Reptiles of Bhutan    Bibliography of the reptiles of Bhutan Taxonomic history of the reptiles of Bhutan Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus palustris Mugger Whitaker & Andrews 2003   Crocodylus porosus Saltwater Crocodile Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022  Family Gavialidae (Gharials) Genus Gavialis (Indian Gharial)   EX Gavialis gangeticus Indian Gharial Bauer & Gunther 1992 Das 1996 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2011 Wangyal 2019 Whitaker & Andrews 2003

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Calotes (South Asian Forest Agamas)   * Calotes bhutanensis Bhutanese Tree Agama Bauer & Gunther 1992 Biswas 1976 Das 1996 Manthey 2008 Tikader & Sharma 1992 Wangyal 2011 Wangyal 2019   Calotes irawadi Ayeyarwady Forest Agama Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Calotes jerdoni Eastern Green Tree Agama Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019   Calotes maria Assam Forest Agama Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016   Calotes versicolor South Indian Crested Agama Bauer & Gunther 1992 Biswas 1976 Das & Palden 2000 Das 1996 Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019 Genus Cristidorsa (Crested Mountain Agamas)   Cristidorsa otai Mizoram Mountain Agama Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020 Genus Japalura (Western Mountain Agamas)   Japalura andersoniana Dafla Hill Mountain Agama Das 1996   Japalura austeniana Arunachal Mountain Agama Tikader & Sharma 1992 Wangyal 2019 Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Japalura tricarinata Three-keeled Mountain Agama Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Japalura variegata East Himalayan Mountain Agama Bauer & Gunther 1992 Biswas 1976 Das 1996 Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019 Genus Ptyctolaemus (Fold-throated Agamas)   Ptyctolaemus gularis Green Fold-throated Agama Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Wangyal 2019  Family Anguidae (Slowworms, etc.) Genus Dopasia (Oriental Glass Lizards)   Dopasia gracilis Burmese Glass Lizard Wangyal 2013 Wangyal 2019  Family Gekkonidae (Typical Geckos) Genus Cnemaspis (Asian Rock and Forest Geckos)   Cnemaspis assamensis Assam Rock Gecko Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus bhupathyi Bagdogra Bent-toed Gecko Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Cyrtodactylus gubernatoris Darjeeling Rock Gecko Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Cyrtodactylus nagalandensis Nagaland Bent-toed Gecko Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Cyrtodactylus septentrionalis Abhayapuri Lowland Bent-toed Gecko Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020 Genus Gekko (Tokays)   Gekko gecko Tokay Ahmed, Das & Dutta 2009 Wangyal 2012 Wangyal 2019 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus brookii Spotted Half-toed Gecko Bauer & Gunther 1992 Das & Palden 2000 Das 1996 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019   Hemidactylus flaviviridis Yellow-bellied House Gecko Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Hemidactylus frenatus Asian House Gecko Ahmed, Das & Dutta 2009 Bauer & Gunther 1992 Das 1996 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2011 Wangyal 2019   Hemidactylus garnotii Fox Gecko Bauer, Jackman, Greenbaum, Giri & Silva 2010 Das & Palden 2000 Das 1996 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2011 Wangyal 2019   Hemidactylus platyurus Flat-tailed Gecko Ahmed, Das & Dutta 2009 Bauer & Gunther 1992 Das 1996 Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019  Family Lacertidae (Typical Lizards) Genus Takydromus (Oriental Grass Lizards)   Takydromus sikkimensis Sikkim Grass Lizard Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020  Family Scincidae (Skinks) Genus Ablepharus (Eurasian Mountain Skinks)   Ablepharus sikkimensis Sikkim Mountain Skink Bauer & Gunther 1992 Das 1996 Sindaco & Jeremchenko 2008 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019 Genus Eutropis (South Asian Sun Skinks)   Eutropis carinata Brahminy Skink Wangyal 2011 Wangyal 2019   Eutropis macularia Speckled Forest Skink Ahmed, Das & Dutta 2009 Das & Palden 2000 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2011 Wangyal 2019   Eutropis multifasciata Many-lined Sun Skink Ahmed, Das & Dutta 2009   * Eutropis quadratilobus Bhutan Sun Skink Bauer & Gunther 1992 Das 1996 Wangyal 2011 Wangyal 2019   Eutropis trivittata Striped Ground Skink Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 [as dissimilis] Genus Riopa (Asian Gracile Skinks)   Riopa albopunctata White-spotted Gracile Skink Ahmed, Das & Dutta 2009   Riopa punctata Spotted Gracile Skink Wangyal 2011 Wangyal 2019 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus indicus Common Indian Forest Skink Bauer & Gunther 1992 Das & Palden 2000 Das 1996 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019   Sphenomorphus maculatus Spotted Forest Skink Ahmed, Das & Dutta 2009 Bauer & Gunther 1992 Das 1996 Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2012 Wangyal 2019  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus bengalensis Bengal Monitor Ahmed, Das & Dutta 2009 Bauer & Gunther 1992 Das 1996 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2011 Wangyal 2019   Varanus flavescens Yellow Monitor Wangyal 2011 Wangyal 2019   Varanus salvator Asian Water Monitor Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla flavescens Yellow Whip Snake Ahmed, Das & Dutta 2009 [as A. prasina] Das 1996 [as A. p. prasina] Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 [as A. prasina] Mitra 2009 [as A. p. prasina] Tshewang & Letro 2018 [as A. prasina] Wallach, Williams & Boundy 2014 [as A. prasina] Wangyal 2011 [as A. prasina] Wangyal 2019 [as A. prasina]   Ahaetulla nasuta Long-nosed Green Whip Snake Wangyal 2011 Wangyal 2019 Genus Amphiesma (South Asian Keelbacks)   Amphiesma stolatum Buff-striped Keelback Bauer & Gunther 1992 Biswas 1976 Das 1996 Mitra 2009 Wangyal 2011 Wangyal 2019 Genus Blythia (Asian Iridescent Snakes)   Blythia reticulata Assam Iridescent Snake Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020 Genus Boiga (Australasian Cat Snakes)   Boiga cyanea Green Cat Snake Ahmed, Das & Dutta 2009 Orlov & Ryabov 2002 Tillack 2006 Wangyal 2011 Wangyal 2019   Boiga gocool Eastern Gamma Cat Snake Choure, Kashyap, Adhikari & Lalremsanga 2020 Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Mitra 2009 Wangyal 2019   Boiga multifasciata Many-banded Cat Snake Koirala, Gurung, Lhendup & Phuntsho 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Orlov & Ryabov 2002 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019   Boiga ochracea Tawny Cat Snake Bauer & Gunther 1992 Das 1996 Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Orlov & Ryabov 2002 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019   Boiga quincunciata Assam Cat Snake Chaida, Das, Tshering & Wangdi 2020   Boiga siamensis Grey Cat Snake Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea ornata Ornate Flying Tree Snake Ahmed, Das & Dutta 2009 Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2012 Wangyal 2019 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus helena Indian Trinket Snake Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Coelognathus radiatus Copper-headed Rat Snake Ahmed, Das & Dutta 2009 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis cyanochloris Green Bronzeback Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019   Dendrelaphis proarchos Northern Painted Bronzeback Ahmed, Das & Dutta 2009 [as pictus] Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Wangyal 2019 [as D. pictus]   Dendrelaphis tristis Common Indian Bronzeback Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2012 Wangyal 2019 Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe cantoris Eastern Trinket Snake Hobcroft & Schulz 2010 Koirala, Gurung, Lhendup & Phuntsho 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019   Elaphe hodgsonii Himalayan Trinket Snake Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Elaphe taeniura Stripe-tailed Rat Snake Koirala, Gurung, Lhendup & Phuntsho 2016 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019 Genus Fowlea (Fishing Keelbacks)   Fowlea piscator Checkered Keelback Bauer & Gunther 1992 Das 1996 Mitra 2009 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2011 Wangyal 2019 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma frenatum Khasi Hills Rat Snake Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Gonyosoma prasinum Green Bush Rat Snake Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020 Genus Hebius (Oriental Equatorial Keelbacks)   Hebius clerki Kachin Keelback Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Hebius parallelus Parallel-striped Keelback Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Wangyal 2019 Genus Herpetoreas (Himalayan Mountain Keelbacks)   Herpetoreas platyceps Himalayan Flat-headed Keelback Bauer & Gunther 1992 Das 1996 Koirala, Gurung, Lhendup & Phuntsho 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Ren, Jiang, Huang, David & Li 2022 Tshewang & Letro 2018 Wallach, Williams & Boundy 2014 Wangyal 2011 Wangyal 2013 Wangyal 2019   Herpetoreas sieboldii Sikkim Keelback Ren, Jiang, Huang, David & Li 2022 Wallach, Williams & Boundy 2014 Wangyal 2011 Wangyal 2019 Genus Liopeltis (Ringneck Snakes)   Liopeltis rappii Himalayan Ringneck Snake Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020 Genus Lycodon (Oriental Wolf Snakes)   Lycodon aulicus Indian Wolf Snake Ahmed, Das & Dutta 2009 Ganesh & Vogel 2018 Koirala, Gurung, Lhendup & Phuntsho 2016 Tshewang & Letro 2018 Wangyal 2012 Wangyal 2019   Lycodon davisonii Burmese Bridle Snake Wangyal 2011 [as "cf. davisonii"] Wangyal 2019 [as "Dryocalamus cf. davisonii"]   Lycodon fasciatus Yellow-banded Wolf Snake Wangyal 2012 Wangyal 2019   Lycodon gammiei Sikkim Wolf Snake Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Wangyal 2013 Wangyal 2019   Lycodon jara Twin-spotted Wolf Snake Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2012 Wangyal 2019   Lycodon septentrionalis Northern Big-toothed Snake Das & Vasudevan 2015 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019   Lycodon zawi Sagaing Wolf Snake Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020 Genus Oligodon (Kukri Snakes)   Oligodon albocinctus White-barred Kukri Snake Ahmed, Das & Dutta 2009 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Wangyal 2011 Wangyal 2019   Oligodon chinensis Chinese Kukri Snake Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Oligodon cinereus Black-barred Kukri Snake Wangyal 2011 Wangyal 2019   Oligodon cyclurus Northeast Indian Kukri Snake Mitra 2009 Wangyal 2019   Oligodon dorsalis Spot-tailed Kukri Snake Das & Palden 2000 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2011 Wangyal 2019   Oligodon juglandifer Darjeeling Kukri Snake Mitra 2009 Sapkota & Sharma 2017 Wangyal 2019   Oligodon taeniolatus Variegated Kukri Snake Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Tshewang & Letro 2018 Wangyal 2019   Oligodon venustus Black-spotted Kukri Snake Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020 Genus Oreocryptophis (Red Mountain Rat Snake)   Oreocryptophis porphyraceus Red Mountain Rat Snake Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Wangyal 2011 Wangyal 2019 Genus Pseudoxenodon (Mock Cobras)   Pseudoxenodon macrops Common Large-eyed Mock Cobra Bauer & Gunther 1992 Das 1996 Koirala, Gurung, Lhendup & Phuntsho 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas korros Indochinese Rat Snake Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019   Ptyas mucosa Common Oriental Rat Snake Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Wangyal 2019   Ptyas nigromarginata Oriental Green Rat Snake Bauer & Gunther 1992 Das 1996 Koirala, Gurung, Lhendup & Phuntsho 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Tshewang & Letro 2018 Vogel & Hauser 2013 Wangyal 2011 Wangyal 2019 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis helleri Northern Red-necked Keelback Ahmed, Das & Dutta 2009 [as subminiatus] Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 [as subminiatus] Wangyal 2012 [as subminiatus] Wangyal 2019 [as R. subminiatus]   Rhabdophis himalayanus Common Himalayan Keelback Ahmed, Das & Dutta 2009 Das 1996 Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Koirala, Gurung, Lhendup & Phuntsho 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019 Genus Sibynophis (Oriental Black-headed Snakes)   Sibynophis collaris Collared Black-headed Snake Koirala, Gurung, Lhendup & Phuntsho 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019   Sibynophis sagittarius Bengal Black-headed Snake Das & Palden 2000 Mitra 2009 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2011 Wangyal 2019 Genus Trachischium (Oriental Slender Snakes)   Trachischium fuscum Darjeeling Slender Snake Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 [as "T. cf. fuscum"] Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Trachischium guentheri Rose-bellied Slender Snake Bauer & Gunther 1992 Das 1996 Mitra 2009 Wangyal 2011 Wangyal 2019   Trachischium laeve Olive Slender Snake Wangyal 2011 [as "cf. laeve"] Wangyal 2019 [as "T. cf. laeve"]   Trachischium monticola Assam Slender Snake Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020   Trachischium tenuiceps Orange-bellied Slender Snake Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Wangyal 2019  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Bungarus (Kraits)   Bungarus bungaroides Northeastern Hill Krait Koirala, Gurung, Lhendup & Phuntsho 2016 Wangyal 2011 Wangyal 2019   Bungarus caeruleus Blue Krait Tshewang & Letro 2018   Bungarus fasciatus Banded Krait Ahmed, Das & Dutta 2009 Das & Palden 2000 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2011 Wangyal 2019   Bungarus lividus Lesser Black Krait Lalbiakzuala, Purkayastha, Rathee & Lalremsanga 2021 Tshewang & Letro 2018   Bungarus niger Greater Black Krait Ahmed, Das & Dutta 2009 Bauer & Gunther 1992 Das 1996 Koirala, Gurung, Lhendup & Phuntsho 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Wangyal 2011 Wangyal 2019   Bungarus walli Faizabad Krait Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020 Genus Naja (Typical Cobras)   Naja kaouthia Monocellate Cobra Ahmed, Das & Dutta 2009 Koirala, Gurung, Lhendup & Phuntsho 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019   Naja naja Indian Spectacled Cobra Das & Palden 2000 Mitra 2009 Sindaco, Venchi & Grieco 2013 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2011 Wangyal 2012 Wangyal 2019 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Ahmed, Das & Dutta 2009 Bauer & Gunther 1992 Biswas 1976 Das 1996 Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019 Genus Sinomicrurus (East Asian Coral Snakes)   Sinomicrurus macclellandii Southeast Asian Coral Snake Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019  Family Homalopsidae (Australasian Mud Snakes) Genus Enhydris (Asian Mud Snakes)   Enhydris enhydris Striped Mud Snake Wangyal 2011 Wangyal 2019  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Ahmed, Das & Dutta 2009 Das & Palden 2000 Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2011 Wangyal 2019  Family Pareidae (Oriental Slug Eaters) Genus Pareas (Typical Oriental Slug Eaters)   Pareas monticola Assam Slug Eater Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Genus Python (Typical Pythons)   Python bivittatus Burmese Python Ahmed, Das & Dutta 2009 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Tshewang & Letro 2018 Wangyal 2012 Wangyal 2019   Python molurus Indian Python Bauer & Gunther 1992 Das 1996 Mitra 2009 Wangyal 2011 Wangyal 2019  Family Typhlopidae (Typical Blind Snakes) Genus Argyrophis (Southeast Asian Blind Snakes)   Argyrophis diardii Greater Blind Snake Ahmed, Das & Dutta 2009 Wangyal 2019 Genus Indotyphlops (South Asian Blind Snakes)   Indotyphlops jerdoni Khasi Hills Blind Snake Das & Palden 2000 Wangyal 2011 Wangyal 2019 Genus Virgotyphlops (Brahminy Blind Snake)   Virgotyphlops braminus Brahminy Blind Snake Bauer & Gunther 1992 Das 1996 Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Wallach 2009 Wallach 2020 Wangyal 2011 Wangyal 2012 Wangyal 2019  Family Viperidae (Vipers) Genus Daboia (Daboias)   Daboia russelii Indian Chain Viper David & Ineich 1999 Mitra 2009 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2019 Genus Gloydius (Mamushis)   Gloydius himalayanus Himalayan Mamushi Chaudhuri, Muhkerjee, Chowdhury & Purkayastha 2018 Koirala, Gurung, Lhendup & Phuntsho 2016 Wangyal 2011 Wangyal 2019 Genus Himalayophis (Himalayan Pitvipers)   Himalayophis tibetanus Tibetan Bamboo Pitviper Wangyal 2019 Genus Ovophis (Asian Mountain Pitvipers)   Ovophis monticola Western Mountain Pitviper Ahmed, Das & Dutta 2009 Koirala, Gurung, Lhendup & Phuntsho 2016 Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Tshewang & Letro 2018 Wangyal 2011 Wangyal 2019 Genus Popeia (South Asian Bamboo Pitvipers)   Popeia popeiorum Northern Bamboo Pitviper Das, Sharma, Surendran, Nath, Ghosh, Dutta, Mondol & Wangdi 2016 Mirza, Lalremsanga, Bhosale, Gowande, Patel, Idiatullina & Poyarkov 2023 Genus Protobothrops (Habus and allies)   Protobothrops himalayanus Himalayan Habu Koirala, Gurung, Lhendup & Phuntsho 2016 Pan, Chettri, Yang, Jiang, Wang, Zhang & Vogel 2013 Tshewang & Letro 2018 Wangyal 2019   Protobothrops jerdonii Spotted Mountain Pitviper Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Mitra 2009 Wangyal 2011 Wangyal 2019   Protobothrops kaulbacki Burmese Lance-headed Pitviper Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020 Genus Trimeresurus (Oriental Pitvipers)   Trimeresurus albolabris Southeast Asian White-lipped Tree Pitviper Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2012 Wangyal 2019   Trimeresurus erythrurus Red-tailed Pitviper Mitra 2009 Wangyal 2011 Wangyal 2019   Trimeresurus salazar Seijosa Pitviper Koirala, Jamtsho, Wangdi, Tshering, Wangdi, Norbu, Phuntsho, Lhendup & Nidup 2021 [as "T. cf. salazar"] Genus Viridovipera (Green Tree Pitvipers)   Viridovipera yunnanensis Yunnan Green Tree Pitviper Wangyal 2019 [as "Trimeresurus cf. stejnegeri yunnanensis"]

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wangyal 2019 Wangyal, Wangchuk & Das 2012   Cuora mouhotii Keeled Box Turtle Das, McCormack, Dijk, Hoang & Struijk 2016 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Tshewang & Letro 2018 Wangyal 2019 Wangyal, Wangchuk & Das 2012 Genus Cyclemys (Asian Leaf Turtles)   Cyclemys gemeli Assam Leaf Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wangyal 2019 Wangyal, Wangchuk & Das 2012 Genus Melanochelys (Indian Black Turtles)   Melanochelys tricarinata Tricarinate Hill Turtle Ahmed, Das & Dutta 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wangyal 2019 Wangyal, Wangchuk & Das 2012   Melanochelys trijuga Indian Black Turtle Wangyal 2013 Wangyal 2019 Genus Pangshura (Roofed Turtles)   Pangshura sylhetensis Assam Roofed Turtle Wangyal, Bower, Sherub, Tshewang, Wangdi & al. 2020  Family Testudinidae (Tortoises) Genus Indotestudo (Asian Tortoises)   Indotestudo elongata Elongated Tortoise Drukpa, Kinley, Tamang, Gyeltshen, Dema, Phuntsho, Thapa, Dorji & Letro 2020 Ihlow, Dawson, Hartmann & Som 2016 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Tenzin, Nidup, Gyeltsen, Tshomo, Nepal & Wangyal 2022 Wangyal 2011 Wangyal 2019 Wangyal, Wangchuk & Das 2012