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Taxonomy of the family Colubridae
Bibliography of the genus Hebius
Biodiversity of the family Colubridae








Genus
Hebius

Oriental Equatorial Keelbacks, Oriental Water Snakes

Gebirgswassernattern

Ækvatorialasiatiske Kølsnoge

1913 Hebius Thompson (type species: Tropidonotus vibakari Boie 1826)
1934 Parahelicops Bourret (type species: Parahelicops annamensis Bourret 1934; syn. Kizirian, Nguyen, Ngo & Le 2018)
1934 Pararhabdophis Bourret (type species: Pararhabdophis chapaensis Bourret 1934; syn. Kizirian, Nguyen, Ngo & Le 2018)
1984 Paranatrix Mahendra (type species: Tropidonotus modestus Günther 1875; syn. Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014)
Contents: 44 species, of which 24 (54.5%) are endemic.
Remarks: Previously included in the genus Amphiesma (e.g., Williams & Wallach 1989; David, Vogel & Pauwels 1999). Revalidated as a separate genus by Guo, Zhu, Liu, Zhang, Li, Huang & Pyron (2014).
Distribution: S. and E. Asia, Malay Archipelago.
Reported from: Bangladesh, Brunei, Cambodia, China (Anhui, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Heilongjiang, Henan, Hong Kong, Hubei, Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shanxi, Sichuan, Tibet, Yunnan, Zhejiang), India (Andaman & Nicobar Islands [Nicobar Islands], Arunachal Pradesh, Assam, Goa, Himachal Pradesh, Karnataka, Kerala, Maharashtra, Meghalaya, Mizoram, Nagaland, Sikkim, Tamil Nadu, West Bengal), Indonesia (Buton, Kalimantan, Sula Islands [Sanana], Sulawesi, Sumatra), Japan (Honshu, Kyushu, Shikoku, Izu Islands, Ryukyu Islands [Okinawa Islands, Sakishima Islands (incl. Miyako Islands [incl. Miyako], Yaeyama Islands), Satsunan Islands (incl. Tokunoshima)]), Laos, Malaysia (Sabah, Sarawak, West Malaysia), Myanmar, Nepal, North Korea, Russia, South Korea, Taiwan, Thailand, Vietnam.

Hebius andreae

Quang Binh Keelback, (Andrea's Keelback)

Quang-Binh-Kielnatter

Quang Binh-kølsnog

2006 Amphiesma andreae Ziegler & Quyet
Hebius andreae Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: Vietnam.


Hebius annamensis

Annam Keelback

Annam-Kielnatter

Annam-kølsnog

1934 Parahelicops annamensis Bourret
Opisthotropis annamensis Smith 1943
Amphiesma annamense Teynié & David 2010
Hebius annamensis Kizirian, Nguyen, Ngo & Le 2018

Remarks: Previously placed in a separate genus, Parahelicops, which was synonymized with Hebius by Kizirian, Nguyen, Ngo & Le (2018).
Distribution: Laos, Vietnam.


Hebius arquus

Arch-lipped Keelback

Bogenlippen-Kielnatter

Buelæbet Kølsnog

2010 Amphiesma arquus David & Vogel
Hebius arquus Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Remarks: Exact distribution within Borneo unknown, but possibly Kalimantan (Indonesia).
Distribution: Borneo.


Hebius atemporalis

Tonkin Keelback, (Mountain Keelback)

Tonkin-Kielnatter

Tonkin-kølsnog

1934 Natrix atemporalis Bourret
Amphiesma atemporalis Malnate 1960
Hebius atemporale Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: China (Guangdong, Guangxi, Guizhou, Hong Kong, Yunnan), Laos, Vietnam.


Hebius beddomei

Nilgiri Keelback, (Beddome's Keelback)

Nilgiri-Kielnatter

Nilgiri-kølsnog

1863 Spilotes vittatus Beddome [not Coluber vittatus Linnaeus 1758]
1864 Tropidonotus beddomei Günther
Rhabdophis beddomei Wall 1923
Natrix beddomei Smith 1943
Amphiesma beddomei Malnate 1960
Hebius beddomei Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: India (Goa, Karnataka, Kerala, Maharashtra, Tamil Nadu).


Hebius bitaeniatus

Two-striped Keelback, Kutkai Keelback

Zweistreifige Kielnatter

Tostribet Kølsnog

1925 Natrix bitæniata Wall
Amphiesma bitaeniata Malnate 1960
Hebius bitaeniatum Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Remarks: Previously regarded as a synonym of parallelum (e.g., Smith 1943).
Distribution: China (Guangdong, Guangxi, Guizhou, Hunan, Yunnan), Myanmar, Thailand, Vietnam.


Hebius boulengeri

White-browed Keelback, Tai-yong Keelback, (Boulenger's Keelback)

Weißbrauen-Kielnatter

Hvidbrynet Kølsnog

1937 Natrix boulengeri Gressitt
Amphiesma boulengeri Malnate 1960
Hebius boulengeri Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Remarks: Previously confused with khasiense and modestum, and, more rarely, inas and parallelum (David, Vogel & Rooijen 2013).
Distribution: Cambodia, China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Jiangxi, Yunnan), Laos, Thailand, Vietnam.


Hebius celebicus

Sulawesi Keelback

Sulawesi-Kielnatter

Sulawesi-kølsnog

1878 Styporhynchus celebicus Peters & Doria
Tropidonotus celebicus Boulenger 1893
Natrix celebica Barbour 1912
Amphiesma celebica Malnate 1960
Hebius celebicum Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: Indonesia (Buton, Sula Islands [Sanana], Sulawesi).


Hebius chapaensis

Chapa Flat-nosed Snake, Sapa Flat-nosed Snake

Chapa-Wassernatter

Chapa-kølsnog

1934 Pararhabdophis chapaensis Bourret
Hebius chapaensis Kizirian, Nguyen, Ngo & Le 2018

Remarks: Previously placed in a separate genus, Pararhabdophis, which was synonymized with Hebius by Kizirian, Nguyen, Ngo & Le (2018) and Ren, Wang, Nguyen, Hoang, Zhong, Jiang, Guo & Li (2018).
Distribution: China (Yunnan), Laos, Vietnam.


Hebius clerki

Kachin Keelback

Kachin-Wassernatter

Kachin-kølsnog

1925 Natrix clerki Wall
Amphiesma clerki David, Agarwal, Athreya, Mathew, Vogel & Mistry 2015
Hebius clerki (new combination, this work)

Remarks: Previously regarded as a synonym of parallelus (e.g., Smith 1943). Revalidated as a separate species by David, Agarwal, Athreya, Mathew, Vogel & Mistry (2015).
Distribution: China (Yunnan), India (Arunachal Pradesh, Nagaland, Sikkim, West Bengal), Myanmar.


Hebius concelarus

Miyako Islands Keelback

Miyako-Kielnatter

Miyako-kølsnog

1963 Amphiesma pryeri concelara Malnate
Amphiesma concelarum Ota & Iwanaga 1997
Hebius concelarum Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: Japan (Ryukyu Islands: Sakishima Islands: Miyako Islands: Miyako).


Hebius craspedogaster

Kuatun Keelback

Kuatun-Kielnatter

Kuatun-kølsnog

1899 Tropidonotus craspedogaster Boulenger
Natrix craspedogaster Stejneger 1907
Amphiesma craspedogaster Malnate 1960
Hebius craspedogaster Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014
1922 Tropidonotus gastrotaenia Werner (Zhao & Adler 1993)
1931 Natrix chrysarga chekiangensis Mell (Zhao & Adler 1993)

Distribution: China (Anhui, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Shaanxi, Shanxi, Sichuan, Tibet, Yunnan, Zhejiang), Vietnam.


Hebius deschauenseei

Northern Keelback, (Deschauensee's Keelback)

Nördliche Kielnatter

Nordlig Kølsnog

1934 Natrix deschauenseei Taylor
Amphiesma deschauenseei Malnate 1960
Hebius deschauenseei Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Remarks: Previously regarded as a synonym of modestum (e.g., Smith 1943). Status of Vietnamese records is uncertain (Nguyen, Ho & Nguyen 2009).
Distribution: China (Guizhou, Yunnan), Thailand, Vietnam.


Hebius flavifrons

White-fronted Keelback, White-fronted Water Snake, White-nosed Keelback, (Sabah Keelback)

Weißstirn-Kielnatter, Weißkopf-Gebirgswassernatter

Hvidpandet Kølsnog

1887 Tropidonotus flavifrons Boulenger
Natrix flavifrons Barbour 1912
Amphiesma flavifrons Malnate 1960
Hebius flavifrons Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: Brunei, Indonesia (Kalimantan), Malaysia (Sabah, Sarawak).


Hebius frenatus

Bridled Keelback

Zügel-Kielnatter

Tøjle-kølsnog

1923 Natrix frenata Dunn
Amphiesma frenata Stuebing 1991
Hebius frenatum Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Remarks: Placed in Amphiesma on the basis of Stuebing (1991), Manthey & Grossmann (1997), and Das (2006, 2010). Not included in Amphiesma by David, Vogel & Pauwels (1998).
Distribution: Malaysia (Sarawak).


Hebius groundwateri

Ranong Keelback, (Groundwater’s Keelback)

Ranong-Kielnatter

Ranong-kølsnog

1922 Natrix groundwateri Smith "1921"?#
Amphiesma groundwateri Malnate 1960
Hebius groundwateri Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: Thailand.


Hebius inas

Malayan Mountain Keelback, Mount Inas Keelback, Gunung Inas Keelback

Malaiische Kielnatter, Malaiische Gebirgswassernatter

Malayisk Bjergkølsnog

1901 Tropidonotus inas Laidlaw
Natrix inas Smith 1930
Amphiesma inas Malnate 1960
Hebius inas Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Remarks: A record from Sumatra (Teynié, David & Ohler 2010) was based on the second known specimen of kerinciense (David, Vogel & Rooijen 2013). In Thailand, limited to the peninsular region. Records from north of the Isthmus of Kra refer to boulengeri or khasiense (David, Vogel & Rooijen 2013).
Distribution: Malaysia (West Malaysia), Thailand.


Hebius ishigakiensis

Yaeyama Water Snake

Yaeyama-Kielnatter

Yaeyama-kølsnog

1960 Natrix pryeri ishigakiensis Malnate & Munsterman
Amphiesma pryeri ishigakiensis Welch 1988
Amphiesma ishigakiense Ota & Iwanaga 1997
Hebius ishigakiense Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: Japan (Ryukyu Islands: Sakishima Islands [Miyako Islands, Yaeyama Islands]).


Hebius johannis

South Chinese Keelback

Südchinesische Kielnatter

Sydkinesisk Kølsnog

1908 Tropidonotus johannis Boulenger
Natrix johannis Schmidt 1927
Amphiesma johannis Malnate 1960
Hebius johannis Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: China (Guizhou, Sichuan, Yunnan).


Hebius kerinciensis

Kerinchi Keelback, Gunung Kerinchi Keelback

Kerinchi-Kielnatter

Kerinchi-kølsnog

2003 Amphiesma kerinciense David & Das
Hebius kerinciense Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: Indonesia (Sumatra).


Hebius khasiensis

Khasi Hills Keelback

Khasi-Hills-Kielnatter

Khasi Hills-kølsnog

1890 Tropidonotus khasiensis Boulenger
Natrix khasiensis Wall 1923
Amphiesma khasiensis Malnate 1960
Sinonatrix khasinensis [sic] Murthy 1994
Hebius khasiense Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014
1925 Natrix gilhodesi Wall (Smith 1943)

Remarks: Regarded as a synonym of modestum by Mahendra (1984). Validity confirmed by David, Vogel & Rooijen (2013). Now restricted to northestern India, Burma, western Yunnan, northern Laos, and northern and western Thailand. Reports from other parts of Laos and Thailand, and from southeastern China, Cambodia, and Vietnam refer to boulengeri (Grismer, Neang, Chav, Wood, Oaks, Holden, Grismer, Szutz & Youmans 2008; David, Vogel & Rooijen 2013). In India, the occurrence of khasiense in Mizoram, Sikim, and West Bengal was not mentioned by David, Vogel & Rooijen (2013), possibly due to a lack of voucher specimens.
Distribution: China (Guangxi, Tibet, Yunnan), India (Arunachal Pradesh, Assam, Meghalaya, Mizoram, Nagaland, Sikkim, West Bengal), Laos, Myanmar, Thailand.


Hebius lacrima

Mourning Keelback, Crying Keelback

Trauern-Kielnatter

Sørge-kølsnog

2019 Hebius lacrima Purkayastha & David

Distribution: India (Arunachal Pradesh).





Hebius leucomystax

White-lipped Keelback

Weißlippen-Kielnatter

Hvidlæbet Kølsnog

2007 Amphiesma leucomystax David, Bain, Nguyen, Orlov, Vogel, Vu & Ziegler
Hebius leucomystax Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: Cambodia, Laos, Thailand, Vietnam.


Hebius metusius

Sichuan Keelback, Wa Shan Keelback

Sichuan-Kielnatter

Sichuan-kølsnog

1990 Amphiesma metusia Inger, Zhao, Schaffer & Wu
Hebius metusium Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: China (Sichuan).


Hebius miyajimae

Taiwanese Striped Keelback, (Maki's Keelback)

Taiwanische Gestreifte Kielnatter

Taiwansk Stribet Kølsnog

1931 Natrix miyajimae Maki
Amphiesma miyajimae Zhao & Jiang 1986
Hebius miyajimae Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: Taiwan.


Hebius modestus

Modest Keelback, (Günther's Keelback)

Indische Gebirgs-Kielnatter

Indisk Bjerg-kølsnog

1875 Tropidonotus modestus Günther
Nerodia modesta Wall 1923
Natrix modesta Bourret 1936
Amphiesma modesta Malnate 1960
Paranatrix modesta Mahendra 1984
Sinonatrix modesta Murthy 1994
Hebius modestum Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Remarks: Previously included deschauenseei (e.g., Smith 1943) and khasiense (e.g., Mahendra 1984). According to David, Vogel & Pauwels (2005), the presence of modestum in Vietnam needs confirmation, although it was listed by Nguyen, Ho & Nguyen (2009). Records from Thailand (e.g., Smith 1943; Cox, Dijk, Nabhitabhata & Thirakhupt 1998) refer to khasiense (David, Vogel & Rooijen 2013). A record from Himachal Pradesh (Agrawal 1979) is regarded as doubtful.
Distribution: Cambodia, China (Guangdong, Guangxi, Guizhou, Yunnan), India (Arunachal Pradesh, Assam, Meghalaya, Nagaland), Myanmar.


Hebius monticolus

Yellow-collared Forest Keelback, Hill Keelback, Green Western Keelback, (Forest Keelback)

Gelbhals-Wald-Kielnatter

Gulhalset Skovkølsnog

1853 Tropidonotus monticolus Jerdon
Rhabdophis monticola Wall 1923
Natrix monticola Smith 1943
Amphiesma monticola Malnate 1960
Hebius monticola Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: India (Goa, Karnataka, Kerala, Tamil Nadu).


Hebius nicobarensis

Nicobar Keelback

Nicobarische Kielnatter

Nicobarisk Kølsnog

1891 Tropidonotus nicobarensis Sclater
Natrix nicobarensis Wall 1910
Amphiesma nicobariensis Malnate 1960
Xenochrophis nicobariensis Biswas & Sanyal 1980
Paranatrix nicobarensis Mahendra 1984
Sinonatrix nicobarensis Murthy 1994
Hebius nicobarense Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014
1891 Tropidonotus nicobaricus Sclater (Smith 1891)

Distribution: India (Nicobar Islands).


Hebius octolineatus

Eight-striped Keelback

Achtstreifige Kielnatter

Ottestribet Kølsnog

1904 Tropidonotus octolineatus Boulenger
Natrix octolineatus Schmidt 1927
Amphiesma octolineata Malnate 1960
Hebius octolineatum Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014
1904 Tropidonotus pleurotænia Boulenger (Zhao & Adler 1993)
1904 Tropidonotus quadrilineatus Boulenger (Zhao & Adler 1993)
1913 Tropidonotus parallelus sublævis Despax (Zhao & Adler 1993)
1925 Natrix septemlineata Schmidt (Zhao & Adler 1993)
1932 Natrix nigroreticulata Chang (Zhao & Adler 1993)

Distribution: China (Guangxi, Guizhou, Sichuan, Yunnan).


Hebius optatus

Mount Omei Keelback, Mount Emei Keelback

Omei-Kielnatter

Omei-kølsnog

1966 Natrix optata Hu & Zhao
Amphiesma optata Zhao & Jiang 1986
Hebius optatum Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: China (Chongqing, Guangdong, Guangxi, Guizhou, Hunan, Sichuan), Vietnam.


Hebius parallelus

Parallel-striped Keelback, (Boulenger's Striped Keelback, Boulenger's Keelback, Yunnan Keelback)

Parallelstreifige Kielnatter

Parallelstribet Kølsnog

1890 Tropidonotus parallelus Boulenger
Natrix parallelus Wall 1923
Amphiesma parallela Malnate 1960
Paranatrix parallela Mahendra 1984
Sinonatrix parallelus Murthy 1994
Hebius parallelum Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Remarks: Previously included bitaeniatum and clerki, and confused with other species as well. Now restricted to India (Meghalaya, Nagaland). Older records from China (Yunnan), Myanmar, and other Indian states (Arunachal Pradesh, Sikkim, West Bengal) refer to clerki (David, Agarwal, Athreya, Mathew, Vogel & Mistry 2015), in spite of the fact that the type locality of parallelus is "Sikkim" (probably in error). Reported without exact locality from Nepal (e.g., Swan & Leviton 1962; Das 1994, 1996; Schleich & Kästle [eds.] 2002), but no specimens with locality are known from the country. Nepalese reports are most likely due to misidentifications of Herpetoreas sieboldi (Kramer 1977; Ao, David, Bordoloi & Ohler 2004; David, Vogel & Pauwels 2005). Some reports from China, i.e., Sichuan (e.g., Deuve 1970) and Yunnan (e.g., Zhao & Adler 1993) were based on misidentified octolineatum or sieboldi (Ao, David, Bordoloi & Ohler 2004). Records from Vietnam (e.g., David, Vogel & Pauwels 2005) refer to bitaeniatum (Nguyen, Ho & Nguyen 2009). Records from Cambodia (e.g., Bourret 1934, 1936) refer to boulengeri.
Distribution: India (Meghalaya, Mizoram, Nagaland), Nepal.


Hebius petersii

Sarawak Spotted Keelback, Yellow-lipped Water Snake, (Peter's Keelback)

Sarawak-Flecken-Kielnatter

Plettet Sarawak-kølsnog

1871 Tropidonotus maculatus Peters [not Tropidonotus maculatus Edeling 1864] (Iskandar & Colijn 2001)
1872 Tropidonotus sundanensis Peters [not Tropidonotus sundanensis Bleeker 1857] (Iskandar & Colijn 2001)
1893 Tropidonotus petersi Boulenger [substitute name for Tropidonotus maculatus Peters 1871]
Natrix petersi Smedley 1931
Amphiesma petersi Malnate 1960
Hebius petersii Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Remarks: Present in the past, but not confirmed recently from Singapore (Baker & Lim 2008). For unknown reasons, Wallach, Williams & Boundy (2014) listed Tropidonotus maculatus Peters 1871 as a synonym of sarawacensis.
Extinct in: Singapore.
Distribution: Indonesia (Sumatra), Malaysia (Sabah, Sarawak, West Malaysia).


Hebius popei

Chinese Mountain Keelback, (Pope's Mountain Keelback, Pope's Keelback)

Chinesische Gebirgs-Kielnatter

Kinesisk Bjergkølsnog

1925 Natrix popei Schmidt
Natrix vibakari popei Mell 1930
Amphiesma popei Malnate 1960
Hebius popei Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: China (Guangdong, Guangxi, Guizhou, Hainan, Hunan, Yunnan), Vietnam.


Hebius pryeri

Ryukyu Keelback, Ryukyu Water Snake

Ryukyu-Kielnatter

Ryukyu-kølsnog

1887 Tropidonotus pryeri Boulenger
Natrix pryeri Stejneger 1907
Amphiesma pryeri Malnate 1960
Hebius pryeri Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Remarks: Previously included concelarum and ishigakiense.
Distribution: Japan (Ryukyu Islands [Okinawa Islands, Sakishima Islands, Satsunan Islands (incl. Amami Islands [incl. Tokunoshima])]).


Hebius sanguineus

Red Mountain Keelback, (Smedley's Keelback)

Rote Kielnatter, Rote Gebirgswassernatter

Rød Bjergkølsnog

1932 Natrix sanguinea Smedley
Amphiesma sanguinae [sic] Malnate 1960
Hebius sanguineum Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: Malaysia (West Malaysia).


Hebius sangzhiensis

Hunan Mountain Keelback

Hunan-Kielnatter

Hunan-bjergkølsnog

2019 Hebius sangzhiensis Zhou, Sun, Qi, Lu, Lyu, Wang, Li & Ma

Distribution: China (Hunan).





Hebius sarasinorum

Mount Bonthain Keelback, (Sarasin's Keelback)

Bonthain-Kielnatter

Bonthain-bjergkølsnog

1896 Tropidonotus sarasinorum Boulenger
Natrix sarasinorum Barbour 1912
Amphiesma sarasinora Malnate 1960
Hebius sarasinorum Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: Indonesia (Sulawesi).


Hebius sarawacensis

Sarawak Mountain Keelback, Sarawak Grass Snake, Checker-bellied Keelsnake, (Sarawak Keelback)

Sarawak-Gebirgs-Kielnatter, Sarawak-Wassernatter

Sarawak-bjergkølsnog

1872 Tropidonotus sarawacensis Günther
Tropidonotus saravacensis [sic] Boulenger 1893
Natrix saravacensis [lapsus] Barbour 1912
Amphiesma saravacensis [sic] Malnate 1960
Rhabdophis sarawacensis Wallach, Williams & Boundy 2014
Hebius sarawacense Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Remarks: For unknown reasons, Wallach, Williams & Boundy (2014) assigned sarawacensis to the genus Rhabdophis. They also listed Tropidonotus maculatus Peters 1871 as a synonym of sarawacensis, although maculatus is a senior (but unavailable) synonym of petersi.
Distribution: Brunei, Indonesia (Kalimantan), Malaysia (Sabah, Sarawak, West Malaysia).


Hebius sauteri

Kosempo Keelback, (Sauter's Keelback)

Kosempo-Kielnatter

Kosempo-kølsnog

1909 Tropidonotus sauteri Boulenger
Natrix sauteri Pope 1935
Amphiesma sauteri Malnate 1960
Hebius sauteri Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014
1909 Natrix copei Van Denburgh (Zhao & Adler 1993)
1962 Amphiesma sauteri bourreti Malnate
1962 Amphiesma sauteri maximus Malnate

Distribution: China (Anhui, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hubei, Hunan, Jiangxi, Sichuan, Yunnan), Laos, Taiwan, Vietnam.


Hebius taronensis

Taron Keelback

Taron-Kielnatter

Taron-kølsnog

1940 Natrix venningi taronensis Smith
Amphiesma venningi taronensis Welch 1988
Amphiesma taronense Das 2010

Remarks: Previously regarded as a subspecies of venningi (e.g., Smith 1943). Revalidated by Das (2010), followed by others (Zug, Leviton, Vindum, Wogan & Koo 2010; Wallach, Williams & Boundy 2014). Not mentioned by Guo, Zhu, Liu, Zhang, Li, Huang & Pyron (2014).
Distribution: Myanmar.


Hebius venningi

Chin Hills Keelback, (Venning's Keelback)

Chin-Hills-Kielnatter

Chin Hills-kølsnog

1910 Natrix venningi Wall
Amphiesma venningi Malnate 1960
Paranatrix venningi Mahendra 1984
Hebius venningi Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014
1925 Natrix nigriventer Wall (Smith 1943)

Remarks: Previously included taronense. Records from Meghalaya, India, (e.g., Gayen 2001; Whitaker & Captain 2004) refer to Herpetoreas xenura (Mathew & Meetei 2004).
Distribution: Bangladesh, China (Guangxi, Yunnan), India (Arunachal Pradesh, Mizoram), Myanmar.


Hebius vibakari

Northeastern Forest Keelback, Hibakari, Vibakari Snake

Nordöstliche Wald-Kielnatter

Nordøstlig Skovkølsnog

1826 Tropidonotus vibakari Boie
Natrix vibakari Stejneger 1907
Hebius vibakari Thompson 1913
Amphiesma vibakari Malnate 1960
1876 Tropidonotus martensii Hilgendorf (Stejneger 1907)
1923 Natrix vibakari ruthveni Van Denburgh
Amphiesma vibakari ruthveni Malnate 1962
1928 Natrix vibakari nikolskii Emeljanov (Malnate 1962)
1986 Amphiesma vibakari danjoensis Toriba

Other common names:
danjoense: Danjo Hibakari
Remarks: A record from Sakhalin Island (Malnate 1962) appears to be erroneous. The island was not included in the distribution by neither Szczerbak (2003), nor Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov (2006).
Introduced to: Japan (Izu Islands: Miyakejima).
Distribution: China (Heilongjiang, Jilin, Liaoning), Japan (Honshu, Kyushu, Shikoku, Ryukyu Islands [Satsunan Islands]), North Korea, Russia, South Korea.


Hebius viperinus

Viper-like Keelback

Vipern-Kielnatter

Hugorme-kølsnog

1901 Xenochrophis viperinus Schenkel
Amphiesma viperina Malnate & Minton 1965
Hebius viperinum Guo, Zhu, Liu, Zhang, Li, Huang & Pyron 2014

Distribution: Indonesia (Sumatra).


Hebius yanbianensis

Yanbian Keelback

Yanbian-Kielnatter

Yanbian-kølsnog

2018 Hebius yanbianensis Liu, Zhong, Wang, Liu & Guo

Distribution: China (Sichuan).