Reptiles of Hainan [China] Related checklists:  China Bibliography of the reptiles of Hainan Taxonomic history of the reptiles of Hainan Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthosaura (Pricklenapes)   Acanthosaura lepidogaster Brown Pricklenape Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Manthey 2008 [as "cf. lepidogaster"] Zhao & Adler 1993 Zhao 2005 Genus Calotes (South Asian Forest Agamas)   Calotes versicolor South Indian Crested Agama Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Manthey 2008 Zhao & Adler 1993 Zhao 2005 Genus Draco (Gliding Dragons)   Draco maculatus Black-spotted Gliding Dragon Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Cox, Dijk, Nabhitabhata & Thirakhupt 1998 Lazell 1999 Lu, Wang & Shi 2004 Manthey 2008 [as "cf. maculatus"] McGuire & Kiew 2001 Musters 1983 Wermuth 1967 Zhao & Adler 1993 Zhao 2005 Genus Leiolepis (Butterfly Agamas)   Leiolepis reevesii Eastern Butterfly Agama Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Peters 1971 Zhao & Adler 1993 Zhao 2005 Genus Pseudocalotes (Long-headed Agamas)   Pseudocalotes microlepis Small-scaled Long-headed Agama Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Hallermann & Boehme 2000 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993  Family Anguidae (Slowworms, etc.) Genus Dopasia (Oriental Glass Lizards)   Dopasia hainanensis Hainan Glass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Nguyen, Boehme, Nguyen, Le, Pahl, Haus & Ziegler 2011 Wang, Shi, Wang, Xie, Li & Zou 2007 Zhao & Adler 1993   Dopasia harti South Chinese Glass Lizard Wang, Shi, Wang, Xie, Li & Zou 2007  Family Dibamidae (Blind Skinks) Genus Dibamus (Oriental Blind Skinks)   Dibamus sp. (Oriental Blind Skink) Lu, Wang & Shi 2004  Family Eublepharidae (Eyelid Geckos) Genus Goniurosaurus (East Asian Leopard Geckos)   * Goniurosaurus bawanglingensis Bawangling Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Grismer, Shi, Orlov & Ananjeva 2002 Lu, Wang & Shi 2004 Seufer, Kaverkin & Kirschner (eds.) 2005   * Goniurosaurus hainanensis Hainan Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Grismer, Shi, Orlov & Ananjeva 2002 Grismer, Viets & Boyle 1999 Lazell 1999 [as lichtenfelderi] Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005 [as lichtenfelderi]   * Goniurosaurus sinensis Kwang Hua Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhou, Peng, Hou & Yuan 2019 Zhu & He in Zhu, Shen, Liu, Chen, Li & He 2020 [as kwanghua]   * Goniurosaurus zhoui Zhou's Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhou, Wang, Chen & Liang 2018  Family Gekkonidae (Typical Geckos) Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005 Genus Gekko (Tokays)   Gekko gecko Tokay Lu, Wang & Shi 2004   Gekko reevesii Chinese Black Tokay Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021   Gekko similignum Hainan Narrow-disked Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 [as chinensis] Lyu, Lin, Ren, Jiang, Zhang, Qi & Wang 2021 Ota, Lau, Weidenhoefer, Yasukawa & Bogadek 1995 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Zhao 2005 [as chinensis] Genus Hemidactylus (Half-toed Geckos)   Hemidactylus bowringii Hong Kong House Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005   Hemidactylus frenatus Asian House Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005   Hemidactylus garnotii Fox Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005 Genus Hemiphyllodactylus (Slender Geckos)   Hemiphyllodactylus typus Indo-Pacific Slender Gecko Zug 2010  Family Lacertidae (Typical Lizards) Genus Takydromus (Oriental Grass Lizards)   Takydromus kuehnei Kanshirei Grass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993   Takydromus sexlineatus Six-lined Grass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005  Family Scincidae (Skinks) Genus Ateuchosaurus (East Asian Short-legged Skinks)   Ateuchosaurus chinensis Chinese Short-Legged Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005 Genus Eutropis (South Asian Sun Skinks)   Eutropis longicaudata Long-tailed Sun Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005   Eutropis multifasciata Many-lined Sun Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 O'Shea, Kusuma & Kaiser 2018 Zhao & Adler 1993 Zhao 2005 Genus Lygosoma (Southeast Asian Writhing Skinks)   Lygosoma siamense Siamese Writhing Skink Qi, Ren, Wang, Yang & Jiang 2022 Genus Plestiodon (Blue-tailed Skinks)   Plestiodon chinensis Eastern Chinese Long Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993   Plestiodon quadrilineatus Four-striped Long Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005 Genus Scincella (South Asian and American Ground Skinks)   Scincella modesta Slender Ground Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021   Scincella reevesii Speckled Ground Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus incognitus Koshun Forest Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005   Sphenomorphus indicus Common Indian Forest Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993   Sphenomorphus tonkinensis Tonkin Forest Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Nguyen, Schmitz, Nguyen, Orlov, Boehme & Ziegler 2011 Wang, Yang & Liu 2013 Genus Tropidophorus (Stream Skinks)   Tropidophorus hainanus Hainan Stream Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Guo, Shu, Wu, Chen, Hou, Shi & Deng 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus salvator Asian Water Monitor Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lu, Wang & Shi 2004 Zhao & Adler 1993 Zhao 2005

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Acrochordidae (Australasian File Snakes) Genus Acrochordus (Australasian File Snakes)   Acrochordus granulatus Lesser File Snake Wallach, Williams & Boundy 2014 Zhao & Adler 1993  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake David, Pauwels, Lays & Lenglet 2006 Genus Amphiesma (South Asian Keelbacks)   Amphiesma stolatum Buff-striped Keelback Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Amphiesmoides (White-eyed Keelback)   Amphiesmoides ornaticeps White-eyed Keelback Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Genus Boiga (Australasian Cat Snakes)   Boiga kraepelini Kelung Cat Snake Orlov & Ryabov 2002 Zhao & Adler 1993 Zhao, Zhao, Shi & Wang 2004   Boiga multomaculata Marbled Cat Snake Orlov & Ryabov 2002 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Calamaria (Oriental Reed Snakes)   Calamaria pavimentata Brown Reed Snake Zhao & Adler 1993   Calamaria septentrionalis Northern Reed Snake Inger & Marx 1965 Zhao & Adler 1993 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea ornata Ornate Flying Tree Snake Zhao & Adler 1993 Zhao & Borkii 1990 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis ngansonensis Nganson Bronzeback Biakzuala, Lalremsanga, Tariang, Vabeiryureilai, Muansanga, Hrima, Kumar, Kundu, Purkayastha & Vogel 2022 Nicodemo & Bain 2007 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020   Dendrelaphis proarchos Northern Painted Bronzeback Biakzuala, Lalremsanga, Tariang, Vabeiryureilai, Muansanga, Hrima, Kumar, Kundu, Purkayastha & Vogel 2022 Zhao & Adler 1993 [as pictus] Zhao 2005 [as pictus] Zhao, Zhao, Shi & Wang 2004 [as pictus] Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe taeniura Stripe-tailed Rat Snake Schulz 1996 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Fowlea (Fishing Keelbacks)   Fowlea flavipunctata Yellow-spotted Keelback Smith 1943 Vogel & David 2012 Zhao & Adler 1993 [as piscator] Zhao 2005 [as piscator] Zhao, Zhao, Shi & Wang 2004 [as piscator] Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma coeruleum Blue-eyed Bush Rat Snake David, Campbell, Deuti, Hauser, Luu, Nguyen, Orlov, Pauwels, Scheinberg, Sethy, Smits, Teynie & Vogel 2022 Schulz 1996 [as prasinum] Zhao & Adler 1993 [as prasinum] Zhao 2005 [as prasinum] Zhao, Zhao, Shi & Wang 2004 [as prasinum]   * Gonyosoma hainanense Hainan Rhinoceros Snake Peng, Zhang, Huang, Burbrink, Chen, Hou, Zhu, Yang & Wang 2021 Zhao & Adler 1993 [as Rhynchophis boulengeri] Zhao 2005 [as Rhynchophis boulengeri] Genus Hebius (Oriental Equatorial Keelbacks)   Hebius boulengeri White-browed Keelback David, Bain, Nguyen, Orlov, Vogel, Vu & Ziegler 2007 [as "cf. boulengeri"] David, Vogel & Rooijen 2013 Lalremsanga, Bal, Vogel & Biakzuala 2022 Purkayastha & David 2019 Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Zhao, Zhao, Shi & Wang 2004   Hebius popei Chinese Mountain Keelback Lalremsanga, Bal, Vogel & Biakzuala 2022 Malnate 1962 Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Zhao 2005 Zhou, Sun, Qi, Lu, Lyu, Wang, Li & Ma 2019   Hebius sauteri Kosempo Keelback David, Bain, Nguyen, Orlov, Vogel, Vu & Ziegler 2007 David, Vogel & Rooijen 2013 Purkayastha & David 2019 Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Zhao, Zhao, Shi & Wang 2004 Genus Lycodon (Oriental Wolf Snakes)   Lycodon capucinus Island Wolf Snake Zhang, Jiang, Vogel & Rao 2011 [as aulicus]   Lycodon flavozonatus Yellow Big-toothed Snake Zhao & Adler 1993   Lycodon rosozonatus Rosy-banded Big-toothed Snake Guo, Liu, Li, Zhong, Chen & Wang 2012 Hu & Zhao 1975 Luo, Ryabov, Luo, Gao, Luo & Hu 2010 Orlov & Ryabov 2004 Wang, Qi, Lyu, Zeng & Wang 2020 Zhao & Adler 1993 Zhao 2005   Lycodon rufozonatus Red-banded Big-toothed Snake Zhao & Adler 1993   Lycodon ruhstrati Taiwanese Wolf Snake Zhang, Jiang, Vogel & Rao 2011   Lycodon subcinctus Malayan Banded Wolf Snake Wang, Qi, Lyu, Zeng & Wang 2020 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Oligodon (Kukri Snakes)   * Oligodon bivirgatus Two-striped Kukri Snake Qian, Qi, Shi, Lu, Jenkins, Mo & Li 2021   Oligodon chinensis Chinese Kukri Snake Zhao & Adler 1993 Zhao, Zhao, Shi & Wang 2004   Oligodon cinereus Black-barred Kukri Snake David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012 Qian, Qi, Shi, Lu, Jenkins, Mo & Li 2021 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004   Oligodon formosanus Taiwanese Kukri Snake Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Opisthotropis (Mountain Stream Keelbacks)   Opisthotropis lateralis Bicoloured Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Genus Oreocryptophis (Red Mountain Rat Snake)   Oreocryptophis porphyraceus Red Mountain Rat Snake Schulz & Helfenberger 1998 Schulz 1996 Smith 1943 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Pseudoxenodon (Mock Cobras)   Pseudoxenodon bambusicola Bamboo Mock Cobra Zhao & Adler 1993 Zhao 2005   Pseudoxenodon karlschmidti Common Chinese Mock Cobra Zhao & Adler 1993 Zhao 2005 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas korros Indochinese Rat Snake Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004   Ptyas major Greater Green Snake Zhao & Adler 1993 Zhao 2005   Ptyas mucosa Common Oriental Rat Snake Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004   Ptyas multicincta Many-banded Green Snake Hauser 2018 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Rhabdophis (Asian Keelbacks)   * Rhabdophis adleri Lingshui Keelback Guo, Liu, Li, Zhong, Chen & Wang 2012 Zhao 1997 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Zhu, Wang, Takeuchi & Zhao 2014   Rhabdophis callichroma Bavi Keelback Zhu, Wang, Takeuchi & Zhao 2014   * Rhabdophis confusus Hainan Keelback David & Vogel 2021 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 [as subminiatus] Zhao & Adler 1993 [as subminiatus] Zhao 2005 [as subminiatus] Zhao, Zhao, Shi & Wang 2004 [as subminiatus] Zhu, Wang, Takeuchi & Zhao 2014 [as subminiatus] Genus Sibynophis (Oriental Black-headed Snakes)   Sibynophis chinensis Chinese Black-headed Snake Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Trimerodytes (Asian Water Snakes)   Trimerodytes aequifasciatus Equal-banded Water Snake Zhao & Adler 1993 Zhao 2005   Trimerodytes annularis Asian Banded Water Snake Zhao & Adler 1993   Trimerodytes balteatus Banded Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019 Zhao & Adler 1993   Trimerodytes percarinatus Chinese Keelback Guo, Zhu, Liu, Wang, Che & Nguyen 2020 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004  Family Cylindrophiidae (Asian Pipe Snakes) Genus Cylindrophis (Oriental Pipe Snakes)   Cylindrophis ruffus Red-tailed Pipe Snake Adler, Zhao & Darevsky 1992 McDiarmid, Campbell & Toure 1999 Zhao & Adler 1993 Zhao & Darevsky 1990  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Bungarus (Kraits)   Bungarus fasciatus Banded Krait Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004   Bungarus multicinctus Many-banded Krait Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Naja (Typical Cobras)   Naja atra Chinese Spitting Cobra Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Sinomicrurus (East Asian Coral Snakes)   Sinomicrurus annularis Hong Kong Coral Snake Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021   Sinomicrurus kelloggi Indochinese Coral Snake Wang, Peng & Huang in Peng, Wang, Ding, Zhu, Luo, Yang, Huang, Lu & Huang 2018 [as houi] Zhao & Adler 1993 Zhao 2005  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis atriceps Black-headed Sea Snake Zhao & Adler 1993 [as H. fasciatus] Zhao 2005   Hydrophis curtus Short Sea Snake Zhao & Adler 1993 Zhao 2005   Hydrophis cyanocinctus Blue-banded Sea Snake Zhao & Adler 1993 Zhao 2005   Hydrophis ornatus Reef Sea Snake Zhao & Adler 1993 Zhao 2005   Hydrophis platurus Yellow-bellied Sea Snake Zhao & Adler 1993   Hydrophis viperinus Viperine Sea Snake Zhao & Adler 1993 Zhao 2005 Genus Microcephalophis (Small-headed Sea Snakes)   Microcephalophis gracilis Graceful Small-headed Sea Snake Zhao & Adler 1993 Zhao 2005  Family Homalopsidae (Australasian Mud Snakes) Genus Hypsiscopus (Orange-bellied Mud Snake and Matanna Mud Snake)   Hypsiscopus murphyi Northern Orange-bellied Mud Snake Bernstein, Voris, Stuart, Phimmachak, Seateun, Sivongxay, Neang, Karns, Andrews, Osterhage, Phipps & Ruane 2022 Gyi 1970 [as plumbea] Zhao & Adler 1993 [as plumbea] Zhao 2005 [as plumbea] Zhao, Zhao, Shi & Wang 2004 [as plumbea] Genus Myrrophis (Indochinese Mud Snakes)   Myrrophis bennetti Mangrove Mud Snake Gyi 1970 Murphy & Voris 2014 Zhao & Adler 1993 Zhao, Zhao, Shi & Wang 2004   Myrrophis chinensis Chinese Mud Snake Gyi 1970 Murphy & Voris 2014 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004  Family Pareidae (Oriental Slug Eaters) Genus Pareas (Typical Oriental Slug Eaters)   Pareas formosensis Taiwanese Red-eyed Slug Eater Ding, Chen, Suwannapoom, Nguyen, Pogarkov & Vogel 2020 Le, Tran, Hoang & Stuart 2021   Pareas macularius Black-spotted Slug Eater Hauser 2017 Vogel, Nguyen, Lalremsanga, Biakzuala, Hrima & Poyarkov 2020   Pareas margaritophorus White-spotted Slug Eater Hauser 2017 Vogel, Nguyen, Lalremsanga, Biakzuala, Hrima & Poyarkov 2020 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python bivittatus Burmese Python Barker & Barker 2010 Stimson 1969 Zhao & Adler 1993 Zhao 2005  Family Typhlopidae (Typical Blind Snakes) Genus Argyrophis (Southeast Asian Blind Snakes)   Argyrophis diardii Greater Blind Snake Hahn 1980 Zhao & Adler 1993 Genus Virgotyphlops (Brahminy Blind Snake)   Virgotyphlops braminus Brahminy Blind Snake McDiarmid, Campbell & Toure 1999 McDowell 1974 Nanhoe & Ouboter 1987 Wallach 2009 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004  Family Viperidae (Vipers) Genus Ovophis (Asian Mountain Pitvipers)   Ovophis tonkinensis Tonkin Pitviper David 1995 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Malhotra, Dawson, Guo & Thorpe 2011 Genus Protobothrops (Habus and allies)   Protobothrops mucrosquamatus Brown-spotted Pitviper Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Trimeresurus (Oriental Pitvipers)   Trimeresurus albolabris Southeast Asian White-lipped Tree Pitviper Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Hoge & Romano Hoge 1981 Regenass & Kramer 1981 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004 Genus Viridovipera (Green Tree Pitvipers)   Viridovipera gumprechti Phu Luang Green Tree Pitviper Gumprecht, Tillack, Orlov, Captain & Ryabov 2004   Viridovipera stejnegeri Chinese Green Tree Pitviper David, Vidal & Pauwels 2001 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Guo, Liu, Li, Cao & Wang 2016 Vogel 2006 Zhao & Adler 1993 Zhao 1997 Zhao 2005 Zhao, Zhao, Shi & Wang 2004  Family Xenodermidae (Asian Litter Snakes) Genus Achalinus (East Asian Burrowing Snakes)   * Achalinus hainanus Hainan Burrowing Snake Hu & Zhao 1975 Wallach, Williams & Boundy 2014 Zhao & Adler 1993   Achalinus rufescens Rufous Burrowing Snake Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Zhao, Zhao, Shi & Wang 2004  Family Xenopeltidae (Asian Sunbeam Snakes) Genus Xenopeltis (Asian Sunbeam Snakes)   Xenopeltis hainanensis Hainan Sunbeam Snake Guo, Liu, Li, Zhong, Chen & Wang 2012 McDiarmid, Campbell & Toure 1999 Orlov 2000 Zhao & Adler 1993 Zhao 2005 Zhao, Zhao, Shi & Wang 2004

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Huang in Bjorndal (ed.) 1982 Zhao & Adler 1993 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Huang in Bjorndal (ed.) 1982 Jia, Wang, Balazs & Liu 2019 Zhao & Adler 1993 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Zhao & Adler 1993 Zhao 2005 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Zhao & Adler 1993  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora galbinifrons Northern Indochinese Box Turtle Iverson 1992 Lau & Shi 2000 McCord & Joseph-Ouni 2002 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 Zhao 2005   Cuora mouhotii Keeled Box Turtle Das 1987 Das, McCormack, Dijk, Hoang & Struijk 2016 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 Zhao 2005   Cuora trifasciata Northern Three-striped Box Turtle Blanck, Ting, Protiva, Crow & Tiedemann 2017 Fritz & Havas 2006 Iverson 1992 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 Zhao 2005 Genus Geoemyda (Black-breasted Leaf Turtles)   Geoemyda spengleri Common Black-breasted Leaf Turtle Dawson, Gaillard, Gong, Lin, McCormack, Nanthavong, Nguyen, Nguyen, Pham & Shi 2022 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Yasukawa & Ota 2010 Zhao & Adler 1993 Genus Mauremys (Stripe-necked Turtles)   Mauremys mutica Yellow Pond Turtle Iverson 1992 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 Zhao 2005   Mauremys sinensis Chinese Stripe-necked Turtle Anonymous 2013 Iverson 1992 Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 Zhao 2005 Genus Sacalia (Eastern Eyed Turtles)   Sacalia bealei Eastern Chinese Eyed Turtle Lau & Shi 2000 Zhao & Adler 1993   Sacalia quadriocellata Four-eyed Turtle Iverson 1992 Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 Zhao 2005  Family Platysternidae (Big-headed Turtle) Genus Platysternon (Big-headed Turtle)   Platysternon megacephalum Big-headed Turtle Iverson 1992 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 Zhao 2005  Family Trionychidae (Softshell Turtles) Genus Palea (Wattle-necked Softshell Turtle)   Palea steindachneri Wattle-necked Softshell Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelochelys (Indomalayan Giant Softshell Turtles)   Pelochelys cantorii Asian Giant Softshell Turtle Das 2008 Fritz & Havas 2006 Iverson 1992 [as bibroni] Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 [as bibroni] Zhao 2005 Genus Pelodiscus (East Asian Softshell Turtles)   Pelodiscus variegatus Large-spotted Softshell Turtle Farkas, Ziegler, Pham, Ong & Fritz 2019 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021