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Taxonomy of the family Phrynosomatidae
Bibliography of the genus Sceloporus
Biodiversity of the family Phrynosomatidae








Genus
Sceloporus

Northern Spiny Lizards, Fence Lizards, Swift Lizards, Swifts

Stachelleguane, Stachelschuppenleguane

Tornleguaner

1817 Tropidolepis Cuvier (type species: Stellio undulatus Bosc & Daudin in Sonnini & Latreille 1801; syn. Smith & Taylor 1950)
1828 Sceloporus Wiegmann (type species: Sceloporus torquatus Wiegmann 1828)
1832 Lopherpes Rafinesque (type species: Lopherpes dicyanelus Rafinesque 1832; syn. Schmidt 1953)
1888 Lysoptychus Cope (type species: Lysoptychus lateralis Cope 1888; syn. Cole 1978)
1919 Sator Dickerson (type species: Sator grandaevus Dickerson 1919; syn. Wiens & Reeder 1997)
Contents: 111 species, of which 77 (69.4%) are endemic.
Remarks: The nominal genus Sator has been regarded as a synonym of Sceloporus by several authors (e.g., Wyles & Gorman 1978; Queiroz 1982; Etheridge & Queiroz 1988; Frost & Etheridge 1989; Wiens & Reeder 1997), however, several others have retained it as a valid genus (Wiens 1993a; Schulte, Macey, Larson & Papenfuss 1998). Larsen & Tanner (1975) recognized Lysoptychus as a valid genus, but this was rejected by Cole (1978).
Distribution: USA, Mexico, Central America.
Reported from: Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Aguascalientes, Baja California Norte [incl. Isla Cedros], Baja California Sur [incl. Isla Cerralvo, Isla Santa Catalina], Campeche, Chiapas, Chihuahua, Coahuila, Colima, Distrito Federal, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico, Michoacan, Morelos, Nayarit, Nuevo Leon, Oaxaca, Puebla, Queretaro, Quintana Roo [incl. Cozumel], San Luis Potosi, Sinaloa, Sonora [incl. Isla San Pedro Nolasco], Tabasco, Tamaulipas, Tlaxcala, Veracruz, Yucatan, Zacatecas), Nicaragua, Panama, USA (Alabama, Arizona, Arkansas, California [incl. Channel Islands], Colorado, Delaware, Florida, Georgia, Idaho, Illinois, Indiana, Kansas, Kentucky, Louisiana, Maryland, Mississippi, Missouri, Montana, Nebraska, Nevada, New Jersey, New Mexico, New York, North Carolina, North Dakota, Ohio, Oklahoma, Oregon, Pennsylvania, South Carolina, South Dakota, Tennessee, Texas, Utah, Virginia, Washington, Washington, D.C., West Virginia, Wyoming).

Sceloporus acanthinus

Guatemalan Rough Spiny Lizard, (Bocourt's Spiny Lizard)

Guatemalischer Rauhschuppiger Stachelleguan

Guatemalansk Ruskællet Tornleguan

1873 Sceloporus acanthinus Bocourt
Sceloporus malachiticus acanthinus Smith 1942
1918 Sceloporus guentheri Stejneger (Peters & Donoso-Barros 1970)

Remarks: Revalidated as a separate species by Stuart (1971). Records from El Salvador (e.g., Peters & Donoso-Barros 1970; Smith, Chiszar & Humphrey 2001) have been regarded as doubtful (Köhler & Heimes 2002). Köhler, Veselý & Greenbaum (2006) treated all records of acanthinus from El Salvador as malachiticus, apparently followed by Greenbaum & Komar (2010).
Distribution: Guatemala, Mexico (Chiapas).


Sceloporus adleri

Asoleadero Spiny Lizard, (Adler's Spiny Lizard)

Asoleadero-Stachelleguan

Asoleadero-tornleguan

1974 Sceloporus adleri Smith

Distribution: Mexico (Guerrero).


Sceloporus aeneus

Southern Bunchgrass Lizard, Black-bellied Bunchgrass Lizard

Südlicher Gras-stachelleguan

Sydlig Græstornleguan

1828 Sceloporus aeneus Wiegmann

Remarks: Previously included bicanthalis and subniger.
Distribution: Mexico (Distrito Federal, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico, Michoacan, Morelos, Puebla, Queretaro, Tlaxcala, Veracruz).


Sceloporus albiventris

Western White-bellied Spiny Lizard, White-bellied Spiny Lizard, White-bellied Rough Spiny Lizard

Westlicher Weißbauch-Stachelleguan

Vestlig Hvidbuget Tornleguan

1939 Sceloporus horridus albiventris Smith
Sceloporus albiventris Lemos-Espinal, Smith & Chiszar 2006

Distribution: Mexico (Chihuahua, Durango, Jalisco, Nayarit, Sinaloa, Sonora).


Sceloporus anahuacus

Anahuacan Bunchgrass Lizard, Anahuacan Graphic Lizard

Anahuaca-Gras-stachelleguan

Anahuaca-græstornleguan

1983 Sceloporus anahuacus Lara-Gongora

Remarks: Sites, Camarillo, Gonzalez, Mendoza, Javier, Manzilla & Lara-Gongora (1988) questioned the validity of anahuacus, however Flores-Villela (1993) noted that the species is valid, referring to as yet unpublished material.
Distribution: Mexico (Distrito Federal, Mexico).


Sceloporus angustus

Santa Cruz Island Spiny Lizard, (Santa Cruz Island Sator)

Santa Cruz-Stachelleguan

Santa Cruz-tornleguan

1919 Sator angustus Dickerson
Sceloporus angustus Wiens & Reeder 1997

Distribution: Mexico (Baja California Sur).


Sceloporus arenicolus

Dunes Sagebrush Lizard, Sand Dune Lizard

Sand-Stachelleguan

Sand-tornleguan

1972 Sceloporus graciosus arenicolus Degenhardt & Jones
Sceloporus arenicolous Collins 1991

Remarks: The phylogenetic analysis of Wiens & Reeder (1997) supported the specific recognition of arenicolous suggested by Collins (1991).
Distribution: USA (New Mexico, Texas).


Sceloporus asper

Asperous Spiny Lizard, (Boulenger's Scaly Lizard)

Rauhschuppiger Stachelleguan

Ru Tornleguan

1897 Sceloporus asper Boulenger
1897 Sceloporus obscurus Van Denburgh (Smith & Taylor 1950)

Distribution: Mexico (Colima, Guerrero, Jalisco, Michoacan, Nayarit).


Sceloporus aurantius

Southern Occidental Bunchgrass Lizard

Orangestreifen-Stachelleguan

Orangestribet Tornleguan

2014 Sceloporus aurantius Grummer & Bryson

Distribution: Mexico (Aguascalientes, Jalisco, Zacatecas).


Sceloporus aureolus

Eastern Crevice Spiny Lizard, Eastern Cleft Spiny Lizard

Östlicher Spalten-Stachelleguan

Østlig Spaltetornleguan

1936 Sceloporus mucronatus aureolus Smith
Sceloporus aureolus Martinez-Mendez & Mendez de la Cruz 2007

Distribution: Mexico (Oaxaca, Puebla, Veracruz).


Sceloporus becki

Island Fence Lizard

Channel Islands-Zaunleguan

Channel Islands-tornleguan

1905 Sceloporus becki Van Denburgh
Sceloporus occidentalis becki Grinnell & Camp 1917

Remarks: Revalidated as a separate species by Bell (2001). Wiens & Reeder (1997) had already suggested that becki deserves species status.
Distribution: USA (California: Channel Islands).


Sceloporus bicanthalis

Transvolcanic Bunchgrass Lizard

Transvulkanischer Stachelleguan

Transvulkansk Tornleguan

1937 Sceloporus aeneus bicanthalis Smith
Sceloporus bicanthalis Guillette & Smith 1985

Distribution: Mexico (Distrito Federal, Hidalgo, Mexico, Oaxaca, Puebla, Tlaxcala, Veracruz).


Sceloporus bimaculosus

Twin-spotted Spiny Lizard, (Desert Spiny Lizard)

Zweiflecken-Stachelleguan

Dobbeltplettet Tornleguan

1955 Sceloporus magister bimaculosus Phelan & Brattstrom
Sceloporus bimaculosus Schulte, Macey & Papenfuss 2006

Remarks: Previously regarded as a subspecies of magister. Recognized as a separate species by Schulte, Macey & Papenfuss (2006), followed by others (e.g., Enderson & Bezy 2010), although Leaché & Mulcahy (2007) presented evidence in support of treating the two taxa as conspecific, followed by some authors (e.g., Wilson, Mata-Silva & Johnson 2013).
Distribution: Mexico (Chihuahua, Coahuila, Durango), USA (Arizona, Colorado, New Mexico, Texas).


Sceloporus brownorum

Durango Bunchgrass Lizard, (Brown's Bunchgrass Lizard)

Durango-Gras-stachelleguan

Durango-græstornleguan

1997 Sceloporus scalaris brownorum Smith, Watkins-Colwell, Lemos-Espinal & Chiszar
Sceloporus brownorum Grummer & Bryson 2014

Distribution: Mexico (Aguascalientes, Durango, Jalisco, Nayarit, Zacatecas).


Sceloporus bulleri

Jaliscan Spiny Lizard, (Buller's Scaly Lizard)

Jalisco-Stachelleguan

Jalisco-tornleguan

1894 Sceloporus bulleri Boulenger

Remarks: Records from Michoacan (e.g., Duellman 1965) and adjacent Jalisco refer to insignis (Webb 1967), however, it was still listed for Michoacan by Alvarado-Díaz, Suazo-Ortuño, Wilson & Medina-Aguilar (2013).
Distribution: Mexico (Colima, Durango, Jalisco, Sinaloa).


Sceloporus caeruleus

Blue Ornate Spiny Lizard

Blaue Pracht-Stachelleguan

Blå Pragttornleguan

1936 Sceloporus ornatus caeruleus Smith
Sceloporus caeruleus Martinez-Mendez & Mendez de la Cruz 2007

Remarks: Sometimes regarded as a subspecies of ornatus (e.g., Lemos-Espinal & Smith 2007).
Distribution: Mexico (Coahuila).


Sceloporus carinatus

Keeled Spiny Lizard

Gekielter Stachelleguan

Kølskællet Tornleguan

1936 Sceloporus carinatus Smith

Distribution: Guatemala, Mexico (Chiapas, Oaxaca).


Sceloporus cautus

Shy Spiny Lizard, Wary Spiny Lizard

Scheuer Stachelleguan

Sky Tornleguan

1938 Sceloporus cautus Smith

Distribution: Mexico (Coahuila, Nuevo Leon, San Luis Potosi, Tamaulipas, Zacatecas).


Sceloporus chaneyi

Peña Nevada Spiny Lizard, (Chaney's Spiny Lizard, Chaney's Bunchgrass Lizard)

Peña Nevada-Stachelleguan

Peña Nevada-tornleguan

1992 Sceloporus chaneyi Liner & Dixon

Distribution: Mexico (Nuevo Leon, Tamaulipas).


Sceloporus chrysostictus

Yellow-spotted Spiny Lizard, Yucatan Spiny Lizard

Gelbfleckiger Stachelleguan

Gulplettet Tornleguan

1867 Sceloporus chrysostictus Cope

Distribution: Belize, Guatemala, Mexico (Campeche, Quintana Roo, Yucatan).


Sceloporus clarkii

Southwestern Spiny Lizard, (Clark's Spiny Lizard)

Clarks Stachelleguan

Sydvestlig Tornleguan, (Clarks Tornleguan)

1852 Sceloporus clarkii Baird & Girard
1893 Sceloporus boulengeri Stejneger (Smith & Taylor 1950)
Sceloporus clarkii boulengeri Burt 1935
1954 Sceloporus clarki vallaris Shannon & Urbana
1959 Sceloporus clarkii uriquensis Tanner & Robison

Other common names:
boulengeri: Southern Clark's Spiny Lizard, Stejneger's Spiny Lizard
clarki: Sonoran Spiny Lizard
uriquensis: El Fuerte Basin Spiny Lizard
vallaris: Plateau Spiny Lizard
Distribution: Mexico (Aguascalientes, Chihuahua, Durango, Jalisco, Nayarit, Sinaloa, Sonora [incl. Isla San Pedro Nolasco], Zacatecas), USA (Arizona, New Mexico).


Sceloporus consobrinus

Prairie Lizard

Prärie-Stachelleguan

Prærie-tornleguan

1853 Sceloporus consobrinus Baird & Girard
Sceloporus undulatus consobrinus Cope 1900
1882 Sceloporus garmani Boulenger (Leaché & Reeder 2002)
Sceloporus undulatus garmani Smith 1938
1992 Sceloporus undulatus tedbrowni Smith, Bell, Applegarth & Chiszar (Leaché & Reeder 2002)

Other common names:
consobrinus: Southern Prairie Lizard
garmani: Northern Prairie Lizard
tedbrowni: Mescalero Sand Dunes Prairie Lizard
Remarks: Now restricted to USA, Mexican records of consobrinus refer to other species (Leaché 2009; Nevárez-de los Reyes, Lazcano, García-Padilla, Mata-Silva, Johnson & Wilson 2016).
This species corresponds to part of the "central clade" of Leaché & Reeder (2002), which contains populations previously included in the subspecies consobrinus, erythrocheilus, garmani, hyacinthinus, tedbrowni, tristichus, and undulatus of the formerly more wide-ranging species undulatus. As with other members of the undulatus complex, the population boundaries of this species is currently defined by genetic data, which do not always agree with traditional concepts of taxonomy based on morphological or ecological variation (Jones & Lovich [eds.] 2009). Genetic research is still needed to clarify the exact limits of distribution of this species.
Records from Nuevo Leon refer to cowlesi (Nevárez-de los Reyes, Lazcano, García-Padilla, Mata-Silva, Johnson & Wilson 2016). Records from Durango and Zacatecas refer to edbelli, according to Lemos-Espinal, Smith, Ballinger, Smith & Chiszar (1998, as belli), however, the status of these records is uncertain. Nevertheless, Valdez-Lares, Muñiz-Martínez, Gadsden, Aguirre-León, Castañeda-Gaytán & González-Trápaga (2013) provisionally retained consobrinus as part of the Durango fauna, pending further studies.
Populations east of the Mississippi River along the Gulf Coast may represent a separate species (Leaché & Reeder 2002; Crother, Boundy & al. 2003).
Distribution: USA (Arkansas, Colorado, Illinois, Kansas, Louisiana, Mississippi, Missouri, Nebraska, New Mexico, Oklahoma, South Dakota, Texas, Wyoming).


Sceloporus couchii

Santa Catarina Spiny Lizard, (Couch's Spiny Lizard)

Santa Catarina-Stachelleguan

Santa Catarina-tornleguan

1858 Sceloporus couchii Baird
Lysoptychus couchi Larsen & Tanner 1974
1888 Lysoptychus lateralis Cope (Smith & Taylor 1950)

Distribution: Mexico (Coahuila, Nuevo Leon).


Sceloporus cowlesi

Southwestern Fence Lizard

Südwestlicher Zaunleguan

Sydvestlig Hegnsleguan

1956 Sceloporus undulatus cowlesi Lowe & Norris
Sceloporus cowlesi Leaché & Reeder 2002
1995 Sceloporus undulatus speari Smith, Chiszar, Lemos-Espinal & Bell (Leaché & Reeder 2002)

Other common names:
cowlesi: White Sands Prairie Lizard
speari: Cabeza de Vaca Prairie Lizard, (Spear's Prairie Lizard)
Remarks: This species corresponds to Leaché's & Reeder's (2002) "southwestern clade", which encompasses populations previously regarded as the subspecies consobrinus, cowlesi, speari, and tristichus of the wide-ranging undulatus. As with other members of the undulatus complex, the population boundaries of this species is currently defined by genetic data, which do not always agree with traditional concepts of taxonomy based on morphological or ecological variation (Jones & Lovich [eds.] 2009). Genetic research is still needed to clarify the exact limits of distribution of this species.
Distribution: Mexico (Chihuahua, Durango, Nuevo Leon, San Luis Potosi, Sonora, Tamaulipas), USA (Arizona, New Mexico, Texas).


Sceloporus cozumelae

Cozumel Spiny Lizard

Cozumel-Stachelleguan

Cozumel-tornleguan

1927 Sceloporus cozumelae Jones

Distribution: Mexico (Campeche, Quintana Roo [incl. Cozumel], Yucatan).


Sceloporus cryptus

Sierra Juarez Spiny Lizard

Sierra Juarez-Stachelleguan

Sierra Juarez-tornleguan

1967 Sceloporus cryptus Smith & Lynch

Distribution: Mexico (Oaxaca).


Sceloporus cupreus

Golden Long-tailed Spiny Lizard, Upland Long-tailed Spiny Lizard

Kupfer-Stachelleguan

Kobber-tornleguan

1873 Sceloporus cupreus Bocourt
Sceloporus siniferus cupreus Smith & Taylor 1950
1936 Sceloporus cochranae Smith (Smith & Taylor 1950)

Remarks: The phylogenetic analysis of Wiens & Reeder (1997) supported the specific recognition of cupreus.
Distribution: Mexico (Oaxaca).


Sceloporus cyanogenys

Blue-chinned Spiny Lizard, Blue-chinned Rough-scaled Spiny Lizard

Blauer Stachelleguan

Blåkindet Tornleguan, Blå Tornleguan

1885 Sceloporus torquatus cyanogenys Cope
Sceloporus cyanogenys Smith 1938
Sceloporus serrifer cyanogenys Olson 1987
1939 Sceloporus serrifer plioporus Smith (Martinez-Mendez & Mendez de la Cruz 2007)

Other common names:
plioporus: Smith's Blue Spiny Lizard, Western Gulf Rough-scaled Lizard
Remarks: Status of cyanogenys remain unresolved (Wiens & Reeder 1997), but most authors now recognize it as distinct. Not mentioned for San Luis Potosi by Lemos-Espinal, Smith & Woolrich-Piña (2018).
Introduced to: USA (California).
Distribution: Mexico (Coahuila, Hidalgo, Nuevo Leon, San Luis Potosi, Tabasco, Tamaulipas, Veracruz), USA (Texas).


Sceloporus cyanostictus

Eastern Blue-spotted Spiny Lizard, (Blue-spotted Spiny Lizard, Blue Spiny Lizard)

Östlicher Blaufleckiger Stachelleguan

Østlig Blåplettet Tornleguan

1971 Sceloporus jarrovii cyanostictus Axtell & Axtell
Sceloporus cyanostictus Wiens, Reeder & Nieto-Montes de Oca 1999

Remarks: Suggested recognition as a separate species by Wiens & Reeder (1997) and treated as such in subsequent works (e.g., Lemos-Espinal & Smith 2007). Previously included the population now assigned to gadsdeni. Records from Nuevo Leon are based on misidentification. The status of this population is still to be resolved (Díaz-Cárdenas, Ruiz-Sanchez, Castro-Felix, Castañeda-Gaytán, Ruiz-Santana & Gadsden 2017).
Distribution: Mexico (Coahuila).


Sceloporus druckercolini

Graceful Mountain Tree Lizard

Venus-Stachelleguan

Venus-tornleguan

2008 Sceloporus druckercolini Perez-Ramos & Saldana-de la Riva

Distribution: Mexico (Guerrero).


Sceloporus dugesii

Colima Spiny Lizard, (Dugès' Spiny Lizard)

Colima-Stachelleguan

Colima-tornleguan

1873 Sceloporus dugesii Bocourt
1869 Tropidolepis intermedius Dugès (Smith & Taylor 1950; Martínez-Méndez & Méndez-de la Cruz 2007)
Sceloporus intermedius Dugès 1877
Sceloporus dugesii intermedius Smith 1938

Other common names:
dugesii: Common Dugès' Spiny Lizard
intermedius: Western Dugès' Spiny Lizard
Distribution: Mexico (Guanajuato, Jalisco, Mexico, Michoacan, Nayarit, Queretaro, San Luis Potosi, Zacatecas).


Sceloporus edbelli

Durangan Undulate Spiny Lizard, (Bell's Fence Lizard)

Durango-Zaunleguan

Durango-hegnsleguan

1995 Sceloporus undulatus belli Smith, Chiszar & Lemos-Espinal
Sceloporus belli Lemos-Espinal, Smith, Ballinger, Smith & Chiszar 1998
2002 Sceloporus edbelli Smith, Chiszar & Lemos-Espinal [substitute name for Sceloporus undulatus belli Smith, Chiszar & Lemos-Espinal 1995]

Remarks: Status uncertain. Tentatively regarded as a synonym of cowlesi by Leaché & Reeder (2002). Regarded as a synonym of consobrinus by Lemos-Espinal, Smith, Woolrich-Piña & Cruz (2017).
Distribution: Mexico (Chihuahua, Coahuila, Durango, Nuevo Leon, Zacatecas).


Sceloporus edwardtaylori

Ixtepec Spiny Lizard, (Taylor's Spiny Lizard)

Ixtepec-Stachelleguan

Ixtepec-tornleguan

1936 Sceloporus edwardtaylori Smith

Distribution: Mexico (Oaxaca).


Sceloporus esperanzae

Esperanza Spiny Lizard

Esperanza-Stachelleguan

Esperanza-tornleguan

2018 Sceloporus esperanzae McCranie

Remarks: Previously regarded as a population of malachiticus (McCranie 2018).
Distribution: Honduras.





Sceloporus exsul

Queretaran Desert Spiny Lizard

Queretaro-Wüsten-Stachelleguan

Queretaro-ørkentornleguan

1972 Sceloporus exsul Dixon, Ketchersid & Lieb

Distribution: Mexico (Queretaro).


Sceloporus formosus

Veracruz Emerald Spiny Lizard, Mexican Emerald Spiny Lizard, Green Swift

Veracruz-Smaragd-Stachelleguan

Veracruz-smaragdtornleguan

1834 Sceloporus formosus Wiegmann
Tropidolepis formosus Duméril & Bibron 1837
1885 Sceloporus viviparus Cope (Smith & Taylor 1950)
1942 Sceloporus formosus scitulus Smith
Sceloporus scitulus Pérez-Ramos & Saldaña-de La Riva 2008

Other common names:
formosus: Common Mexican Emerald Spiny Lizard, Western Green Swift
scitulus: Striated Mexican Emerald Spiny Lizard, Elegant Green Swift
Remarks: Wiens & Reeder (1997) suggested that scitulus might deserve recognition as a separate species, and some authors have treated it as such (e.g., Pérez-Ramos & Saldaña-de La Riva 2008; Palacios-Aguilar & Flores-Villela 2018).
Distribution: Mexico (Guerrero, Oaxaca, Puebla, Veracruz).


Sceloporus gadoviae

Mezquititlan Spiny Lizard, (Gadow's Spiny Lizard)

Mezquititlan-Stachelleguan

Mezquititlan-tornleguan

1905 Sceloporus gadoviae Boulenger
Lysoptychus gadoviae Larsen & Tanner 1974

Remarks: Occasionally misspelled gadovae (Liner 1996).
Distribution: Mexico (Guerrero, Mexico, Michoacan, Morelos, Oaxaca, Puebla).


Sceloporus gadsdeni

Western Blue-spotted Spiny Lizard, Lagoon Spiny Lizard

Westlicher Blaufleckiger Stachelleguan

Vestlig Blåplettet Tornleguan

2017 Sceloporus gadsdeni Castañeda-Gaytán & Díaz-Cárdenas in Díaz-Cárdenas, Ruiz-Sanchez, Castro-Felix, Castañeda-Gaytán, Ruiz-Santana & Gadsden

Remarks: Previously regarded as a population of cyanostictus.
Distribution: Mexico (Coahuila).


Sceloporus goldmani

San Luis Potosí Bunchgrass Lizard, (Goldman's Bunchgrass Spiny Lizard)

San Luis Potosí-Gras-Stachelleguan

San Luis Potosí-græstornleguan

1937 Sceloporus goldmani Smith

Distribution: Mexico (Aguascalientes, Coahuila, Jalisco, Nuevo Leon, San Luis Potosi, Zacatecas).


Sceloporus graciosus

Common Sagebrush Lizard

Gewöhnlicher Beifußleguan

Almindelig Bynkeleguan

1852 Sceloporus graciosus Baird & Girard
Sceloporus gratiosus Yarrow 1875 [invalid emendation]
1852 Sceloporus gracilis Baird & Girard (Censky 1986)
Sceloporus graciosus gracilis Van Denburgh 1922

Other common names:
gracilis: Western Sagebrush Lizard
graciosus: Northern Sagebrush Lizard
Remarks: Previously included arenicolous and vandenburgianus.
Distribution: Mexico (Baja California Norte), USA (Arizona, California, Colorado, Idaho, Montana, Nebraska, Nevada, New Mexico, North Dakota, Oregon, South Dakota, Utah, Washington, Wyoming).


Sceloporus grammicus

Mesquite Lizard, Northeastern Graphic Lizard, Graphic Spiny Lizard

Mesquite-Stachelleguan

Mesquite-tornleguan

1828 Sceloporus grammicus Wiegmann
1828 Sceloporus microlepidotus Wiegmann (Smith & Taylor 1950)
Sceloporus grammicus microlepidotus Smith & Laufe 1945
1828 Sceloporus pleurostictus Wiegmann (Smith & Taylor 1950)
1852 Sceloporus dispar Baird & Girard (Smith & Taylor 1950)
1867 Sceloporus heterurus Cope (Smith & Taylor 1950)
1890 Sceloporus rubriventris Günther (Smith & Taylor 1950)
1894 Sceloporus microlepis Boulenger [lapsus] (Smith & Taylor 1950)
1916 Sceloporus disparilis Stejneger (Schmidt 1953)
Sceloporus microlepidotus disparilis Dunn 1936
Sceloporus grammicus disparilis Smith & Laufe 1945
1936 Sceloporus pilsbryi Dunn (Smith & Taylor 1950)
1981 Sceloporus grammicus tamaulipensis Sites & Dixon

Other common names:
disparilis: Northern Mesquite Lizard, Northeastern Graphic Lizard
grammicus: Wiegmann's Mesquite Lizard, Southern Plateau Graphic Lizard
microlepidotus: Common Mesquite Lizard
tamaulipensis: Sierra Tamaulipas Mesquite Lizard, Sierra Tamaulipas Graphic Lizard
Remarks: Arechaga-Ocampo, Montalban-Huidobro & Castro-Franco (2008) and García-Vázquez, Canseco-Márquez, Gutiérrez-Mayén & Trujano-Ortega (2009) regarded microlepidotus as a separate species.
Distribution: Mexico (Aguascalientes, Coahuila, Distrito Federal, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico, Michoacan, Morelos, Nayarit, Nuevo Leon, Oaxaca, Puebla, Queretaro, San Luis Potosi, Tamaulipas, Tlaxcala, Veracruz, Zacatecas), USA (Texas).


Sceloporus grandaevus

Cerralvo Island Spiny Lizard, (Cerralvo Island Sator)

Cerralvo-Stachelleguan

Cerralvo-tornleguan

1919 Sator grandaevus Dickerson
Sceloporus grandaevus Wiens & Reeder 1997

Distribution: Mexico (Baja California Sur: Isla Cerralvo).


Sceloporus halli

Oaxacan Large-scaled Spiny Lizard, (Hall's Spiny Lizard)

Großschuppiger Oaxaca-Stachelleguan

Storskællet Oaxaca-tornleguan

1974 Sceloporus megalepidurus halli Dasmann & Smith
Sceloporus halli Wiens & Reeder 1997

Distribution: Mexico (Oaxaca).


Sceloporus heterolepis

Southern Odd-scaled Spiny Lizard, Dorsal-keeled Spiny Lizard, (Jaliscan Spiny Lizard)

Südlicher Gekielter Stachelleguan

Sydlig Kølskællet Tornleguan

1894 Sceloporus heterolepis Boulenger

Distribution: Mexico (Durango, Jalisco, Michoacan, Nayarit, Zacatecas).


Sceloporus hondurensis

Olancho Spiny Lizard

Olancho-Stachelleguan

Olancho-tornleguan

2018 Sceloporus hondurensis McCranie

Remarks: Previously regarded as a population of malachiticus (McCranie 2018). Expected to occur in Nicaragua as well (McCranie 2018).
Distribution: Honduras.





Sceloporus horridus

Mexican Rough Spiny Lizard, Horrible Spiny Lizard

Mexikanischer Rauhschuppiger Stachelleguan

Mexicansk Ru Tornleguan

1834 Sceloporus horridus Wiegmann
1864 Sceloporus oligoporus Cope (Smith & Taylor 1950)
Sceloporus horridus oligoporus Taylor 1938

Remarks: Previously included albiventris. Boyer, Smith & Andreu (1982) suggested that horridus should be a subspecies of spinosus. Smith & Chiszar (1992) demonstrated that they are separate species.
Other common names:
horridus: Southern Rough Spiny Lizard, Wiegmann's Horrible Spiny Lizard
oligoporus: Few-pored Rough Spiny Lizard, Mexican Horrible Spiny Lizard
Distribution: Mexico (Aguascalientes, Colima, Durango, Guerrero, Jalisco, Mexico, Michoacan, Morelos, Nayarit, Oaxaca, Puebla, Zacatecas).


Sceloporus hunsakeri

San Bartolo Spiny Lizard, (Hunsaker's Spiny Lizard)

San Bartolo-Stachelleguan

San Bartolo-tornleguan

1979 Sceloporus hunsakeri Hall & Smith

Distribution: Mexico (Baja California Sur).


Sceloporus insignis

Michoacan Black-collared Spiny Lizard, Coalcoman Black-collared Lizard

Michoacan-Schwarzkragen-Stachelleguan

Sortkravet Michoacan-tornleguan

1967 Sceloporus insignis Webb

Distribution: Mexico (Jalisco, Michoacan).


Sceloporus internasalis

Mail-snouted Spiny Lizard

Oaxaca-Smaragd-Stachelleguan

Oaxaca-smaragdtornleguan

1955 Sceloporus malachiticus internasalis Smith & Bumzahem
Sceloporus internasalis Flores-Villela 1993

Remarks: Regarded as a synonym of salvini by Smith & Pérez-Higareda (1992). Revalidated by Wiens & Reeder (1997).
Distribution: Guatemala, Mexico (Chiapas, Oaxaca, Veracruz).


Sceloporus jalapae

Jalapa Spiny Lizard, Xalapa Spiny Lizard

Jalapa-Stachelleguan

Jalapa-tornleguan

1890 Sceloporus jalapae Günther
Lysoptychus jalapae Larsen & Tanner 1974

Distribution: Mexico (Oaxaca, Puebla, Veracruz).


Sceloporus jarrovii

Mountain Spiny Lizard, (Yarrow's Spiny Lizard)

Gebirgs-Stachelleguan

Bjerg-tornleguan

1875 Sceloporus jarrovii Cope
1950 Sceloporus jarrovii erythrocyaneus Mertens
Sceloporus minor erythrocyaneus Bell, Smith & Chiszar 2003
1989 Sceloporus jarrovi cyaneus Trevino-Saldana [1988]
Sceloporus minor cyaneus Bell, Smith & Chiszar 2003

Other common names:
cyaneus: Blue Mountain Spiny Lizard, Northern Blue Mountain Lizard
erythrocyaneus: Red-spotted Spiny Lizard, Durangan Mountain Spiny Lizard
jarrovi: Common Mountain Spiny Lizard, Common Yarrow's Spiny Lizard
Remarks: Previously included cyanostictus, minor, oberon, sugillatus, and immucronatus. The former four are now regarded as separate species, the latter as a subspecies of minor.
Distribution: Mexico (Aguascalientes, Chihuahua, Coahuila, Durango, Hidalgo, Jalisco, Morelos, Nayarit, Queretaro, Sinaloa, Sonora, Zacatecas), USA (Arizona, New Mexico).


Sceloporus lemosespinali

Chihuahua Graphic Lizard, (Lemos-Espinal's Spiny Lizard)

Chihuahua-Stachelleguan

Chihuahua-tornleguan

2004 Sceloporus lemosespinali Lara-Gongora

Distribution: Mexico (Chihuahua, Durango, Sonora).


Sceloporus licki

Cape Arboreal Spiny Lizard

Kap-Stachelleguan

Kap-tornleguan

1895 Sceloporus licki Van Denburgh
Sceloporus orcutti licki Smith 1939

Remarks: Revalidated as a separate species by Hall & Smith (1979).
Distribution: Mexico (Baja California Sur).


Sceloporus lineatulus

Santa Catalina Island Spiny Lizard

Santa Catalina-Stachelleguan

Santa Catalina-tornleguan

1919 Sceloporus lineatulus Dickerson
Sceloporus magister lineatulus Smith 1939

Remarks: Revalidated as a separate species by Grismer & McGuire (1996), although already Flores-Villela (1993) and Liner (1994) treated it as such.
Distribution: Mexico (Baja California Sur: Isla Santa Catalina).


Sceloporus lineolateralis

Durangan Lined Spiny Lizard, Lined Spiny Lizard, (Durangan Spiny Lizard)

Gestreifter Durango-Stachelleguan

Stribet Durango-tornleguan

1936 Sceloporus lineolateralis Smith
Sceloporus jarrovii lineolateralis Webb & Hensley 1959

Remarks: Regarded as a subspecies of jarrovi by some authors (e.g., Martínez-Méndez & Méndez-de la Cruz 2007). Regarded as a valid species in most recent works (e.g., Wiens & Reeder 1997; Bell, Smith & Chiszar 2003; Wilson & Johnson 2010).
Distribution: Mexico (Coahuila, Durango, Zacatecas).


Sceloporus lunaei

Guatemalan Plateau Lizard

Guatemalischer Plateau-Stachelleguan

Guatemalansk Plateauleguan

1873 Sceloporus lunaei Bocourt
Sceloporus acanthinus lunaei Stuart 1963

Distribution: Guatemala.


Sceloporus lundelli

Belize Spiny Lizard, (Lundell's Spiny Lizard)

Belize-Stachelleguan

Belize-tornleguan

1939 Sceloporus lundelli Smith
1939 Sceloporus lundelli gaigeae Smith

Other common names:
gaigeae: Gaige's Spiny Lizard
lundelli: Common Lundell's Spiny Lizard
Distribution: Belize, Guatemala, Mexico (Campeche, Quintana Roo, Tabasco, Yucatan).


Sceloporus macdougalli

Tehuantepec Spiny Lizard, (MacDougall's Spiny Lizard)

Tehuantepec-Stachelleguan

Tehuantepec-tornleguan

1953 Sceloporus macdougalli Smith & Bumsahem

Distribution: Mexico (Oaxaca).


Sceloporus maculosus

Rio Nazas Spiny Lizard, Spotted Spiny Lizard, Northern Snub-nosed Spiny Lizard

Rio Nazas-Stachelleguan

Rio Nazas-tornleguan

1934 Sceloporus maculosus Smith
Lysoptychus maculosus Larsen & Tanner 1974

Remarks: A record from San Luis Potosi (Flores-Villela, Hernández-Garcia & Nieto-Montes de Oca 1991) refers to consobrinus (Smith & Flores-Villela 1994).
Distribution: Mexico (Coahuila, Durango, Zacatecas).


Sceloporus magister

Northern Desert Spiny Lizard, Desert Spiny Lizard

Nördlicher Wüsten-Stachelleguan

Nordlig Ørkentornleguan

1854 Sceloporus magister Hallowell
Sceloporus spinosus magister Echegoyen 2009
1955 Sceloporus magister cephaloflavus Tanner

Other common names:
cephaloflavus: Orange-headed Spiny Lizard
magister: Purple-backed Spiny Lizard, Sonoran Spiny Lizard
Remarks: Previously included bimaculosus, lineatulus, rufidorsus, uniformis, and zosteromus. Also cephaloflavus have been suggested to represent a separate species (Jones & Lovich (eds.) 2009). The recognition of some of these forms as separate species has been disputed (e.g., Leaché & Mulcahy 2007).
Introduced to: USA (Florida).
Distribution: Mexico (Baja California Norte, Sinaloa, Sonora), USA (Arizona, California, Colorado, Nevada, New Mexico, Utah).


Sceloporus malachiticus

Malachite Spiny Lizard, (Emerald Swift, Green Spiny Lizard)

Malachit-Stachelleguan, Malachitgrüner Stachelleguan

Malakit-tornleguan

1864 Sceloporus malachiticus Cope
1890 Sceloporus irazuensis Günther (Peters & Donoso-Barros 1970)

Remarks: Previously included acanthinus, internasalis, salvini, schmidti, smaragdinus, taeniocnemis, and the populations now assigned to esperanzae and hondurensis. E. Smith (2001, in an unpublished dissertation), as well as Acevedo, Wilson, Cano & Vázquez-Almazán (2010) and McCranie (2015) treated schmidti as a separate species.
Distribution: Costa Rica, El Salvador, Guatemala, Nicaragua, Panama.


Sceloporus megalepidurus

Veracruz Large-scaled Spiny Lizard, Cursorial Spiny Lizard, (Large-scaled Spiny Lizard)

Ostmexikanischer Großschuppiger Stachelleguan

Østmexicansk Storskællet Tornleguan

1934 Sceloporus megalepidurus Smith

Remarks: Previously included halli and pictus. Records from Oaxaca (e.g., Casas-Andreu, Mendez-de la Cruz & Aguilar-Miguel 2004) refer to these two species.
Distribution: Mexico (Hidalgo, Mexico, Oaxaca, Puebla, Tlaxcala, Veracruz).


Sceloporus melanorhinus

Black-nosed Spiny Lizard, Black-nosed Lizard, Pastel Tree Lizard, Pastel Spiny Lizard

Schwarzschnäuziger Stachelleguan

Sortsnudet Tornleguan

1876 Sceloporus melanorhinus Bocourt
1939 Sceloporus melanorhinus calligaster Smith
1947 Sceloporus melanorhinus stuarti Smith

Other common names:
calligaster: Pastel-bellied Spiny Lizard
melanorhinus: Southern Black-nosed Lizard
stuarti: Stuart's Pastel Tree Lizard
Remarks: The subspecies stuarti probably deserves specific recognition (Wiens & Reeder 1997).
Distribution: Guatemala, Mexico (Chiapas, Colima, Durango, Guerrero, Jalisco, Mexico, Michoacan, Morelos, Nayarit, Oaxaca, Puebla).


Sceloporus merriami

Canyon Lizard

Canyon-Stachelleguan

Canyon-tornleguan

1904 Sceloporus merriami Stejneger
Lysoptychus merriami Larsen & Tanner 1974
1937 Sceloporus merriami annulatus Smith
1960 Sceloporus merriami australis Williams, Smith & Chrapliwy
1973 Sceloporus merriami longipunctatus Olson
2000 Sceloporus merriami williamsi Lemos-Espinal, Chiszar & Smith
2001 Sceloporus merriami ballingeri Lemos-Espinal & Smith in Lemos-Espinal, Smith, Auth & Chiszar
2003 Sceloporus merriami sanojae Lemos-Espinal in Smith, Lemos-Espinal & Chiszar

Other common names:
annulatus: Big Bend Canyon Lizard
australis: Southern Canyon Lizard, Southeastern Canyon Lizard
ballingeri: Ballinger's Canyon Lizard
longipunctatus: Presidio Canyon Lizard
merriami: Merriam's Canyon Lizard
sanojae: Sanoja's Canyon Lizard
williamsi: Williams' Canyon Lizard
Distribution: Mexico (Chihuahua, Coahuila, Durango, Nuevo Leon), USA (Texas).


Sceloporus minor

Lesser Spiny Lizard, Minor Spiny Lizard, (Cope's Mountain Spiny Lizard)

Kleiner Stachelleguan

Lille Tornleguan

1885 Sceloporus torquatus minor Cope
Sceloporus jarrovi minor Smith 1938
Sceloporus minor Webb & Axtell 1994
1936 Sceloporus jarrovii immucronatus Smith
Sceloporus minor immucronatus Lemos-Espinal & Smith 2007

Other common names:
immucronatus: Southern Blue Minor Spiny Lizard, (Blue Yarrow's Spiny Lizard)
minor: Red Minor Spiny Lizard
Distribution: Mexico (Coahuila, Guanajuato, Hidalgo, Jalisco, Mexico, Nuevo Leon, Queretaro, San Luis Potosi, Tamaulipas, Veracruz, Zacatecas).


Sceloporus mucronatus

Oaxacan Crevice Spiny Lizard, Cleft Lizard, (Southern Crevice Spiny Lizard)

Oaxaca-Spalten-Stachelleguan

Oaxaca-spaltetornleguan

1885 Sceloporus torquatus mucronatus Cope
Sceloporus mucronatus Smith 1938
2002 Sceloporus mucronatus olsoni Webb, Lemos-Espinal & Smith

Other common names:
mucronatus: Central Crevice Spiny Lizard
olsoni: Olson's Crevice Spiny Lizard
Remarks: Olson (1998) included mucronatus in the synonymy of poinsetti, however, this arrangement has not been followed in subsequent publications (e.g., Köhler & Heimes 2002; Webb, Lemos-Espinal & Smith 2002; Martinez-Mendez & Mendez de la Cruz 2007). Previously included aureolus and omiltemanus (e.g., Köhler & Heimes 2002).
Distribution: Mexico (Distrito Federal, Hidalgo, Mexico, Oaxaca, Puebla, Tlaxcala, Veracruz).


Sceloporus nelsoni

Sinaloan Spiny Lizard, (Nelson's Spiny Lizard)

Sinaloa-Stachelleguan

Sinaloa-tornleguan

1923 Sceloporus nelsoni Cochran
1960 Sceloporus nelsoni coeruleus Tanner & Robison [not Sceloporus ornatus caeruleus Smith 1936]
1960 Sceloporus nelsoni barrancorum Tanner & Robison [substitute name for Sceloporus nelsoni coeruleus Tanner & Robison 1960]
Sceloporus nelsoni barrancarum Bell, Smith & Chiszar 2003 [emendation]

Other common names:
barrancarum: Northern Nelson's Spiny Lizard, Nelson's Barranca Spiny Lizard
nelsoni: Southern Nelson's Spiny Lizard
Remarks: According to Bell, Smith & Chiszar (2003), the correct spelling of the specific epithet barrancorum should be barrancarum.
Distribution: Mexico (Chihuahua, Durango, Jalisco, Nayarit, Sinaloa, Sonora).


Sceloporus oberon

Royal Mountain Spiny Lizard, Royal Lesser Minor Lizard, Black Spiny Lizard

König-Stachelleguan

Konge-tornleguan

1941 Sceloporus jarrovii oberon Smith & Brown
Sceloporus oberon Wiens, Reeder & Nieto-Montes de Oca 1999
Sceloporus ornatus oberon Martinez-Mendez & Mendez de la Cruz 2007

Remarks: Previously regarded as a subspecies of jarrovi and, more recently, of ornatus. Treated here as a separate species, following several recent publications (e.g., Köhler & Heimes 2002; Lemos-Espinal & Smith 2007; Wiens, Kuczynski, Arif & Reeder 2010).
Distribution: Mexico (Coahuila, Nuevo Leon).


Sceloporus occidentalis

Western Fence Lizard

Westlicher Zaunleguan

Vestlig Tornleguan

1852 Sceloporus occidentalis Baird & Girard
1854 Sceloporus biseriatus Hallowell (Schmidt 1953)
Sceloporus occidentalis biseriatus Camp 1916
1858 Sceloporus longipes Baird
Sceloporus occidentalis longipes Liner 1994
1885 Sceloporus undulatus bocourtii Boulenger (Schmidt 1953)
Sceloporus occidentalis bocourti Queiroz & Reeder in Crother (ed.) 2008
1916 Sceloporus occidentalis taylori Camp

Other common names:
becki: Island Fence Lizard
biseriatus: San Joaquin Fence Lizard
bocourti: Coast Range Fence Lizard
longipes: Great Basin Fence Lizard
occidentalis: Northwestern Fence Lizard
taylori: Sierra Fence Lizard
Remarks: Previously included becki. In Montana, known from only a single population which may have been introduced (Werner, Maxell, Hendricks & Flath 2004).
Introduced to: Guam.
Distribution: Mexico (Baja California Norte [incl. Isla Cedros]), USA (California, Idaho, Montana, Nevada, Oregon, Utah, Washington).


Sceloporus ochoterenae

Queretaran Spiny Lizard, (Ochoterena's Snub-nosed Spiny Lizard)

Queretaro-Stachelleguan

Queretaro-tornleguan

1934 Sceloporus ochoterenae Smith
Lysoptychus ochoterenae Larsen & Tanner 1974

Distribution: Mexico (Guerrero, Mexico, Morelos, Oaxaca, Puebla).


Sceloporus olivaceus

Texas Spiny Lizard

Texas-Stachelleguan

Texas-tornleguan

1934 Sceloporus olivaceus Smith
1893 Sceloporus spinosus floridanus Stejneger [not Sceloporus floridanus Baird 1858] (Smith & Taylor 1950)

Distribution: Mexico (Coahuila, Nuevo Leon, San Luis Potosi, Tamaulipas), USA (Oklahoma, Texas).


Sceloporus olloporus

Central American Rose-bellied Lizard

Zentralamerikanischer Rosabaüchiger Stachelleguan

Centralamerikansk Rosabuget Tornleguan

1937 Sceloporus variabilis olloporus Smith
Sceloporus olloporus Mendoza-Quijano, Flores-Villela & Sites 1998

Remarks: Mendoza-Quijano, Flores-Villela & Sites (1998) regarded olloporus as a separate species, but they based this taxonomic decision on a single examined specimen. Several authors chose not to follow this, pending further studies (e.g., Köhler 2001, 2008; Köhler & Heimes 2002; Savage 2002; Bolaños, Savage & Gerardo 2010; Wilson & Johnson 2010). Validity as a separate species was supported by Solis-Zurita, Luna & González (2019).
Distribution: Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Chiapas), Nicaragua.





Sceloporus omiltemanus

Guerreran Crevice Spiny Lizard, Southern Cleft Spiny Lizard

Guerrero-Spalten-Stachelleguan

Guerrero-spaltetornleguan

1890 Sceloporus omiltemanus Günther
Sceloporus torquatus omiltemanus Taylor 1931
Sceloporus mucronatus omiltemanus Smith 1936

Remarks: Revalidated as a separate species by Martinez-Mendez & Mendez de la Cruz (2007).
Distribution: Mexico (Guerrero, Oaxaca).


Sceloporus orcutti

Granite Spiny Lizard

Granit-Stachelleguan

Granit-tornleguan

1893 Sceloporus orcutti Stejneger
1899 Sceloporus digueti Mocquard (Weintraub 1980)

Distribution: Mexico (Baja California Norte, Baja California Sur), USA (California).


Sceloporus ornatus

Ornate Spiny Lizard

Gewöhnlicher Pracht-Stachelleguan

Almindelig Pragttornleguan

1858 Sceloporus ornatus Baird

Remarks: Previously included caeruleus and oberon.
Distribution: Mexico (Coahuila, Nuevo Leon).


Sceloporus palaciosi

Zempoala Bunchgrass Lizard, (Palacios' Bunchgrass Lizard, Palacios' Graphic Lizard)

Zempoala-Gras-Stachelleguan

Zempoala-græstornleguan

1983 Sceloporus palaciosi Lara-Gongora

Distribution: Mexico (Distrito Federal, Guerrero, Mexico, Michoacan, Morelos, Puebla).


Sceloporus parvus

Blue-bellied Spiny Lizard, (Dwarf Spiny Lizard)

Blaubäuchiger Stachelleguan

Blåbuget Tornleguan

1934 Sceloporus parvus Smith
Lysoptychus parvus Larsen & Tanner 1974
1937 Sceloporus parvus scutulatus Smith

Other common names:
parvus: Northern Blue-bellied Spiny Lizard
scutulatus: Southern Blue-bellied Spiny Lizard
Remarks: A record from Durango (Flores-Villela, Hernández-Garcia & Nieto-Montes de Oca 1991) refer to bimaculosus (Smith & Flores-Villela 1994).
Distribution: Mexico (Coahuila, Hidalgo, Nuevo Leon, Queretaro, San Luis Potosi, Tamaulipas, Zacatecas).


Sceloporus pictus

Painted Spiny Lizard, Painted Cursorial Spiny Lizard

Gemalter Stachelleguan

Malet Tornleguan

1936 Sceloporus pictus Smith
Sceloporus megalepidurus pictus Smith & Smith 1976

Remarks: Sites, Archie, Cole & Villela (1992) questioned the distinctness of pictus and megalepidurus. Some authors regard pictus as a subspecies of megalepidurus (e.g., Smith, Auth, Chiszar, Lintz & Brown 2000). Wiens & Reeder (1997) tentatively retained them as separate species.
Distribution: Mexico (Hidalgo, Mexico, Oaxaca, Veracruz).


Sceloporus poinsettii

Northern Crevice Spiny Lizard, (Crevice Spiny Lizard)

Nördlicher Spalten-Stachelleguan

Nordlig Spaltetornleguan

1852 Sceloporus poinsettii Baird & Girard
Sceloporus torquatus poinsettii Cope 1885
1958 Sceloporus poinsetti macrolepis Smith & Chrapliwy
1958 Sceloporus poinsetti polylepis Smith & Chrapliwy
1987 Sceloporus poinsetti robisoni Tanner
2006 Sceloporus poinsettii axtelli Webb

Other common names:
axtelli: Texas Crevice Spiny Lizard
macrolepis: Large-scaled Crevice Spiny Lizard, Western Crevice Spiny Lizard
poinsetti: New Mexico Crevice Spiny Lizard, Northern Crevice Spiny Lizard
polylepis: Southern Crevice Spiny Lizard, Small-scaled Crevice Spiny Lizard
robisoni: Robison's Crevice Spiny Lizard
Remarks: Olson (1998) included mucronatus in the synonymy of poinsetti, however, this arrangement has not been followed by all authors (e.g., Köhler & Heimes 2002; Webb, Lemos-Espinal & Smith 2002). Included for Jalisco on the basis of Cruz-Sáenz, Guerrero, Lazcano & Téllez-López (2009) and Cruz-Sáenz, Muñoz-Nolasco, Mata-Silva, Johnson, García-Padilla & Wilson (2017), although Chávez-Avila, Casas-Andreu, García-Aguayo, Cifuentes-Lemus & Cupul-Magaña (2015) condidered the presence of the species in the state as unconfirmed.
Introduced to: USA (Florida).
Distribution: Mexico (Chihuahua, Coahuila, Durango, Jalisco, Nayarit, Nuevo Leon, San Luis Potosi, Sinaloa, Sonora, Zacatecas), USA (New Mexico, Texas).


Sceloporus prezygus

Chiapan Blue Spiny Lizard, Chiapan Rough-scaled Spiny Lizard

Blauer Chiapas-Stachelleguan

Blå Chiapas-tornleguan

1942 Sceloporus prezygus Smith
Sceloporus serrifer prezygus Smith & Alvarez del Toro 1964

Remarks: The phylogenetic analysis of Wiens & Reeder (1997) supported the specific recognition of prezygus. Martínez-Méndez, Mejía, Rocha-Gómez & Méndez-de la Cruz (2012) retained it as a subspecies of serrifer, pending further studies.
Distribution: Belize, Guatemala, Mexico (Chiapas).


Sceloporus pyrocephalus

Boulder Spiny Lizard, Red-headed Spiny Lizard

Rotkopf-Stachelleguan

Rødhovedet Tornleguan

1864 Sceloporus pyrocephalus Cope
Sceloporus pyrrhocephalus Cope 1885 [emendation]

Distribution: Mexico (Colima, Guerrero, Jalisco, Mexico, Michoacan).


Sceloporus salvini

Veracruz Green Spiny Lizard, (Salvin's Spiny Lizard)

Grüner Veracruz-Stachelleguan

Grøn Veracruz-tornleguan

1890 Sceloporus salvini Günther
Sceloporus malachiticus salvini Smith 1942

Remarks: Previously included internasalis.
Distribution: Guatemala, Mexico (Oaxaca, Veracruz).


Sceloporus samcolemani

Nuevo Leon Bunchgrass Lizard, (Coleman's Bunchgrass Lizard)

Nuevo Leon-Gras-Stachelleguan

Nuevo Leon-græstornleguan

1974 Sceloporus scalaris samcolemani Smith
Sceloporus samcolemani Smith, Watkins-Colwell, Liner & Chiszar 1996

Distribution: Mexico (Coahuila, Nuevo Leon).


Sceloporus scalaris

Light-bellied Bunchgrass Lizard

Hellbäuchiger Gras-Stachelleguan

Lysbuget Græstornleguan

1828 Sceloporus scalaris Wiegmann

Remarks: Previously included brownorum, samcolemani, slevini, unicanthalis, and the population now known as aurantius. Records from Chihuahua, Coahuila, Durango, Nuevo Leon, and Zacatecas refer to these other species. Records from USA (Arizona) refer to slevini.
Distribution: Mexico (Distrito Federal, Guanajuato, Hidalgo, Jalisco, Mexico, Michoacan, Morelos, Nayarit, Puebla, Queretaro, San Luis Potosi, Tamaulipas, Tlaxcala).


Sceloporus schmidti

Sula Spiny Lizard

Sula-Stachelleguan

Sula-tornleguan

1927 Sceloporus schmidti Jones

Remarks: Previously regarded as a synonym of smaragdinus (e.g., Smith 1939) and/or malachiticus (e.g., Meyer 1969). Revalidated as a separate species by McCranie (2018).
Distribution: Guatemala, Honduras.





Sceloporus serrifer

Southern Blue Spiny Lizard, Rough-scaled Spiny Lizard, (Blue-spotted Spiny Lizard)

Südlicher Blauer Stachelleguan

Sydlig Blå Tornleguan

1866 Sceloporus serrifer Cope
1952 Sceloporus serrifer cariniceps Martin

Other common names:
serrifer: Yucatecan Rough-scaled Spiny Lizard, Eastern Blue Spiny Lizard
Remarks: Previously included prezygus (e.g., Smith & Alvarez del Toro 1964), cyanogenys (e.g., Olson 1987) and plioporus Smith 1939, the latter now regarded as a synonym of cyanogenys (Martinez-Mendez & Mendez de la Cruz 2007). Older records from Chiapas refer to prezygus, but serrifer was documented from the state by León & Reynoso (2005).
Distribution: Belize, Guatemala, Mexico (Campeche, Chiapas, Hidalgo, Queretaro, Quintana Roo, San Luis Potosi, Tabasco, Tamaulipas, Veracruz, Yucatan).


Sceloporus shannonorum

Northern Odd-scaled Spiny Lizard, (Shannon's Spiny Lizard)

Nördlicher Gekielter Stachelleguan

Nordlig Kølskællet Tornleguan

1959 Sceloporus shannonorum Langebartel
Sceloporus heterolepis shannonorum Webb 1969

Remarks: Revalidated as a separate species by Wiens & Reeder (1997).
Distribution: Mexico (Durango, Jalisco, Nayarit, Sinaloa, Zacatecas).


Sceloporus siniferus

Long-tailed Spiny Lizard

Langschwanz-Stachelleguan

Langhalet Tornleguan

1869 Sceloporus siniferus Cope
1873 Sceloporus humeralis Bocourt (Peters & Donoso-Barros 1970)

Remarks: Although included for Jalisco by Cruz-Sáenz, Guerrero, Lazcano & Télles-López (2009), presence in the state is considered undocumented herein, following Cruz-Sáenz, Muñoz-Nolasco, Mata-Silva, Johnson, García-Padilla & Wilson (2017), who did not list the species for Jalisco.
Distribution: Guatemala, Mexico (Chiapas, Guerrero, Michoacan, Morelos, Oaxaca).


Sceloporus slevini

Arizona Bunchgrass Lizard, (Slevin's Bunchgrass Lizard)

Arizona-Gras-Stachelleguan

Arizona-græstornleguan

1937 Sceloporus scalaris slevini Smith
Sceloporus slevini Tanner 1987

Remarks: Validity as separate species confirmed by Smith, Watkins-Colwell, Liner & Chiszar (1996).
Distribution: Mexico (Chihuahua, Durango, Sonora), USA (Arizona, New Mexico).


Sceloporus smaragdinus

Chiapan Malachite Spiny Lizard, (Emerald Spiny Lizard, Bocourt's Emerald Spiny Lizard)

Chiapas-Smaragd-Stachelleguan

Chiapas-smaragdtornleguan

1873 Sceloporus smaragdinus Bocourt
Sceloporus formosus smaragdinus Smith 1939
Sceloporus malachiticus smaragdinus Smith 1942
Sceloporus smaragdinus Stuart 1971

Distribution: Guatemala, Mexico (Chiapas).


Sceloporus smithi

Oaxacan Rose-bellied Lizard, (Smith's Rose-bellied Lizard)

Rosabaüchiger Oaxaca-Stachelleguan

Rosabuget Oaxaca-tornleguan

1937 Sceloporus variabilis smithi Hartweg & Oliver
Sceloporus smithi Smith, Perez-Higareda & Chiszar 1993

Distribution: Mexico (Chiapas, Oaxaca).


Sceloporus spinosus

Eastern Mexican Spiny Lizard, Eastern Spiny Lizard

Gewöhnlicher Ostmexikanischer Stachelleguan

Almindelig Østmexicansk Tornleguan

1828 Sceloporus spinosus Wiegmann
1936 Sceloporus spinosus caeruleopunctatus Smith
1951 Sceloporus spinosus apicalis Smith & Smith

Other common names:
apicalis: Miahuatlán Spiny Lizard, Oaxacan Spiny Lizard
caeruleopunctatus: Blue-spotted Spiny Lizard
spinosus: Common Eastern Spiny Lizard, (Wiegmann's Spiny Lizard)
Remarks: The two subspecies apicalis and spinosus probably deserve specific recognition (Wiens & Reeder 1997). Boyer, Smith & Andreu (1982) suggested that horridus should be a subspecies of spinosus. Smith & Chiszar (1992) demonstrated that they are separate species.
Introduced to: Mexico (Chiapas).
Distribution: Mexico (Aguascalientes, Distrito Federal, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico, Michoacan, Morelos, Nuevo Leon, Oaxaca, Puebla, Queretaro, San Luis Potosi, Sinaloa, Tamaulipas, Veracruz, Zacatecas).


Sceloporus squamosus

Central American Dwarf Spiny Lizard, (Dwarf Spiny Lizard, Mexican Spiny Lizard, Bocourt's Spiny Lizard)

Centralamerikanischer Zwerg-Stachelleguan

Centralamerikansk Dværgtornleguan

1874 Sceloporus squamosus Bocourt
1874 Sceloporus fulvus Bocourt (Peters & Donoso-Barros 1970)

Distribution: Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Chiapas, Oaxaca), Nicaragua.


Sceloporus stejnegeri

Guerreran Black-collared Spiny Lizard, (Stejneger's Black-collared Spiny Lizard)

Guerrero-Schwarzkragen-Stachelleguan

Sortkravet Guerrero-tornleguan

1942 Sceloporus stejnegeri Smith

Distribution: Mexico (Guerrero).


Sceloporus subniger

Tolucan Bunchgrass Lizard, Plateau Bunchgrass Lizard

Toluca-Gras-Stachelleguan

Toluca-græstornleguan

1958 Sceloporus aeneus subniger Poglayen & Smith
Sceloporus subniger Guilette & Smith 1985

Remarks: Already Guilette & Smith (1985) suggested species status for subniger, a suggestion followed by Liner (1994) and Wiens & Reeder (1997).
Distribution: Mexico (Guanajuato, Mexico).


Sceloporus subpictus

Paint-bellied Spiny Lizard

Bauchgemalter Stachelleguan

Bugmalet Tornleguan

1965 Sceloporus subpictus Lynch & Smith

Distribution: Mexico (Oaxaca).


Sceloporus sugillatus

Zempoala Spiny Lizard, Zempoala Bar-bellied Lizard, Variable Mountain Spiny Lizard

Zempoala-Stachelleguan

Zempoala-tornleguan

1942 Sceloporus jarrovii sugillatus Smith
Sceloporus sugillatus Wiens, Reeder & Nieto-Montes de Oca 1999

Remarks: Recognition as a separate species was suggested by Wiens & Reeder (1997), and followed by most subsequent works (e.g., Lemos-Espinal & Smith 2007; Mendoza-Hernandez, Garcia-Vazquez & Nieto-Montes de Oca 2008).
Distribution: Mexico (Distrito Federal, Mexico, Morelos).


Sceloporus taeniocnemis

Nuclear Emerald Spiny Lizard

Guatemalischer Smaragd-Stachelleguan

Guatemalansk Smaragdtornleguan

1885 Sceloporus taeniocnemis Cope
Sceloporus malachiticus taeniocnemis Smith 1942
1971 Sceloporus taeniocnemis hartwegi Stuart
Sceloporus hartwegi Pérez-Ramos & Saldaña-de La Riva 2008

Remarks: Revalidated as a separate species by Stuart (1971). Pérez-Ramos & Saldaña-de La Riva (2008) listed hartwegi as a separate species.
Other common names:
hartwegi: Hartweg's Emerald Spiny Lizard
taeniocnemis: Guatemalan Emerald Spiny Lizard
Distribution: Guatemala, Mexico (Chiapas).


Sceloporus tanneri

Juquila Spiny Lizard, (Tanner's Spiny Lizard)

Juquila-Stachelleguan

Juquila-tornleguan

1975 Sceloporus tanneri Smith & Larsen

Distribution: Mexico (Oaxaca).


Sceloporus teapensis

Teapen Rose-bellied Lizard

Teapen-Stachelleguan

Teapen-tornleguan

1890 Sceloporus teapensis Günther
Sceloporus variabilis teapensis Cole 1978

Remarks: Status of teapensis remains unresolved (Mendoza-Quijano, Flores-Villela & Sites 1998). Most recent works have treated it as a valid species (e.g., Campbell & Vannini 1989; Smith, Perez-Higareda & Chiszar 1993; Liner 1994, 2007; Köhler & Heimes 2002). Records from Belize (e.g.; Campbell & Vannini 1989; Smith, Perez-Higareda & Chiszar 1993; Lee 1996) refer to variabilis (Stafford & Meyer 2000).
Distribution: Belize, Guatemala, Mexico (Campeche, Chiapas, Oaxaca, Tabasco, Veracruz).


Sceloporus torquatus

Torquate Spiny Lizard, Torquate Lizard, Crevice Swift

Kragen-Stachelleguan

Krave-tornleguan

1828 Sceloporus torquatus Wiegmann
1885 Sceloporus ferrariperezi Cope (Smith & Taylor 1950)
1885 Sceloporus melanogaster Cope (Smith & Taylor 1950)
Sceloporus ferrariperezi melanogaster Smith 1939
Sceloporus torquatus melanogaster Liner 1994
1936 Sceloporus binocularis Dunn (Smith & Taylor 1950)
Sceloporus ferrariperezi binocularis Smith 1939
Sceloporus torquatus binocularis Liner 1994
1974 Sceloporus torquatus mikeprestoni Smith & Alvarez
1986 Sceloporus torquatus madrensis Olson

Other common names:
binocularis: Nuevo Leon Torquate Spiny Lizard
madrensis: Sierra Madre Torquate Spiny Lizard, Lesser Torquate Lizard
melanogaster: Black-bellied Torquate Spiny Lizard, Central Plateau Torquate Lizard
mikeprestoni: Preston's Torquate Spiny Lizard
torquatus: Common Torquate Spiny Lizard, Plateau Crevice Swift
Distribution: Mexico (Aguascalientes, Distrito Federal, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, Mexico, Michoacan, Morelos, Nayarit, Nuevo Leon, Puebla, Queretaro, San Luis Potosi, Tamaulipas, Veracruz, Zacatecas).


Sceloporus tristichus

Western Plateau Lizard, Plateau Fence Lizard

Westlicher Plateau-Stachelleguan

Vestlig Plateau-leguan

1875 Sceloporus tristichus Cope
Sceloporus undulatus tristichus Cope 1900
1890 Sceloporus elongatus Stejneger (Leaché & Reeder 2002)
Sceloporus undulatus elongatus Burt 1933
1956 Sceloporus undulatus erythrocheilus Maslin (Leaché & Reeder 2002)

Other common names:
elongatus: Northern Plateau Lizard
erythrocheilus: Red-lipped Plateau Lizard, Red-lipped Prairie Lizard
tristichus: Southern Plateau Lizard
Remarks: This species corresponds to Leaché's & Reeder's (2002) "western clade", which encompasses populations previously regarded as the subspecies elongatus, erythrocheilus, and tristichus of the wide-ranging undulatus. As with other members of the undulatus complex, the population boundaries of this species is currently defined by genetic data, which do not always agree with traditional concepts of taxonomy based on morphological or ecological variation (Jones & Lovich [eds.] 2009). Genetic research is still needed to clarify the exact limits of distribution of this species.
Distribution: USA (Arizona, Colorado, Nevada, New Mexico, Utah, Wyoming).


Sceloporus undulatus

Eastern Fence Lizard, Undulate Spiny Lizard

Östlicher Zaunleguan

Østlig Tornleguan

1801 Stellio undulatus Bosc & Daudin in Sonnini & Latreille
Sceloporus undulatus Wiegmann 1828
Agama undulatus Harlan 1829
Tropidolepis undulatus Holbrook 1838
1818 Lacerta fasciata Green [not Lacerta fasciata Linnaeus 1758] (Schmidt 1953)
1818 Lacerta hyacinthina Green (Schmidt 1953)
Sceloporus undulatus hyacinthinus Smith 1948
1832 Lopherpes dicyanelus Rafinesque (Schmidt 1953)
1852 Sceloporus thayeri Baird & Girard (Schmidt 1953)
1858 Sceloporus floridanus Baird (Schmidt 1953)

Other common names:
hyacinthinus: Northern Fence Lizard
undulatus: Southern Fence Lizard
Remarks: Previously included consobrinus, cowlesi, edbelli, tristichus, and virgatus. Apart from virgatus, the status of these forms is contentious. Leaché & Reeder (2002) recommended treating four different clades as evolutionary species (consobrinus, cowlesi, tristichus, undulatus), including belli (=edbelli) as a tentative synonym of cowlesi. While many recent publications have followed this (e.g., Brennan & Holycross 2006; Rorabaugh 2008; Enderson, Quijada-Mascarenas, Turner, Rosen & Bezy 2009; Jones & Lovich (eds.) 2009), some authors have retained them as subspecies of undulatus, pending further studies to clarify the exact limits of these species and their distribution (e.g., Bell, Smith & Chiszar 2003; Lemos-Espinal & Smith 2007). Based on molecular studies, Miles, Noecker, Roosenburg & White (2002) did not support the recognition of subspecies. This species corresponds to Leaché's & Reeder's (2002) "eastern clade", which encompasses some, but not all, populations which were previously regarded as the subspecies hyacinthinus and undulatus of the wide-ranging undulatus. As with other members of the undulatus complex, the population boundaries of this species is currently defined by genetic data, which do not always agree with traditional concepts of taxonomy based on morphological or ecological variation (Jones & Lovich [eds.] 2009). Genetic research is still needed to clarify the exact limits of the distribution of this species.
Distribution: USA (Alabama, Delaware, Florida, Georgia, Indiana, Kentucky, Maryland, Nebraska, New Jersey, New York, North Carolina, Ohio, Oklahoma, Pennsylvania, South Carolina, Tennessee, Virginia, Washington, D.C., West Virginia).


Sceloporus unicanthalis

Southwestern Bunchgrass Lizard

Südwestlicher Gras-Stachelleguan

Sydvestlig Græstornleguan

1937 Sceloporus scalaris unicanthalis Smith
Sceloporus unicanthalis Grummer & Bryson 2014

Distribution: Mexico (Colima, Durango, Jalisco, Nayarit).


Sceloporus uniformis

Yellow-backed Spiny Lizard

Gelbrücken-Stachelleguan

Gulrygget Tornleguan

1955 Sceloporus magister uniformis Phelan & Brattstrom
Sceloporus uniformis Schulte, Macey & Papenfuss 2006
1955 Sceloporus magister transversus Phelan & Brattstrom (Schulte, Macey & Papenfuss 2006)

Other common names:
transversus: Barred Spiny Lizard
Remarks: Previously regarded as a subspecies of magister. Recognized as a separate species by Schulte, Macey & Papenfuss (2006), followed by others (e.g., Enderson & Bezy 2010), although Leaché & Mulcahy (2007) presented evidence in support of treating the two taxa as conspecific.
Distribution: USA (Arizona, California, Nevada, Utah).


Sceloporus utiformis

West Mexican Large-scaled Spiny Lizard

Westmexikanischer Großschuppiger Stachelleguan

Vestmexicansk Storskællet Tornleguan

1864 Sceloporus utiformis Cope

Distribution: Mexico (Colima, Guerrero, Jalisco, Michoacan, Morelos, Nayarit, Sinaloa, Zacatecas).


Sceloporus vandenburgianus

Southern Sagebrush Lizard, (Van Denburgh's Pine Lizard)

Südlicher Beifußleguan

Sydlig Bynkeleguan

1896 Sceloporus vandenburgianus Cope
Sceloporus graciosus vandenburgianus Camp 1916

Remarks: The phylogenetic analysis of Wiens & Reeder (1997) supported the specific recognition of vandenburgianus suggested by Collins (1991).
Distribution: Mexico (Baja California Norte), USA (California).


Sceloporus variabilis

Southern Rose-bellied Lizard, (Common Rose-bellied Lizard)

Südlicher Rosabaüchiger Stachelleguan

Sydlig Rosabuget Tornleguan

1834 Sceloporus variabilis Wiegmann
Tropidolepis variabilis Duméril & Bibron 1837
1852 Sceloporus delicatissimus Hallowell (Mather & Sites 1985)
1852 Sceloporus marmoratus Hallowell (Dixon & Lemos-Espinal 2010)
Sceloporus variabilis marmoratus Smith 1934

Other common names:
marmoratus: Northern Rose-bellied Lizard
Remarks: Previously included olloporus, smithi and teapensis. Records from Belize refer to teapensis (Stafford & Meyer 2000). Mendoza-Quijano, Flores-Villela & Sites (1998) revalidated marmoratus as a separate species, followed by some authors (e.g., Wilson & Johnson 2010; Wilson, Mata-Silva & Johnson 2013; Nevárez-de los Reyes, Lazcano, García-Padilla, Mata-Silva, Johnson & Wilson 2016), but this was not regarded as justified by Crother (ed.) (2008), Dixon & Lemos-Espinal (2010), and Lemos-Espinal (2015). Records from El Salvador, Honduras, Nicaragua, and Costa Rica refer to olloporus (Smith, Pérez-Higareda & Chiszar 1993; Herrera, Henríquez & Greenbaum 2007). Although included for Jalisco by some authors (e.g., Auth, Smith, Brown & Lintz 2000; Cruz-Sáenz, Guerrero, Lazcano & Télles-López 2009), presence in the state is considered undocumented herein, following Cruz-Sáenz, Muñoz-Nolasco, Mata-Silva, Johnson, García-Padilla & Wilson (2017), who did not list the species for Jalisco.
Distribution: Guatemala, Mexico (Chiapas, Coahuila, Guanajuato, Hidalgo, Nuevo Leon, Oaxaca, Puebla, Queretaro, San Luis Potosi, Tabasco, Tamaulipas, Veracruz), USA (Texas).


Sceloporus virgatus

Striped Plateau Lizard

Gestreifter Plateau-Stachelleguan

Stribet Plateau-leguan

1938 Sceloporus undulatus virgatus Smith
Sceloporus virgatus Cole 1963

Distribution: Mexico (Chihuahua, Sinaloa, Sonora), USA (Arizona, New Mexico).


Sceloporus woodi

Florida Scrub Lizard

Florida-Stachelleguan

Florida-tornleguan

1918 Sceloporus woodi Stejneger
Sceloporus undulatus woodi Burt 1935

Distribution: USA (Florida).


Sceloporus zosteromus

Baja California Spiny Lizard

Niederkalifornischer Stachelleguan

Baja California-tornleguan

1863 Sceloporus zosteromus Cope
Sceloporus clarkii zosteromus Cope 1875
Sceloporus magister zosteromus Linsdale 1932
1882 Sceloporus rufidorsum Yarrow (Grismer & McGuire 1996)
Sceloporus magister rufidorsum Linsdale 1932
1921 Sceloporus monserratensis Van Denburgh & Slevin (Grismer & McGuire 1996)
Sceloporus magister monserratensis Smith 1939
Sceloporus zosteromus monserratensis Bell, Smith & Chiszar 2003

Other common names:
monserratensis: Monserrat Island Spiny Lizard, Southern Spiny Lizard, Central Baja Spiny Lizard
rufidorsum: Red-backed Spiny Lizard, Mexican Desert Spiny Lizard
zosteromus: San Lucan Spiny Lizard
Distribution: Mexico (Baja California Norte [incl. Isla Cedros], Baja California Sur).