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© Rune Midtgaard


Taxonomy of the family Colubridae
Bibliography of the genus Lycodon
Biodiversity of the family Colubridae








Genus
Lycodon

Oriental Wolf Snakes, Asian Wolf Snakes

Wolfzahnnattern, (Großzahnnattern [syn. Dinodon])

Orientalske Ulvesnoge

1826 Lycodon Boie in Fitzinger (type species: Coluber aulicus Linnaeus 1758)
1826 Nympha Fitzinger (type species: Coluber nympha Daudin 1803; syn. Williams & Wallach 1989)
1830 Cercaspis Wagler (type species: Hurria carinata Kuhl 1820; syn. Pyron, Kandambi, Hendry, Pushpamal, Burbrink & Somaweera 2013)
1830 Ophites Wagler (type species: Lycodon subcinctus Boie 1827)
1853 Odontomus Dumeril [not Odontomus Kirby in Richardson 1837 (Insecta)] (type species: Coluber nympha Daudin 1803)
1854 Dinodon Dumeril, Bibron & Dumeril (type species: Dinodon cancellatum Dumeril 1853; syn. Guo, Zhang, Liu, Li, Pyron Jiang & Burbrink 2013; Siler, Oliveiros, Santanen & Brown 2013)
1858 Dryocalamus Günther (type species: Dryocalamus tristrigatus Günther 1858; syn. Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016)
1858 Hemidipsas Günther (type species: Hemidipsas ocellata = Odontomus sub-annulatus Dumeril, Bibron & Dumeril 1854)
1858 Leptorhytaon Günther (type species: Coluber jara Shaw 1802)
1858 Tetragonosoma Günther (type species: Lycodon effraenis Cantor 1847)
1860 Eumesodon Cope (type species: Dinodon semicarinatus Mell 1922; syn. Guo, Zhang, Liu, Li, Pyron Jiang & Burbrink 2013 [by implication])
1860 Euprotodon Cope (type species: Coluber aulicus Linnaeus 1758)
1861 Lepidocephalus Hallowell (type species: Lepidocephalus fasciatus Hallowell 1860; syn. Guo, Zhang, Liu, Li, Pyron Jiang & Burbrink 2013 [by implication])
1862 Hydrophobus Günther (type species: Hydrophobus semifasciatus Günther 1862)
1864 Nymphophidium Günther (type species: Nymphodidium maculatum Günther 1864)
1868 Tytleria Theobald (type species: Tytleria hypsirhinoides Theobald 1868)
1878 Ulupe Blanford (type species: Ulupe davisonii Blanford 1878)
1893 Anoplophallus Cope (type species: Lycodon subcinctus Boie 1827)
1900 Haplonodon Griffin (type species: Haplonodon philippinensis Griffin 1910; syn. Williams & Wallach 1989)
1900 Lepturophis Boulenger (type species: Lepturophis borneensis Boulenger 1900; syn. Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016)
1925 Adiastema Werner (type species: Adiastema cervinum Werner 1925; syn. Guo, Zhang, Liu, Li, Pyron Jiang & Burbrink 2013 [by implication])
Contents: 66 species, of which 37 (56.1%) are endemic.
Other common names:
Dinodon: Big-toothed Snakes, Large-toothed Snakes, Odd-toothed Snakes, False Wolf Snakes
Dryocalamus: Bridle Snakes, Bridal Snakes
Remarks: Closely related to the genus Stegonotus. The taxonomic relationship between the two genera is unclear (Kaiser, Kaiser & O'Shea 2018).
Distribution: Asia, Malay Archipelago.
Reported from: Bangladesh, Bhutan, Brunei, Cambodia, China (Anhui, Beijing, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hebei, Heilongjiang, Henan, Hong Kong [incl. Shek Kwu Chau Island], Hubei, Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Shanxi, Sichuan, Tibet, Yunnan, Zhejiang), East Timor (incl. Atauro Island), India (Andaman & Nicobar Islands [Andaman Islands, Nicobar Islands], Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Chhattisgarh, Delhi, Goa, Gujarat, Haryana, Himachal Pradesh, Jammu & Kashmir, Jharkhand, Karnataka, Kerala, Lakshadweep, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Punjab, Rajasthan, Sikkim, Tamil Nadu, Telangana, Tripura, Uttar Pradesh, Uttarakhand, West Bengal), Indonesia (Adonara, Alor, Ambon, Babar Islands, Bali, Banyak Islands, Ambon, Banda Islands, Bawean Islands, Buru, Buton, Ende, Flores, Java, Kabaena, Kalao, Kalimantan, Karimunjawa Islands, Kisar, Komodo, Lembata, Lombok, Muna, Natuna Islands, Nias, Padar, Pantar, Penida, Riau Islands, Rinca, Rote, Savu, Selayar, Semau, Seram, Serua, Siberut, Simeulue, Sulawesi, Sumatra, Sumba, Sumbawa, West Timor, Wetar), Iran, Japan (Hokkaido, Honshu, Kyushu, Shikoku, Tsushima Islands, Izu Islands, Ryukyu Islands [Okinawa Islands, Sakishima Islands (incl. Miyako Islands, Yaeyama Islands), Satsunan Islands], Volcano Islands), Laos, Malaysia (Sabah, Sarawak, West Malaysia [incl. Langkawi Archipelago, Seribuat Archipelago (incl. Tioman)]), Maldives, Myanmar, Nepal, North Korea, Pakistan, Philippines (Babuyan Islands [Babuyan Claro, Balabac, Calayan, Camiguin Norte, Dalupiri], Bantayan, Batan Islands [Batan], Camiguin Sur, Carabao, Catanduanes, Cebu, Dinagat, Cuyo, Leyte, Luzon, Masbate, Mindanao, Mindoro, Negros, Palawan, Panay, Polillo, Samar, Sibuyan, Sulu Islands [Basilan], Tablas), Russia (incl. Kuril Islands), Singapore, South Korea, Sri Lanka, Taiwan, Tajikistan, Thailand, Turkmenistan, Uzbekistan, Vietnam (incl. Cat Ba Island).

Lycodon albofuscus

Slender Wolf Snake, Slender-tailed Wolf Snake

Schlank-Wolfzahnnatter

Sunda-ulvesnog

1854 Sphecodes albo-fuscus Dumeril, Bibron & Dumeril
Ophites albofuscus Günther 1858
Lycodon albofuscus Boulenger 1893
Lepturophis albofuscus David & Vogel 1996
1900 Lepturophis borneensis Boulenger (Wallach, Williams & Boundy 2014)

Remarks: Previously assigned to a separate, monotypic genus, Lepturophis, by some authors, whereas others have included the species in Lycodon (e.g., Lanza 1999; Grismer & Aun 2008; Das 2010). This synonymization was supported by Figueroa, KcKelvy, Grismer, Bell & Lailvaux (2016). Das & Yaakob (2007) listed Lepturophis borneensis as a valid species.
Distribution: Indonesia (Kalimantan, Nias, Sumatra), Malaysia (Sabah, Sarawak, West Malaysia [incl. Seribuat Archipelago (incl. Tioman)]), Thailand.


Lycodon alcalai

Batan Wolf Snake, (Alcala's Wolf Snake)

Batan-Wolfzahnnatter

Batan-ulvesnog

1994 Lycodon alcalai Ota & Ross

Distribution: Philippines (Babuyan Islands [Babuyan Claro, Calayan, Camiguin Norte], Batan Islands [Batan]).


Lycodon anamallensis

Anaimalai Wolf Snake

Anaimalai-Wolfzahnnatter

Anaimalai-ulvesnog

1864 Lycodon anamallensis Günther
1950 Lycodon osmanhilli Taylor (Ganesh & Vogel 2018)
Lycodon aulicus osmanhilli Schleich & Kästle (eds.) 2002

Other common names:
osmanhilli: Flowered Wolf Snake
Remarks: Previously regarded as a synonym of aulicus. Revalidated as a separate species by Ganesh & Vogel (2018).
Distribution: India (Kerala, Tamil Nadu), Sri Lanka.





Lycodon aulicus

Indian Wolf Snake, Common Wolf Snake, (White-spotted Wolf Snake, House Snake)

Indische Wolfzahnnatter, Gewöhnliche Wolfzahnnatter

Indisk Ulvesnog

1758 Coluber aulicus Linnaeus
Lycodon aulicus Boie 1827
Ophites aulicus Wall 1921
1839 Lycodon subfuscus Cantor (Smith 1943)
1909 Lycodon aulicus oligozonatus Wall (Smith 1943)

Remarks: Previously included anamallensis. Sometimes considered conspecific with capucinus (see remarks under that species). Now considered restricted to South Asia (Wallach, Williams & Boundy 2014; O'Shea, Kusuma & Kaiser 2018). A large number of records of aulicus outside this area are therefore almost certainly based on capucinus. This includes a single record from the Comoro Archipelago which is regarded as doubtful, but indicates that it may have been introduced without having established a population (Hawlitschek, Bruckmann, Berger, Green & Glaw 2011, as aulicus).
Distribution: Bangladesh, Bhutan, China (Yunnan), India (Andhra Pradesh, Assam, Bihar, Chhattisgarh, Delhi, Goa, Gujarat, Haryana, Himachal Pradesh, Jammu & Kashmir, Jharkhand, Karnataka, Kerala, Lakshadweep, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Punjab, Rajasthan, Sikkim, Tamil Nadu, Tripura, Uttar Pradesh, Uttarakhand, West Bengal), Myanmar, Nepal, Pakistan, Sri Lanka.


Lycodon banksi

Khammouane Wolf Snake, (Banks' Wolf Snake)

Khammouane-Wolfzahnnatter

Khammouane-ulvesnog

2018 Lycodon banksi Luu, Bonkowski, Nguyen, Le, Calame & Ziegler

Distribution: Laos.





Lycodon bibonius

Camiguin Norte Wolf Snake, (Crombie's Wolf Snake)

Camiguin-Norte-Wolfzahnnatter

Camiguin Norte-ulvesnog

1994 Lycodon bibonius Ota & Ross

Distribution: Philippines (Babuyan Islands [Babuyan Claro, Camiguin Norte]).


Lycodon butleri

Malaysian Mountain Wolf Snake, (Butler's Wolf Snake)

Malaiische Gebirgs-Wolfzahnnatter

Malaysisk Bjerg-ulvesnog

1900 Lycodon Butleri Boulenger

Distribution: Malaysia (West Malaysia), Thailand.


Lycodon capucinus

Island Wolf Snake, Oriental House Snake, Asian House Snake, Indian Wolf Snake, (Common Asian Wolf Snake)

Kapuzen-Wolfzahnnatter

Kapuciner-ulvesnog

1827 Lycodon capucinus Boie
Lycodon aulicus capucinus Boettger 1898
1827 Lycodon unicolor Boie (Iskandar & Colijn 2001)
Boaedon unicolor Duméril, Bibron & Duméril 1854
1839 Lycodon atropurpureus Cantor (Iskandar & Colijn 2001)

Remarks: Status uncertain (Siler, Oliveiros, Santanen & Brown 2013). Lanza (1999) did not consider existing evidence strong enough to separate capucinus from aulicus, however, he continued to treat them in separate accounts. Regarded also by some other authors as a subspecies or a synonym of aulicus (e.g., Smith 1943; Ota & Ross 1994; Jackson & Fritts 2004), but as a separate species by others (e.g., Taylor 1965; David & Vogel 1996; Wallach, Williams & Boundy 2014; O'Shea, Kusuma & Kaiser 2018). The distributions of the two species apparently overlap in Myanmar, but the exact range limits are unclear (O'Shea, Kusuma & Kaiser 2018).
Previously included hypsirhinoides (e.g., Iskandar & Colijn 2001) to which records of capucinus from the Andaman Islands refer (Vogel & Harikrishnan 2013). The Maldives population is provisionally assigned to capucinus herein, although it is unclear whether it could represent aulicus (O'Shea, Kusuma & Kaiser 2018). The species has also been listed for Gansu and Hunan (Wallach, Williams & Boundy 2014). The first record is almost certainly an error or based on an introduction (O'Shea, Kusuma & Kaiser 2018), the second record is provisionally regarded as doubtful herein, since it is also quite far from the nearest records listed below. Philippine populations may have been introduced (Lanza 1999). Leviton, Siler, Weinell & Brown (2018) did not include Sibuyan in the distribution of the species. Presence in the Comoro Archipelago is regarded as doubtful, although a single museum specimen indicates that the species may have been introduced without having established a population (Hawlitschek, Bruckmann, Berger, Green & Glaw 2011, as aulicus).
Introduced to: Christmas Island, Mauritius, Micronesia (Pohnpei), Reunion.
Distribution: Cambodia, China (Fujian, Guangdong, Hainan, Hong Kong [incl. Shek Kwu Chau Island]; see remarks), East Timor (incl. Atauro Island), Indonesia (Adonara, Alor, Ambon, Babar Islands, Bali, Banda Islands, Bawean Islands, Buru, Buton, Ende, Flores, Java, Kabaena, Kalao, Karimunjawa Islands, Kisar, Komodo, Lembata, Lombok, Muna, Padar, Pantar, Penida, Rinca, Rote, Savu, Selayar, Semau, Seram, Serua, Sulawesi, Sumatra, Sumba, Sumbawa, West Timor, Wetar), Laos, Malaysia (Sabah, West Malaysia [incl. Seribuat Archipelago], Maldives, Myanmar, Philippines (Bantayan, Camiguin Sur, Carabao, Cebu, Cuyo, Dinagat, Leyte, Luzon, Masbate, Mindanao, Mindoro, Negros, Palawan, Panay, Samar, Sibuyan, Sulu Islands, Tablas), Singapore, Thailand, Vietnam.


Lycodon cardamomensis

Cardamom Mountains Wolf Snake

Cardamom-Wolfzahnnatter

Cardamom-ulvesnog

2002 Lycodon cardamomensis Daltry & Wüster

Distribution: Cambodia, Thailand, Vietnam.


Lycodon carinatus

Sri Lanka Wolf Snake

Srilankanische Wolfzahnnatter

Srilankansk Ulvesnog

1820 Hurria carinata Kuhl
Cercaspis carinata Günther 1864
Lycodon carinatus Boulenger 1890

Distribution: Sri Lanka.


Lycodon cathaya

Huaping Wolf Snake

Huaping-Wolfzahnnatter

Huaping-ulvesnog

2020 Lycodon cathaya Wang, Qi, Lyu, Zeng & Wang

Distribution: China (Guangxi).





Lycodon cavernicolus

Cave Wolf Snake

Höhlen-Wolfzahnnatter

Hule-ulvesnog

2014 Lycodon cavernicolus Grismer, Quah, Anuar, Muin, Wood & Nor

Distribution: Malaysia (West Malaysia).


Lycodon chithrasekarai

Sri Lankan Bridle Snake

Srilankanische Zügelnatter

Srilankansk Tøjlesnog

2020 Dryocalamus chithrasekarai Wickramasinghe, Vidanapathirana, Pushpamal & Wickramasinghe
Lycodon chithrasekarai Midtgaard 2020 (this work)

Distribution: Sri Lanka.





Lycodon chrysoprateros

Dalupiri Wolf Snake, (Ross's Wolf Snake)

Dalupiri-Wolfzahnnatter

Dalupiri-ulvesnog

1994 Lycodon chrysoprateros Ota & Ross

Distribution: Philippines (Babuyan Islands: Dalupiri).


Lycodon davidi

Vientiane Wolf Snake, (David's Wolf Snake)

Vientiane-Wolfzahnnatter

Vientiane-ulvesnog

2012 Lycodon davidi Vogel, Nguyen, Kingsada & Ziegler

Distribution: Laos.


Lycodon davisonii

Burmese Bridle Snake, (Blanford's Bridal Snake, Common Bridal Snake)

Burmesische Zügelnatter

Burmesisk Tøjlesnog

1878 Ulupe davisonii Blanford
Hydrophobus davisonii Boulenger 1890
Dryocalamus davisonii Boulenger 1893
Lycodon davisonii Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016

Remarks: Listed for China by Bain & Hurley (2011), but no other records have been found to verify this.
Distribution: Bhutan, Cambodia, Laos, Myanmar, Thailand, Vietnam.


Lycodon deccanensis

Deccan Wolf Snake

Dekkan-Wolfzahnnatter

Deccan-ulvesnog

2020 Lycodon deccanensis Ganesh, Deuti, Punith, Achyuthan, Mallik, Adhikari & Vogel

Remarks: Previously regarded as a population of travancoricus.
Distribution: India (Andhra Pradesh, Karnataka, Tamil Nadu).





Lycodon dumerilii

Mindanao Wolf Snake, (Dumeril's Wolf Snake)

Mindanao-Wolfzahnnatter

Mindanao-ulvesnog

1858 Lycodon mülleri Günther [not Lycodon mülleri Dumeril, Bibron & Dumeril 1854]
1893 Stegonotus dumerilii Boulenger
Lycodon dumerilii Leviton 1965
1922 Dryocalamus mccroryi Taylor (Leviton 1962)
Dryocalamus mccroryae Michels & Bauer 2004 [justified emendation]

Distribution: Philippines (Dinagat, Mindanao, Samar, Sulu Islands [Basilan]).


Lycodon effraenis

Scarce Wolf Snake, Brown Wolf Snake

Braune Wolfzahnnatter

Brun Ulvesnog

1847 Lycodon effraenis Cantor
Tetragonosoma effrene Günther 1864

Distribution: Brunei, Indonesia (Kalimantan, Sumatra), Malaysia (Sarawak, West Malaysia [incl. Seribuat Archipelago (incl. Tioman)]), Thailand.


Lycodon fasciatus

Yellow-banded Wolf Snake, Indian Banded Wolf Snake, (Anderson's Wolf Snake, Banded Wolf Snake)

Gelbbändige Wolfzahnnatter

Gulbåndet Ulvesnog

1879 Ophites fasciatus Anderson
Lycodon fasciatus Boulenger 1890
1922 Dinodon yunnanensis Werner (Zhao & Adler 1993)

Remarks: Previously confused with gammiei (e.g., Wall 1911). Validity as a separate species confirmed by Mistry, Vogel & Tillack (2007), although recognized also by other, previous works.
Distribution: Bangladesh, Bhutan, China (Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Henan, Hubei, Hunan, Shaanxi, Sichuan, Tibet, Yunnan), India (Arunachal Pradesh, Meghalaya, Mizoram, Nagaland, West Bengal), Laos, Myanmar, Thailand, Vietnam.


Lycodon fausti

Panay Wolf Snake, (Faust's Wolf Snake)

Panay-Wolfzahnnatter

Panay-ulvesnog

2002 Lycodon fausti Gaulke

Distribution: Philippines (Panay).


Lycodon ferroni

Samar Wolf Snake, (Ferron's Wolf Snake)

Samar-Wolfzahnnatter

Samar-ulvesnog

1999 Lycodon ferroni Lanza

Distribution: Philippines (Samar).


Lycodon flavicollis

Yellow-collared Wolf Snake

Halsband-Wolfzahnnatter

Halsbånd-ulvesnog

2007 Lycodon flavicollis Mukherjee & Bhupathy

Distribution: India (Karnataka, Kerala, Tamil Nadu).


Lycodon flavomaculatus

Yellow-spotted Wolf Snake

Gelbfleckige Wolfzahnnatter

Gulplettet Ulvesnog

1907 Lycodon flavomaculatus Wall
Ophites flavomaculatus Wall 1923

Remarks: Mukherjee & Bhupathy (2007) suggested that some previous records of flavomaculatus may refer to flavicollis (e.g., Murthy 1991).
Distribution: India (Chhattisgarh, Gujarat, Karnataka, Madhya Pradesh, Maharashtra, Tamil Nadu, Telangana).


Lycodon flavozonatus

Yellow Big-toothed Snake

Gelbe Wolfzahnnatter

Gul Ulvesnog

1922 Dinodon semicarinatus Mell (not Eumesodon semicarinatus Cope 1860)
1928 Dinodon flavozonatum Pope
Lycodon flavozonatum Guo, Zhang, Liu, Li, Pyron Jiang & Burbrink 2013

Distribution: China (Anhui, Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hunan, Jiangxi, Sichuan, Zhejiang), Myanmar.


Lycodon futsingensis

Fujian Wolf Snake

Fujian-Wolfzahnnatter

Fujian-ulvesnog

1928 Dinodon futsingensis Pope
Lycodon futsingensis Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler 2009

Remarks: Previously regarded as a synonym of ruhstrati (e.g., Zhao & Adler 1993). Revalidated by Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler (2009). Exact distribution remains unclear, as many literature records cannot with certainty be assigned to futsingensis or ruhstrati abditus (Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler 2009).
Distribution: China (Fujian, Guangdong, Hong Kong), Laos, Vietnam.


Lycodon gammiei

Sikkim Wolf Snake, Sikkim False Wolf Snake

Sikkim-Wolfzahnnatter

Sikkim-ulvesnog

1878 Ophites gammiei Blanford
Lycodon gammiei Boulenger 1890
Dinodon gammiei Wall 1923

Remarks: Previously confused with fasciatus (e.g., Wall 1911). Validity as a separate species confirmed by Mistry, Vogel & Tillack (2007), although recognized also by other, previous works. Based on confusion regarding the type locality ("Cinchona Plantations, British Sikkim", later corrected to "Darjeeling"), gammiei has frequently been listed for Sikkim (e.g., Smith 1943; Murthy 1985; Sanyal, Sur & Gayen 2006), although the type locality is in Darjeeling, West Bengal. Mistry, Vogel & Tillack (2007) pointed out that there was no documented record from Sikkim, but later, Chettri & Bhupathy (2009) provided the first vouchered record from the state.
Distribution: Bhutan, China (Tibet), India (Arunachal Pradesh, Sikkim, West Bengal).


Lycodon gibsonae

Khorat Wolf Snake

Khorat-Wolfzahnnatter

Khorat-ulvesnog

2019 Lycodon gibsonae Vogel & David

Distribution: Thailand.





Lycodon gongshan

Gaoligong Mountains Wolf Snake

Gaoligong-Wolfzahnnatter

Gaoligong-ulvesnog

2011 Lycodon gongshan Vogel & Luo

Distribution: China (Yunnan).


Lycodon gracilis

Slender Bridle Snake, Scarce Bridal Snake, (Günther's Bridal Snake)

Grazile Zügelnatter

Slank Tøjlesnog

1864 Odontomus gracilis Günther
Hydrophobus gracilis Boulenger 1890
Dryocalamus gracilis Boulenger 1893
Lycodon gracilis Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016
1888 Odontomus fergusonii Haly (Smith 1943)

Remarks: Not mentioned for Kerala by Palot (2015).
Distribution: India (Kerala, Odisha, Tamil Nadu), Myanmar, Sri Lanka.


Lycodon hypsirhinoides

Andaman Wolf Snake

Andamanische Wolfzahnnatter

Andamansk Ulvesnog

1868 Tytleria hypsirhinoides Theobald
Lycodon hypsirhinoides Vogel & Harikrishnan 2013

Remarks: Previously regarded as a synonym of capucinus (e.g., Iskandar & Colijn 2001). Revalidated as a separate species by Vogel & Harikrishnan (2013).
Distribution: India (Andaman Islands).


Lycodon jara

Twin-spotted Wolf Snake, Yellow-striped Wolf Snake, Yellow-speckled Wolf Snake

Doppelpunkt-Wolfzahnnatter

Gulspættet Ulvesnog

1802 Coluber jara Shaw
Leptorhytaon jara Günther 1864
Lycodon jara Stoliczka 1871
Ophites jara Wall 1923
1839 Coluber bipunctatus Cantor (Smith 1943)
2014 Lycodon odishii Mallik, Parida, Mohanty, Mallik, Purohit, Mohanty, Nanda, Sindura, Purohit, Mishra & Sahoo (Chaudhuri, Sharma & Purkayastha 2015)

Distribution: Bangladesh, Bhutan, India (Arunachal Pradesh, Assam, Haryana, Jharkhand, Meghalaya, Mizoram, Nagaland, Odisha, Sikkim, Tripura, Uttar Pradesh, Uttarakhand, West Bengal), Myanmar, Nepal.


Lycodon kundui

Burmese Wolf Snake, (Kundu's Wolf Snake)

Burmesische Wolfzahnnatter

Burmesisk Ulvesnog

1943 Lycodon kundui Smith

Distribution: Myanmar.


Lycodon laoensis

Laotian Wolf Snake, Indochinese Wolf Snake

Laotische Wolfzahnnatter

Laotisk Ulvesnog

1864 Lycodon laoensis Günther
Ophites laoensis Zhao & Adler 1993

Distribution: Cambodia, China (Guangxi, Yunnan), India (Arunachal Pradesh), Laos, Malaysia (West Malaysia), Thailand, Vietnam.


Lycodon liuchengchaoi

Tangjiahe Wolf Snake

Tangjiahe-Wolfzahnnatter

Tangjiahe-ulvesnog

2011 Lycodon liuchengchaoi Zhang, Jiang, Vogel & Rao

Distribution: China (Anhui, Sichuan).


Lycodon mackinnoni

Musoorie Wolf Snake, (Mackinnon's Wolf Snake)

Musoorie-Wolfzahnnatter

Musoorie-ulvesnog

1906 Lycodon mackinnoni Wall
Ophites mackinnoni Wall 1923

Distribution: India (Jammu & Kashmir, Uttarakhand).


Lycodon meridionalis

Vietnamese Big-toothed Snake, Southern Big-toothed Snake

Vietnamesische Wolfzahnnatter

Vietnamesisk Ulvesnog

1935 Dinodon rufozonatum meridionale Bourret
Dinodon meridionale Orlov & Ryabov 2004
Lycodon meridionale Guo, Zhang, Liu, Li, Pyron Jiang & Burbrink 2013

Remarks: Previously regarded as a synonym of flavozonatus (e.g., Smith 1943). Revalidated as a separate species by Orlov & Ryabov (2004).
Distribution: China (Guangxi, Yunnan), Laos, Vietnam (incl. Cat Ba Island).


Lycodon muelleri

Luzon Wolf Snake, (Müller's Wolf Snake)

Luzon-Wolfzahnnatter

Luzon-ulvesnog

1854 Lycodon mülleri Dumeril, Bibron & Dumeril
Odontomus mülleri Boettger 1886
1898 Stegonotus dumerili Boettger 1898 [not Stegonotus dumerilii Boulenger 1893]
1910 Haplonodon philippinensis Griffin 1910 (Leviton 1965)

Distribution: Philippines (Batan Islands [Batan], Catanduanes, Luzon, Mindoro, Polillo).


Lycodon multifasciatus

Ryukyu Mountain Wolf Snake, Sakishima Plum Blossom Snake

Vielbändige Wolfzahnnatter

Mangebåndet Ulvesnog

1931 Dinodon septentrionale multifasciatum Maki
Lycodon ruhstrati multifasciatus Mori 1984
Lycodon multifasciatus Vogel & Brachtel 2008

Remarks: Previously regarded as a subspecies of ruhstrahti (e.g., Goris & Maeda 2004; Nguyen, Ho & Nguyen 2009). Raised to species status by Vogel & Brachtel (2008). Validity confirmed by Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler (2009).
Distribution: Japan (Ryukyu Islands: Sakishima Islands [Miyako Islands, Yaeyama Islands]).


Lycodon multizonatus

Luding Wolf Snake, (Luding Kukri Snake)

Luding-Wolfzahnnatter

Luding-ulvesnog

1981 Oligodon multizonatum Zhao & Jiang
Lycodon multizonatum Lei, Sun, Jiang, Vogel, Booth & Ding 2014

Remarks: Not mentioned by Zhang, Jang, Li, Hou & Rao (2011).
Distribution: China (Chongqing, Gansu, Sichuan).


Lycodon namdongensis

Thanh Hoa Wolf Snake

Thanh-Hoa-Wolfzahnnatter

Thanh Hoa-ulvesnog

2019 Lycodon namdongensis Luu, Ziegler, Ha, Le & Hoang

Distribution: Vietnam.





Lycodon nympha

Vellore Bridle Snake, (Common Bridal Snake)

Nymphen-Zügelnatter

Nymfetøjlesnog

1803 Coluber nympha Daudin
Odontomus nympha Günther 1964
Hydrophobus nympha Boulenger 1890
Dryocalamus nympha Boulenger 1893
Lycodon nympha Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016
1862 Hydrophobus semifasciatus Günther (Smith 1943)
Odontomus semifasciatus Günther 1964
1887 Dryocalamus nympha ceylonensis Müller (Smith 1943)
1924 Cochliophagus isolepis Müller (Kucharzewski & Tillack 2008)

Distribution: India (Andhra Pradesh, Karnataka, Kerala, Odisha, Tamil Nadu), Sri Lanka.


Lycodon ophiophagus

Snake-eater Wolf Snake

Schlangenfressende Wolfzahnnatter

Slangeædende Ulvesnog

2009 Lycodon ophiophagus Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler

Remarks: This species was previously regarded as a population of ruhstrati.
Distribution: Thailand.


Lycodon orientalis

East Asian Wolf Snake, Eastern False Wolf Snake, White-banded Snake, Oriental Big-toothed Snake, Shiromadara

Ostasiatische Wolfzahnnatter

Østasiatisk Ulvesnog

1880 Ophites orientalis Hilgendorf
Dinodon orientale Stejneger 1907
Lycodon orientalis Guo, Zhang, Liu, Li, Pyron Jiang & Burbrink 2013
1880 Ophites japonicus Günther (Stejneger 1907)
Dinodon japonicus Boulenger 1893

Distribution: Japan (Hokkaido, Honshu, Kyushu, Shikoku, Izu Islands, Ryukyu Islands [Satsunan Islands], Volcano Islands), Russia (Kuril Islands).


Lycodon paucifasciatus

Annam Wolf Snake, (Rendahl's Wolf Snake)

Annam-Wolfzahnnatter

Annam-ulvesnog

1943 Lycodon paucifasciatus Rendahl in Smith

Distribution: Vietnam.


Lycodon philippinus

Philippine Bridle Snake

Philippinische Zügelnatter

Filippinsk Tøjlesnog

1909 Dryocalamus philippinus Griffin
Lycodon philippinus Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016

Remarks: Not mentioned by Brown & Alcala (1970) or Alcala (1986), who only mentioned tristrigatus as part of the Philippine snake fauna, presumably considering philippinus a synonym of tristrigatus. Listed (with a question mark) as a synonym of tristrigatus by Iskandar & Colijn (2001). Listed as a valid species by the Reptile Database (viewed 3 March 2015).
Distribution: Philippines (Balabac, Palawan).


Lycodon pictus

Painted Wolf Snake

Gemalte Wolfzahnnatter

Malet Ulvesnog

2019 Lycodon pictus Janssen, Pham, Ngo, Le, Nguyen & Ziegler

Distribution: Vietnam.





Lycodon rosozonatus

Rosy-banded Big-toothed Snake, Rose Big-toothed Snake, Pink Big-toothed Snake

Rosenbändige Wolfzahnnatter

Rosenbåndet Ulvesnog

1972 Dinodon rosozonatum Hu & Zhao
Lycodon rosozonatus Guo, Zhang, Liu, Li, Pyron Jiang & Burbrink 2013

Distribution: China (Hainan), Vietnam.


Lycodon rufozonatus

Red-banded Big-toothed Snake, Red Big-toothed Snake, Red-banded Snake, Red-banded Odd-tooth Snake, Chinese Red-banded Snake, Banded Red Snake, Sakishima King Snake, Asian King Snake, Akamadara

Rot-Schwarze Wolfzahnnatter

Rødbåndet Ulvesnog

1842 Lycodon rufo-zonatus Cantor
Dinodon rufozonatus Peters 1881
1853 Dinodon cancellatum Duméril (Zhao & Adler 1993)
1856 Coronella striata Hallowell (Zhao & Adler 1993)
1885 Dinodon rufozonatus formosana Boettger (Zhao & Adler 1993)
1907 Dinodon rufozonatum walli Stejneger
1925 Adiastema cervinum Werner (Iskandar & Colijn 2001)
1925 Dinodon rufozonatum williamsi Schmidt (Zhao & Adler 1993)
1931 Dinodon rufozonatum yunnanense Mell (Zhao & Adler 1993)

Other common names:
walli: Sakishima Banded Snake
Remarks: Records from Laos (e.g., Deuve 1970) refer to meridionalis (Orlov & Ryabov 2004). Included for Russia based on a number of sources (e.g., Kudrjavtsev & Mamet 1989; Maslov & Kotlobay 1998; Szczerbak 2003; Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006), although Zhao & Adler (1993) claim that presence in Russia is unconfirmed.
Distribution: China (Anhui, Beijing, Chongqing, Fujian, Gansu, Guangdong [incl. Nan Ao Island], Guangxi, Guizhou, Hainan, Hebei, Heilongjiang, Henan, Hubei, Hunan, Jiangsu, Jiangxi, Jilin, Liaoning, Shaanxi, Shandong, Shanxi, Sichuan, Yunnan, Zhejiang), Japan (Tsushima Islands, Ryukyu Islands [Sakishima Islands (Miyako Islands, Yaeyama Islands)]), North Korea, Russia, South Korea, Taiwan, Vietnam.


Lycodon ruhstrati

Taiwanese Wolf Snake, Mountain Wolf Snake, Formosa Wolf Snake, (Ruhstrat's Wolf Snake)

Taiwanische Wolfzahnnatter

Taiwansk Ulvesnog

1886 Ophites ruhstrati Fischer
Dinodon septentrionale ruhstrati Boulenger 1899
Dinodon ruhstrati Pope 1929
Lycodon ruhstrati Pope 1935
2009 Lycodon ruhstrati abditus Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler

Remarks: Previously included futsingensis and multifasciatus, and populations now assigned to ophiophagus. Exact distribution remains unclear, as many literature records cannot with certainty be assigned to futsingensis or ruhstrati abditus (Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler 2009). Records from Laos (e.g., Nguyen, Ho & Nguyen 2009) refer to fasciatus (Teynié & David 2010).
Distribution: China (Anhui, Beijing, Chongqing, Fujian, Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangsu, Jiangxi, Shaanxi, Sichuan, Yunnan, Zhejiang), Laos, Taiwan, Vietnam.


Lycodon sealei

Palawan Wolf Snake, (Seale's Banded Asian Wolf Snake)

Palawan-Wolfzahnnatter

Palawan-ulvesnog

1955 Lycodon subcinctus sealei Leviton
Lycodon sealei Leviton, Siler, Weinell & Brown 2018

Remarks: Previously regarded as a subspecies of subcinctus. Regarded herein as a separate species, following Leviton, Siler, Weinell & Brown (2018).
Distribution: Brunei, Indonesia (Kalimantan), Malaysia (Sabah, Sarawak), Philippines (Palawan).


Lycodon semicarinatus

Tricolored Banded Snake

Dreifarbige Wolfzahnnatter

Trefarvet Ulvesnog

1860 Eumesodon semicarinatus Cope
Dinodon semicarinatum Boulenger 1892
Lycodon semicarinatus Guo, Zhang, Liu, Li, Pyron Jiang & Burbrink 2013
1860 Lepidocephalus fasciatus Hallowell (Stejneger 1907)

Distribution: Japan (Ryukyu Islands [Okinawa Islands, Satsunan Islands]).


Lycodon septentrionalis

Northern Big-toothed Snake, White-banded Wolf Snake, Northern Large-toothed Snake, Hill Wolf Snake, (Günther's False Wolf Snake)

Nördliche Wolfzahnnatter

Nordlig Ulvesnog

1875 Ophites septentrionalis Günther
Lycodon septentrionalis Boulenger 1890
Dinodon septentrionalis Boulenger 1893
1933 Dinodon septentrionale chapaense Angel & Bourret (Smith 1943)

Remarks: Records from Japan (Riu Kiu Islands) and Taiwan are based on misidentified ruhstrahti (Ota 1988; Zhao & Adler 1993; Nguyen, Ho & Nguyen 2009). Although this species was reported from Jammu & Kashmir by Sahi & Duda (1985), it was not listed in a subsequent work by the same authors (Sahi & Duda 1996). It is unclear whether the authors intentionally removed the species from the state list in their latter work, but since the closest records are in NE. India, its presence in the state is here regarded as doubtful.
Distribution: Bhutan, Cambodia, China (Yunnan), India (Arunachal Pradesh, Assam, Mizoram, Nagaland, West Bengal), Laos, Myanmar, Thailand, Vietnam.


Lycodon sidiki

Aceh Wolf Snake

Aceh-Wolfzahnnatter

Aceh-ulvesnog

2017 Lycodon sidiki Wostl, Hamidy, Kurniawan & Smith

Distribution: Indonesia (Sumatra).


Lycodon solivagus

Pale-banded Wolf Snake, Northern Luzon Wolf Snake

Hellbändige Wolfzahnnatter

Lysbåndet Ulvesnog

1994 Lycodon solivagus Ota & Ross

Distribution: Philippines (Luzon).


Lycodon stormi

Sulawesi Wolf Snake

Sulawesi-Wolfzahnnatter

Sulawesi-ulvesnog

1892 Lycodon stormi Boettger

Distribution: Indonesia (Sulawesi).


Lycodon striatus

Striped Wolf Snake, Barred Wolf Snake, (Shaw's Wolf Snake, White-spotted Wolf Snake)

Gestreifte Wolfzahnnatter

Stribet Ulvesnog

1802 Coluber striatus Shaw
Lycodon striatus Stoliczka 1870
Ophites striatus Wall 1921
1803 Coluber galathea Daudin (Smith 1943)
1803 Coluber malignus Daudin (Smith 1943)
1854 Lycodon napei Duméril & Bibron (status uncertain; Smith 1943)
1903 Contia bicolor Nikolsky
Lycodon striatus bicolor Terentjev & Chernov 1949
1955 Lycodon striatus nordicus Deraniyagala (Lanza 1999)
1955 Lycodon striatus sinhaleyus Deraniyagala

Other common names:
bicolor: Golden-spotted Wolf Snake, Golden Wolf Snake
striatus: Northern Barred Wolf Snake, Northern Wolf Snake, Indus Valley Wolf Snake
Distribution: India (Andhra Pradesh, Bihar, Chhattisgarh, Goa, Gujarat, Haryana, Himachal Pradesh, Jammu & Kashmir, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Odisha, Punjab, Rajasthan, Tamil Nadu, Uttar Pradesh, Uttarakhand, West Bengal), Iran, Nepal, Pakistan, Sri Lanka, Tajikistan, Turkmenistan, Uzbekistan.


Lycodon subannulatus

Malayan Bridle Snake, Half-banded Bridled Snake, Saddle-backed Dwarf Tree Snake

Gebänderte Zügelnatter

Malayisk Tøjlesnog

1854 Odontomus sub-annulatus Dumeril, Bibron & Dumeril
Nymphophidium subannulatum Blanford 1881
Hydrophobus subannulatus Boulenger 1890
Dryocalamus subannulatus Boulenger 1893
Lycodon subannulatus Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016
1864 Nymphodidium maculatum Günther (Iskandar & Colijn 2001)
1986 Dryocalamus davisonii tungsongensis Nutphand (Pauwels, David & Chan-ard 2006)

Remarks: Reported from Siberut by Haas (1950), but Dring, McCarthy & Whitten (1990) were unable to find any sources supporting this record. Leviton, Siler, Weinell & Brown (2018) did not include Palawan in the distribution of the species.
Distribution: Brunei, Indonesia (Banyak Islands, Java, Riau Islands, Siberut, Sumatra), Malaysia (Sabah, Sarawak, West Malaysia [incl. Langkawi Archipelago, Seribuat Archipelago (incl. Tioman)]), Myanmar, Philippines (Palawan), Singapore, Thailand.


Lycodon subcinctus

Malayan Banded Wolf Snake, White-banded Wolf Snake

Gebänderte Wolfzahnnatter

Malaysisk Båndet Ulvesnog

1827 Lycodon subcinctus Boie
Ophites subcinctus Wagler 1830
1884 Elapoidis annulatus Sauvage (Smith 1943)
1895 Anoplophallus maculatus Cope
1986 Lycodon suratensis Nutphand (Pauwels, David & Chan-ard 2006)

Remarks: Previously included sealei.
Distribution: Brunei, Cambodia, China (Fujian, Guangdong, Guangxi, Hainan, Hong Kong, Yunnan), East Timor, India (Nicobar Islands), Indonesia (Alor, Bali, Flores, Java, Kalimantan, Lombok, Nias, Penida, Siberut, Simeulue, Sumatra, Sumba, Sumbawa), Laos, Malaysia (Sabah, Sarawak, West Malaysia), Myanmar, Singapore, Thailand, Vietnam.


Lycodon synaptor

Dongchuan Wolf Snake, (Böhme's Wolf Snake)

Dongchuan-Wolfzahnnatter

Dongchuan-ulvesnog

2010 Lycodon synaptor Vogel & David

Distribution: China (Yunnan).


Lycodon tessellatus

Manila Wolf Snake

Manila-Wolfzahnnatter

Manila-ulvesnog

1863 Lycodon tessellatus Jan
Ophites tessellatus Griffin 1911

Remarks: Known from only two specimens said to have come from Luzon, Philippines.
Distribution: Philippines (Luzon).


Lycodon tiwarii

Nicobar Wolf Snake, (Tiwari's Wolf Snake)

Nicobarische Wolfzahnnatter

Nicobarisk Ulvesnog

1965 Lycodon tiwarii Biswas & Sanyal

Remarks: Presence in the Andaman Islands needs confirmation. Although the type locality is in the Andamans, there has been no records of this species from these islands since its original description (Harikrishnan, Vasudevan & Choudhury 2010).
Distribution: India (Andaman & Nicobar Islands [Andaman Islands, Nicobar Islands]).


Lycodon travancoricus

Travancore Wolf Snake, South Indian Wolf Snake, (Beddome's Wolf Snake)

Travancore-Wolfzahnnatter

Travancore-ulvesnog

1870 Cercaspis travancoricus Beddome
Lycodon travancoricus Boulenger 1890
Lycodon aulicus travancoricus Constable 1940

Remarks: Presence in Jammu & Kashmir needs confirmation (Whitaker & Captain 2004), although reported by several authors (e.g., Murthy & Sharma 1976; Sahi & Duda 1985; Sharma & Sharma 1975).
Distribution: India (Andhra Pradesh, Goa, Gujarat, Karnataka, Kerala, Lakshadweep, Madhya Pradesh, Maharashtra, Odisha, Tamil Nadu), Pakistan.


Lycodon tristrigatus

Three-banded Bridle Snake, Striped Dwarf Tree Snake

Dreistreifen-Zügelnatter

Trestribet Tøjlesnog

1858 Dryocalamus tristrigatus Günther
Lycodon tristrigatus Figueroa, KcKelvy, Grismer, Bell & Lailvaux 2016

Remarks: See under philippinus. Records from the Philippines are herein assigned to philippinus.
Distribution: Brunei, Indonesia (Kalimantan, Natuna Islands), Malaysia (Sabah, Sarawak).


Lycodon zawi

Sagaing Wolf Snake, (Zaw's Wolf Snake)

Sagaing-Wolfzahnnatter

Sagaing-ulvesnog

2001 Lycodon zawi Slowinski, Pawar, Win, Thin, Tun, Gyi, Oo & Tun

Distribution: Bangladesh, India (Assam, Meghalaya, Mizoram, West Bengal), Myanmar.


Lycodon zoosvictoriae

Phnom Samkos Wolf Snake

Phnom-Samkos-Wolfzahnnatter

Phnom Samkos-ulvesnog

2014 Lycodon zoosvictoriae Neang, Hartmann, Hun, Souter & Furey

Distribution: Cambodia.