Reptiles of the Sula Islands [Indonesia] Related checklists:  Indonesia Bibliography of the reptiles of Sula Islands Taxonomic history of the reptiles of Sula Islands Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus porosus Saltwater Crocodile Kopstein 1926 Kopstein 1927

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama Brongersma 1933 Kopstein 1926 Kopstein 1927 Genus Draco (Gliding Dragons)   Draco beccarii Kandari Gliding Dragon Kopstein 1927 Wermuth 1967  Family Gekkonidae (Typical Geckos) Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus jellesmae Kabaena Bent-toed Gecko Brongersma 1933 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Kopstein 1927 Genus Gekko (Tokays)   Gekko gecko Tokay Kopstein 1926 Kopstein 1927 [as verticillatus]   Gekko monarchus Spotted House Gecko Kopstein 1926 Kopstein 1927   Gekko vittatus White-striped Gecko Brongersma 1933 Kopstein 1927 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus frenatus Asian House Gecko Kopstein 1926 Kopstein 1927   Hemidactylus platyurus Flat-tailed Gecko Kopstein 1927  Family Scincidae (Skinks) Genus Emoia (Emo Skinks)   Emoia caeruleocauda Pacific Blue-tailed Skink Brongersma 1933 [as werneri] Brown 1991 Kopstein 1927 [as cyanura]   Emoia kuekenthali East Indonesian Slender Skink Brown 1991 Kopstein 1926 Kopstein 1927 Genus Eugongylus (Short-legged Giant Skinks)   * Eugongylus sulaensis Sula Giant Skink Kopstein 1927 Genus Eutropis (South Asian Sun Skinks)   Eutropis multifasciata Many-lined Sun Skink Kopstein 1926 Kopstein 1927   Eutropis rudis Three-keeled Sun Skink Brongersma 1933 [as multifasciata] Kopstein 1927 Genus Lamprolepis (Malay Archipelago Tree Skinks)   Lamprolepis smaragdina Emerald Tree Skink Brongersma 1933 Kopstein 1927 Genus Lygisaurus (Sahul Leaf-Litter Skinks)   Lygisaurus sp. (Australian-Papuan Leaf-Litter Skink) Kopstein 1926 [as novaeguineae] Kopstein 1927 [as novaeguineae] Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus nigrolabris Black-lipped Litter Skink Brongersma 1933   Sphenomorphus undulatus Wavy-backed Forest Skink Kopstein 1927 [as dammermani] Genus Tytthoscincus (Dwarf Leaf-litter Skinks)   * Tytthoscincus sananus Sanana Leaf-litter Skink Grismer, Wood, Quah, Anuar, Ngadi, Mohd-Izam & Ahmad 2017 Kopstein 1926  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   * Varanus melinus Quince Monitor Boehme & Ziegler 1997 Boehme 2003 Philipp, Boehme & Ziegler 1999 Weijola, Donnellan & Lindquist 2016 Ziegler & Boehme in Pianka, King & King (eds.) 2004 Ziegler, Schmitz, Koch & Boehme 2007   * Varanus obor Sago Monitor Boehme 2003 [as V. melinus] Weijola & Sweet 2010 Ziegler & Boehme in Pianka, King & King (eds.) 2004 [as V. melinus]   Varanus salvator Asian Water Monitor Mertens 1963 Weijola & Sweet 2010

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake Bosch 1985 Haas 1950 Iskandar & Colijn 2001 Kopstein 1927 Lang 2013 Mertens 1930

Genus Boiga (Australasian Cat Snakes)

Boiga flavescens Yellow Cat Snake

Haas 1950 [as i. irregularis] Iskandar & Colijn 2001 [as irregularis] Kopstein 1927 [as irregularis] Lang 2013 [as irregularis] Orlov & Ryabov 2002 [as irregularis]

Genus Dendrelaphis (Bronzebacks)

Dendrelaphis marenae Philippine Painted Bronzeback

Lang 2013

Dendrelaphis pictus Southern Painted Bronzeback

Haas 1950 Iskandar & Colijn 2001 Kopstein 1927 Lang 2011

Genus Hebius (Oriental Equatorial Keelbacks)

Hebius celebicus Sulawesi Keelback

Bosch 1985 Haas 1950 Iskandar & Colijn 2001 Koch 2012 Kopstein 1926 Kopstein 1927 Lang 2013 Wallach, Williams & Boundy 2014

Genus Oligodon (Kukri Snakes)

Oligodon waandersi Makassar Kukri Snake

Bosch 1985 Haas 1950 Iskandar & Colijn 2001 Koch 2012 Kopstein 1927 Lang 2013

Family Cylindrophiidae (Asian Pipe Snakes)

Genus Cylindrophis (Oriental Pipe Snakes)

Cylindrophis melanotus Black Pipe Snake

Bernstein, Bauer, McGuire, Arida, Kaiser, Kieckbusch & Mecke 2020 Bosch 1985 Iskandar & Colijn 2001 Koch 2012 Lang 2013 McDiarmid, Campbell & Toure 1999

Cylindrophis ruffus Red-tailed Pipe Snake

Bosch 1985 Haas 1950 Kopstein 1926 Kopstein 1927 McDiarmid, Campbell & Toure 1999

Family Homalopsidae (Australasian Mud Snakes)

Genus Cerberus (Dog-faced Water Snakes)

Cerberus schneiderii South East Asian Dog-faced Water Snake

Iskandar & Colijn 2001 [as C. rynchops] Kopstein 1927 [as C. rynchops] Lang 2013

Family Lamprophiidae (House Snakes, etc.)

Genus Psammodynastes (Asian Mock Vipers)

Psammodynastes pulverulentus Dusky Mock Viper

Bosch 1985 Haas 1950 Iskandar & Colijn 2001 Kopstein 1927 Lang 2013

Family Pythonidae (Pythons)

Genus Malayopython (Malayan Pythons)

Malayopython reticulatus Reticulated Python

David, Pauwels, Lays & Lenglet 2006 Iskandar & Colijn 2001 Kopstein 1927 Lang 2013

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle Kopstein 1926 Kopstein 1927 Mertens 1930