Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk).
Latest update: May 12th, 2024.
 
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Genus
Pseudocalotes
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Long-headed Agamas, Long-headed Lizards, False Garden Lizards, False Blood-suckers, Forest Agamids
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Spitzkopfagamen
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Spidshovedagamer
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1843 |  |
Pseudocalotes Fitzinger (type species: Calotes tympanistra [sic] Gray 1831) |
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1935 | |
Mictopholis Smith (type species: Salea austeniana Annandale 1908; syn. Mahony 2010) |
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1939 | |
Paracalotes Bourret (type species: Paracalotes poilani Bourret 1939; syn. Hallermann & Böhme 2000) |
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1997 | |
Pseudocophotis Manthey & Grossmann (type species: Cophotis sumatrana Hubrecht 1879; syn. Hallermann & Böhme 2000) |
Contents:
24 species, of which 18 (75.0%) are endemic.
Endemism: 0%  100%
Other common names:
Pseudocophotis: Prehensile-tailed Agamas
Remarks:
Previously included in the genus
Calotes
(e.g., Wermuth 1967).
Formerly included
Japalura austeniana.
Includes Paracalotes, regarded as a synonym of Pseudocalotes already by Welch, Cooke & Wright (1990), but with no discussion or references.
Synonymy confirmed by (Hallermann & Böhme 2000).
Hypsicalotes kinabaluensis was previously included in Calotes (e.g., Wermuth 1967), or more recently, in Pseudocalotes (e.g., Inger & Stuebing 1994).
Status of Pseudocophotis is uncertain.
It was tentatively recognized by Ananjeva, Orlov, Nguyen & Nazarov (2007), who described a new species of the nominal genus (kontumensis).
Also Das (2010) and Manthey (2010) treated Pseudocophotis as a valid genus.
Until more detailed studies have been published, the review of Hallermann & Böhme (2000) is followed here, treating Pseudocophotis as a synonym of Pseudocalotes.
Distribution:
SE. Asia, Indonesia.
Reported from:
Bhutan,
Cambodia,
China
(Guangdong,
Guangxi,
Guizhou,
Hainan,
Tibet,
Yunnan),
India
(Andaman & Nicobar Islands
[Andaman Islands,
Nicobar Islands],
Arunachal Pradesh),
Indonesia
(Java,
Sumatra),
Laos,
Malaysia
(Sarawak,
West Malaysia),
Myanmar,
Thailand,
Vietnam
(incl.
Cat Ba Island).
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Pseudocalotes andamanensis
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Andaman Long-headed Agama, Green Crestless Forest Agama, (Andaman Green Calotes)
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Andamanische Spitzkopfagame
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Andamansk Spidshovedagam
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1891 | |
Calotes andamanensis Boulenger |
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Pseudocalotes andamanensis Harikrishnan & Vasudevan 2013 |
Remarks:
Ishwar & Das (1998) suggested that the type locality (Andaman Islands) was in error and restricted it to the Western Ghats of Tamil Nadu, India
(repeated by Aengals (2009) and Daniels (2001a)).
Subsequently, Krishnan (2008) showed that the Western Ghats population and andamanensis (at that time still included in
Calotes)
are not the same species, and he described the Western Ghats population as a new species,
Calotes aurantolabium,
and restricted the type locality of andamanensis to the Nicobar Islands, erroneously assuming that the collector of the type specimen, F.A. de Roepstorff,
never visited the Andaman Islands.
This was proved wrong by Harikrishnan & Vasudevan (2013), who concluded that the Andaman Islands is indeed the correct type locality, as originally stated.
Distribution:
India
(Andaman & Nicobar Islands
[Andaman Islands,
Nicobar Islands]),
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Pseudocalotes baliomus
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Barisan Range Long-headed Agama, Spot-shouldered False Garden Lizard
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Barisan-Spitzkopfagame
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Barisan-spidshovedagam
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2017 | |
Pseudocalotes baliomus Harvey, Shaney, Hamidy, Kurniawan & Smith |
Distribution:
Indonesia
(Sumatra).
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Pseudocalotes brevipes
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Short-footed Long-headed Agama, Vietnamese Tree Lizard, Vietnamese False Bloodsucker
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Kurzbeinige Spitzkopfagame
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Kortfodet Spidshovedagam
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1904 | |
Calotes brevipes Werner |
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Pseudocalotes brevipes Welch, Cooke & Wright 1990 |
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1931 | |
Acanthosaura quinquecarinata Fan (syn. Wermuth 1967) |
Distribution:
China
(Guangxi,
Yunnan),
Vietnam
(incl.
Cat Ba Island).
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Pseudocalotes cybelidermus
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Purple-throated Long-headed Agama, Purple-throated False Garden Lizard
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Purpur-Spitzkopfagame
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Purpurstrubet Spidshovedagam
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2014 | |
Pseudocalotes cybelidermus Harvey, Hamidy, Wan, Shaney & Smith |
Distribution:
Indonesia
(Sumatra).
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Pseudocalotes dringi
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Mount Tahan Long-headed Agama, (Dring's Long-headed Agama)
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Drings Spitzkopfagame
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Tahan-spidshovedagam
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2000 | |
Pseudocalotes dringi Hallermann & Böhme |
Distribution:
Malaysia
(West Malaysia).
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Pseudocalotes drogon
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Fraser's Hill Long-headed Agama, (Drogon's False Garden Lizard)
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Fraser's-Hill-Spitzkopfagame
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Fraser's Hill-spidshovedagam
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2016 | |
Pseudocalotes drogon Grismer, Quah, Wood, Anuar, Muin, Davis, Murdoch, Grismer, Cota & Cobos |
Distribution:
Malaysia
(West Malaysia).
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Pseudocalotes flavigula
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Yellow-throated Long-headed Agama, Yellow-throated Forest Agamid
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Gelbkehl-Spitzkopfagame
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Gulstrubet Spidshovedagam
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1924 | |
Calotes flavigula Smith |
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Pseudocalotes flavigula Welch, Cooke & Wright 1990 |
Distribution:
Malaysia
(West Malaysia).
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Pseudocalotes floweri
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Chanthaburi Long-headed Agama, (Flower’s Long-headed Agama, Flower's Forest Agamid, Thai False Bloodsucker)
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Flowers Spitzkopfagame
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Chanthaburi-spidshovedagam
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1912 | |
Calotes floweri Boulenger |
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Pseudocalotes floweri Welch, Cooke & Wright 1990 |
Remarks:
This species is not known from West Malaysia (Hallermann & Böhme 2000). Boulenger's (1912) syntypes of floweri from West Malaysia refer to dringi (a record repeated by Smith (1935), Wermuth (1967), Denzer & Manthey (1991, as "P. cf. floweri"), and probably others as well). A population in Vietnam (e.g., Nguyen, Ho & Nguyen 2009) has now been described as a separate species, ziegleri (Hallermann, Nguyen, Orlov & Ananjeva 2010).
Distribution:
Cambodia,
Thailand.
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Pseudocalotes guttalineatus
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Dash-lined Long-headed Agama, Dash-lined False Garden Lizard
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Strichlierter Spitzkopfagame
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Stiplet Spidshovedagam
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2014 | |
Pseudocalotes guttalineatus Harvey, Hamidy, Wan, Shaney & Smith |
Distribution:
Indonesia
(Sumatra).
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Pseudocalotes jingpo
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Jingpo Long-headed Agama
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Jingpo-Spitzkopfagame
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Jingpo-spidshovedagam
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2024 | |
Pseudocalotes jingpo Xu, Gong, Hou, Weng, Liu, Deng, Hu & Peng |
Distribution:
China
(Yunnan).
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Pseudocalotes kakhienensis
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Kakhien Hills Long-headed Agama, (Burmese Mountain Agamid)
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Kakhyen-Spitzkopfagame
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Kakhien-spidshovedagam
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1870 | |
Oriocalotes discolor Anderson (primary objective synonym) (syn. Mahony 2010) |
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1878 | |
Oriocalotes kakhienensis Anderson |
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Acanthosaura kakhienensis Boulenger 1885 |
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Calotes kakhienensis Pope 1935 |
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Salea kakhiensis [lapsus] Moody 1981 |
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Pseudocalotes kakhienensis Mahony 2010 |
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1887 | |
Calotes feae Boulenger (syn. Mahony 2010) |
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1937 | |
Salea brachydactyla Rendahl (syn. Manthey & Denzer 2000; Mahony 2010) |
Remarks:
Placed in the genus Salea (and misspelled kakhiensis!) by Moody (1981), Welch, Cooke & Wright (1990), and Hallermann, Ananjeva, Orlov & Tillack (2002), but with no discussion or references. Although reported from India by Das (1996), there are no verified records from the country (Venugopal 2010). Although the name discolor has priority over kakhienensis, Mahony (2010) argued that the latter name should be retained for nomenclatural stability.
Distribution:
China
(Yunnan),
Myanmar.
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Pseudocalotes khaonanensis
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Khao Nan Long-headed Agama
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Khaonan-Spitzkopfagame
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Khao Nan-spidshovedagam
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2008 | |
Pseudocalotes khaonanensis Chan-Ard, Cota, Makchai & Laoteow |
Distribution:
Thailand.
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Pseudocalotes kingdonwardi
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Adung Valley Long-headed Agama, (Kingdon-Ward's Forest Lizard)
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Kingdonwards Spitzkopfagame, Kingdonwards Schönechse
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Adung-spidshovedagam
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1935 | |
Calotes kingdon-wardi Smith |
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Pseudocalotes kingdonwardi Mahony 2010 |
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1937 | |
Japalura kaulbacki Smith (syn. Mahony 2010) |
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Pseudocalotes kaulbacki Mahony 2010 (syn. see remarks) |
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1979 | |
Calotes kingdonwardi bapoensis Yang & Su |
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Japalura bapoensis Yang & Rao 2008 |
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Pseudocalotes kingdonwardi bapoensis Mahony 2010 |
Other common names:
bapoensis: Bapo Long-headed Agama
kaulbacki: Kaulback's Mountain Lizard
Remarks:
Although Mahony (2010) synonymized kaulbacki with kingdonwardi, he also (for reasons unknown) used the new combination Pseudocalotes kaulbacki in the same paper.
The subspecies bapoensis may represent a separate species, as suggested by Yang & Rao (2008).
Reports from Tibet are erroneous (Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021).
Distribution:
China
(Yunnan),
India
(Arunachal Pradesh),
Myanmar.
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Pseudocalotes kontumensis
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Kon Tum Long-headed Agama, Kon Tum Prehensile-tailed Agama, Kon Tum Tree Lizard
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Kontum-Greifschwanzagame
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Kon Tum-spidshovedagam
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2007 | |
Pseudocophotis kontumensis Ananjeva, Orlov, Nguyen & Nazarov |
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Pseudocalotes kontumensis (new combination) |
Distribution:
Vietnam.
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Pseudocalotes larutensis
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Larut Long-headed Agama
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Larut-Spitzkopfagame
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Larut-spidshovedagam
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2001 | |
Pseudocalotes larutensis Hallermann & McGuire |
Distribution:
Malaysia
(West Malaysia).
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Pseudocalotes microlepis
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Small-scaled Long-headed Agama, Burmese Long-headed Agama, Small-scaled Forest Lizard, Small-scaled Montane Forest Lizard, Boulenger’s Long-headed Agama, Burmese False Bloodsucker
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Burmesische Spitzkopfagame
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Finskællet Spidshovedagam
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1888 | |
Calotes microlepis Boulenger |
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Pseudocalotes microlepis Welch, Cooke & Wright 1990 |
Remarks:
A record from India (Manipur or "Assam") (e.g., Smith 1935; Das 2003; Ahmed, Das & Dutta 2009) represents an undescribed species, presumably of the
genus Calotes (Raju, Das, Bag & Raha 2020).
Distribution:
China
(Guangdong,
Guangxi,
Guizhou,
Hainan,
Yunnan),
Laos,
Myanmar,
Thailand,
Vietnam.
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Pseudocalotes poilani
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Laotian Long-headed Agama, (Poilane's Long-headed Agama)
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Laotische Spitzkopfagame, (Poilanes Spitzkopfagame)
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Laotisk Spidshovedagam
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1939 | |
Paracalotes poilani Bourret |
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Pseudocalotes poilani Welch, Cooke & Wright 1990 |
Distribution:
Laos.
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Pseudocalotes rhaegal
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Cameron Highlands Long-headed Agama, (Rhaegal's False Garden Lizard)
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Cameron-Highlands-Spitzkopfagame
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Cameron Highlands-spidshovedagam
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2016 | |
Pseudocalotes rhaegal Grismer, Quah, Wood, Anuar, Muin, Davis, Murdoch, Grismer, Cota & Cobos |
Distribution:
Malaysia
(West Malaysia).
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Pseudocalotes rhammanotus
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Stitch-backed Long-headed Agama, Stitched-back False Garden Lizard
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Ranau-Spitzkopfagame
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Ranau-spidshovedagam
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2014 | |
Pseudocalotes rhammanotus Harvey, Hamidy, Wan, Shaney & Smith |
Distribution:
Indonesia
(Sumatra).
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Pseudocalotes saravacensis
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Sarawak Long-headed Agama, Bornean Long-headed Agama
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Sarawak-Spitzkopfagame
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Sarawak-spidshovedagam
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1994 | |
Pseudocalotes saravacensis Inger & Stuebing |
Distribution:
Malaysia
(Sarawak).
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Pseudocalotes sumatrana
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Sumatran Long-headed Agama, Prehensile-tailed Agama
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Sumatra-Greifschwanzagame
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Sumatra-spidshovedagam
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1879 | |
Cophotis sumatrana Hubrecht [nomen protectum; Denzer, Dondorp, Koppetsch, Lahaise, Manthey, Mostadius & Böhme 2021] |
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Pseudocophotis sumatrana Manthey & Grossmann 1997 |
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Pseudocalotes sumatrana Hallerman & Böhme 2000 |
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1855 | |
Calotes nasicornis Hoeven [nomen oblitum] (syn. Denzer, Dondorp, Koppetsch, Lahaise, Manthey, Mostadius & Böhme 2021) |
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1905 | |
Calotes aberrans Rosén (syn. Denzer, Dondorp, Koppetsch, Lahaise, Manthey, Mostadius & Böhme 2021) |
Remarks:
Status uncertain.
Some authors have placed it in a separate, monotypic genus, Pseudocophotis (e.g., Manthey & Grossmann 1997; Manthey & Denzer 2000; Ananjeva, Orlov, Nguyen & Nazarov 2007).
Now considered endemic to Java.
Although the type locality is Sumatra, presence in this island is unconfirmed (Denzer, Dondorp, Koppetsch, Lahaise, Manthey, Mostadius & Böhme 2021).
Distribution:
Indonesia
(Java).
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Pseudocalotes tympanistriga
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Javanese Long-headed Agama, (Kuhl's Long-headed Agama)
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Kuhls Spitzkopfagame
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Javansk Spidshovedagam
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1831 | |
Calotes tympanistra [error typographicus] Gray |
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Bronchocela tympanistriga [emendation] Duméril & Bibron 1837 |
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Calotes tympanistriga Boulenger 1885 |
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Pseudocalotes tympanistriga Welch, Cooke & Wright 1990 |
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1889 | |
Salea rosaceum Thominot (syn. Harvey, Shaney, Sidik, Kurniawan & Smith 2017) |
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1893 | |
Dendragama fruhstorferi Boettger (syn. Wermuth 1967) |
Remarks:
Salea rosaceum was previously regarded as a synonym of Dendragama boulengeri (e.g., Wermuth 1967).
Distribution:
Indonesia
(Java,
Sumatra).
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Pseudocalotes viserion
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Pahang Long-headed Agama, (Viserion’s False Garden Lizard)
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Pahang-Spitzkopfagame
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Pahang-spidshovedagam
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2016 | |
Pseudocalotes viserion Grismer, Quah, Wood, Anuar, Muin, Davis, Murdoch, Grismer, Cota & Cobos |
Distribution:
Malaysia
(West Malaysia).
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Pseudocalotes ziegleri
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Vietnamese Long-headed Agama, (Ziegler's Tree Lizard)
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Vietnamesische Spitzkopfagame
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Vietnamesisk Spidshovedagam
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2010 | |
Pseudocalotes ziegleri Hallermann, Nguyen, Orlov & Ananjeva |
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Pseudocophotis ziegleri Bain & Hurley 2011 |
Remarks:
Previously confused with floweri.
Distribution:
Laos,
Vietnam.
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