]   Prosymna greigerti Sudan Savanna Shovel-snout Chirio, Ineich, Schmitz & Trape 2011   Prosymna meleagris Speckled Shovel-snout Broadley 1980 Chirio, Ineich, Schmitz & Trape 2011 Hulselmans & Verheyen 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis afroccidentalis West African Sand Snake Hagedoorn 2005 [as P. sibilans] Hulselmans & Verheyen 1970 [as P. sibilans] Hulselmans, Vree & Straeten 1971 [as P. sibilans] Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 [as P. sibilans] Trape & Balde 2014 [as P. sibilans] Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019   Psammophis elegans Elegant Sand Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Psammophis phillipsii West African Olive Grass Snake Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Trape, Crochet, Broadley, Sourouille, Mane, Burger, Boehme, Saleh, Karan, Lanza & Mediannikov 2019 Genus Rhamphiophis (Beaked Snakes)   Rhamphiophis oxyrhynchus Western Beaked Snake Chirio & Ineich 1991 Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014  Family Leptotyphlopidae (Thread Snakes) Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis narirostris Sahel Thread Snake Chirio 2012 Hahn 1980 McDiarmid, Campbell & Toure 1999 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Genus Tricheilostoma (West African Forest Thread Snakes)   Tricheilostoma bicolor Bicoloured Thread Snake Adalsteinsson, Branch, Trape, Vitt & Hedges 2009 Hahn & Wallach 1998 Hahn 1980 Hulselmans & Verheyen 1970 McDiarmid, Campbell & Toure 1999 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Tricheilostoma sundewalli Central African Thread Snake Hahn 1980 McDiarmid, Campbell & Toure 1999 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python regius Ball Python Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Joger 1981 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Python sebae Northern Rock Python Broadley 1984 Hulselmans, Roo & Vree 1970 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops lineolatus Lineolate Blind Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Afrotyphlops punctatus Spotted Blind Snake Hahn 1980 Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Wallach, Williams & Boundy 2014 Genus Letheobia (African Gracile Blind Snakes)   Letheobia crossii Nigerian Gracile Blind Snake Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Trape & Segniagbeto 2008  Family Viperidae (Vipers) Genus Atheris (African Bush Vipers)   Atheris chlorechis West African Green Bush Viper Chippaux & Jackson 2019 David & Ineich 1999 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 McDiarmid, Campbell & Toure 1999 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Wallach, Williams & Boundy 2014   Atheris squamigera Southern Green Bush Viper Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014 Villiers 1975 Wallach, Williams & Boundy 2014 Genus Bitis (African Adders)   Bitis arietans Common Puff Adder David & Ineich 1999 Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 McDiarmid, Campbell & Toure 1999 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014   Bitis nasicornis African Nose-horned Viper Chirio & LeBreton 2007 David & Ineich 1999 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Trape & Balde 2014   Bitis rhinoceros Rhinoceros Viper Chippaux & Jackson 2019 Hulselmans & Verheyen 1970 [as B. gabonica] Hulselmans, Roo & Vree 1970 [as B. gabonica] Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014 Genus Causus (Night Adders)   Causus maculatus West African Night Adder David & Ineich 1999 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014   Causus rhombeatus Rhombic Night Adder Hulselmans & Verheyen 1970 Hulselmans, Roo & Vree 1970 Hulselmans, Vree & Straeten 1971 Genus Echis (Carpet Vipers)   Echis ocellatus West African Carpet Viper David & Ineich 1999 Hughes 1976 Hulselmans, Vree & Straeten 1971 McDiarmid, Campbell & Toure 1999 Pook, Joger, Stumpel & Wuster 2009 Segniagbeto, Trape, David, Ohler, Dubois & Glitho 2011 Spawls & Branch 1995 Trape & Balde 2014 Trape 2018

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Assou, Segniagbeto, Radji, Akiti & Pando 2018 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Okangny, Dendi, Fretey & Luiselli 2017 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Assou, Segniagbeto, Radji, Akiti & Pando 2018 Brongersma in Bjorndal (ed.) 1982 King & Burke 1989 Mettler, Clyde-Brockway, Honarvar & Paladino 2019 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Bour, Ohler, Dubois, Roedel, Trape, Fretey, Petrozzi & Luiselli 2014 Segniagbeto, Okangny, Dendi, Fretey & Luiselli 2017 Trape, Trape & Chirio 2012 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Assou, Segniagbeto, Radji, Akiti & Pando 2018 Brongersma in Bjorndal (ed.) 1982 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Bour, Ohler, Dubois, Roedel, Trape, Fretey, Petrozzi & Luiselli 2014 Segniagbeto, Okangny, Dendi, Fretey & Luiselli 2017 Trape, Trape & Chirio 2012 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Assou, Segniagbeto, Radji, Akiti & Pando 2018 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Bour, Ohler, Dubois, Roedel, Trape, Fretey, Petrozzi & Luiselli 2014 Segniagbeto, Okangny, Dendi, Fretey & Luiselli 2017 Trape, Trape & Chirio 2012  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Assou, Segniagbeto, Radji, Akiti & Pando 2018 Brongersma in Bjorndal (ed.) 1982 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Bour, Ohler, Dubois, Roedel, Trape, Fretey, Petrozzi & Luiselli 2014 Segniagbeto, Okangny, Dendi, Fretey & Luiselli 2017 Trape, Trape & Chirio 2012  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa olivacea Sahelian Helmeted Turtle Boycott & Bourquin 2008 [as subrufa] King & Burke 1989 [as subrufa] Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Bour, Ohler, Dubois, Roedel, Trape, Fretey, Petrozzi & Luiselli 2014 [as subrufa] Trape, Trape & Chirio 2012   Pelomedusa variabilis West African Helmeted Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelusios (African Mud Turtles)   Pelusios castaneus Chestnut Mud Turtle Bour, Luiselli, Petrozzi, Segniabeto & Chirio 2016 Chirio & LeBreton 2007 King & Burke 1989 Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Bour, Ohler, Dubois, Roedel, Trape, Fretey, Petrozzi & Luiselli 2014 Trape, Trape & Chirio 2012   Pelusios gabonensis Forest Mud Turtle Chirio & LeBreton 2007 King & Burke 1989  Family Testudinidae (Tortoises) Genus Kinixys (Hinge-back Tortoises)   Kinixys erosa Forest Hinge-back Tortoise Chirio & LeBreton 2007 Luiselli & Diagne 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Bour, Ohler, Dubois, Roedel, Trape, Fretey, Petrozzi & Luiselli 2014 Trape, Trape & Chirio 2012   Kinixys homeana West African Hinge-back Tortoise Chirio & LeBreton 2007 Luiselli & Diagne 2013 Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Bour, Ohler, Dubois, Roedel, Trape, Fretey, Petrozzi & Luiselli 2014 Trape, Trape & Chirio 2012   Kinixys nogueyi Western Savanna Hinge-back Tortoise King & Burke 1989 [as belliana] Loveridge 1952 Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Bour, Ohler, Dubois, Roedel, Trape, Fretey, Petrozzi & Luiselli 2014 Trape, Trape & Chirio 2012 [as belliana]  Family Trionychidae (Softshell Turtles) Genus Cyclanorbis (SubSaharan Flapshell Turtles)   Cyclanorbis elegans Nubian Flapshell Turtle Baker, Diagne & Luiselli 2015 King & Burke 1989 McCord & Joseph-Ouni 2003 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Bour, Ohler, Dubois, Roedel, Trape, Fretey, Petrozzi & Luiselli 2014 Trape, Trape & Chirio 2012 Wermuth & Mertens 1977   Cyclanorbis senegalensis Senegal Flapshell Turtle King & Burke 1989 Maran 2009 McCord & Joseph-Ouni 2003 McGovern, Diagne, Diagne, Luiselli & Meylan 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Bour, Ohler, Dubois, Roedel, Trape, Fretey, Petrozzi & Luiselli 2014 Trape, Trape & Chirio 2012 Genus Trionyx (African Softshell Turtle)   Trionyx triunguis African Softshell Turtle Bonin, Devaux & Dupre 2006 King & Burke 1989 Maran 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Segniagbeto, Bour, Ohler, Dubois, Roedel, Trape, Fretey, Petrozzi & Luiselli 2014