Reptiles of Guangdong [China] Related checklists:  China Bibliography of the reptiles of Guangdong Taxonomic history of the reptiles of Guangdong Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthosaura (Pricklenapes)   Acanthosaura lepidogaster Brown Pricklenape Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Xu 2007 Zhao & Adler 1993 Genus Calotes (South Asian Forest Agamas)   Calotes emma Forest Crested Agama Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Xu 2007 Zhao & Adler 1993   Calotes versicolor South Indian Crested Agama Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Li, Xiao, Qing, Lu & Lazell 2009 Xu 2007 Zhao & Adler 1993 Genus Diploderma (Eastern Mountain Agamas)   Diploderma fasciatum Banded Mountain Agama Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 [as Diploderma szechwanensis] Wang, Qi, Wang, Koehler, Lu, Lyu, Wang, Wang & Che 2022 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Wang, Ren, Wu, Jiang, Jin, Hou, Zheng, Xie, Siler & Che 2021 Genus Leiolepis (Butterfly Agamas)   Leiolepis reevesii Eastern Butterfly Agama Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Zhao & Adler 1993 Genus Physignathus (Indochinese Water Dragon)   Physignathus cocincinus Indochinese Water Dragon Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993 Genus Pseudocalotes (Long-headed Agamas)   Pseudocalotes microlepis Small-scaled Long-headed Agama Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Xu 2007  Family Anguidae (Slowworms, etc.) Genus Dopasia (Oriental Glass Lizards)   Dopasia harti South Chinese Glass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021  Family Dibamidae (Blind Skinks) Genus Dibamus (Oriental Blind Skinks)   Dibamus bourreti Indochinese Blind Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021  Family Eublepharidae (Eyelid Geckos) Genus Goniurosaurus (East Asian Leopard Geckos)   * Goniurosaurus gollum Huaiji Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Qi, Wang, Grismer, Chen, Lyu & Wang 2020   * Goniurosaurus varius Nanling Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Qi, Grismer, Lyu, Zhang, Li & Wang 2020   * Goniurosaurus wangshu Wangshu Leopard Gecko Zhu, Wu, Liu, Bai, Roman-Palacios, Li & He 2022   * Goniurosaurus yingdeensis Yingde Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Wang, Yang & Cui 2010   * Goniurosaurus zhelongi Shimentai Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Wang, Jin, Li & Grismer 2014  Family Gekkonidae (Typical Geckos) Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 David, Pauwels, Lays & Lenglet 2006 Genus Gekko (Tokays)   Gekko chinensis Chinese Narrow-disked Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Li, Xiao, Qing, Lu & Lazell 2009 Lyu, Lin, Ren, Jiang, Zhang, Qi & Wang 2021 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Xu 2007 Zhao & Adler 1993   Gekko gecko Tokay Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Xu 2007 Zhao & Adler 1993   Gekko hokouensis Hokou Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021   Gekko japonicus Japanese Giant Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021   Gekko melli Southern Chinese Palmated Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 [as "cf. subpalmatus"] Lazell, Kolby, Lin, Zhuang & Lu 1999 [as G. subpalmatus] Lyu, Lin, Ren, Jiang, Zhang, Qi & Wang 2021 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 [as G. subpalmatus] Roesler & Tiedemann 2007 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 [as G. subpalmatus] Xu 2007 [as "cf. subpalmatus"] Yang, Wang, Zhang, Sun & Lin 2012 Zhao & Adler 1993 [as G. subpalmatus]   Gekko reevesii Chinese Black Tokay Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lyu, Lin, Ren, Jiang, Zhang, Qi & Wang 2021 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus bowringii Hong Kong House Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lazell, Kolby, Lin, Zhuang & Lu 1999 Lazell, Lau & Lu 1997 Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xu 2007 Zhao & Adler 1993   Hemidactylus frenatus Asian House Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Li, Xiao, Qing, Lu & Lazell 2009 Zhao & Adler 1993   Hemidactylus garnotii Fox Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993   Hemidactylus platyurus Flat-tailed Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993  Family Lacertidae (Typical Lizards) Genus Takydromus (Oriental Grass Lizards)   * Takydromus albomaculosus White-spotted Grass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Wang, Gong, Liu & Wang 2017 Wang, Lyu, Yang, Li & Wang 2019   Takydromus intermedius Sichuan Grass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021   Takydromus kuehnei Kanshirei Grass Lizard Bischoff 1992 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Wang, Lyu, Yang, Li & Wang 2019 Xu 2007 Zhao & Adler 1993   Takydromus septentrionalis Northern Grass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Wang, Lyu, Yang, Li & Wang 2019 Xu 2007 Zhao & Adler 1993   Takydromus sexlineatus Six-lined Grass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Wang, Lyu, Yang, Li & Wang 2019 Xu 2007 Zhao & Adler 1993   Takydromus sylvaticus Fujian Grass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Yang & Wang 2010   * Takydromus yunkaiensis Yunkai Grass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Wang, Lyu, Yang, Li & Wang 2019  Family Scincidae (Skinks) Genus Ateuchosaurus (East Asian Short-legged Skinks)   Ateuchosaurus chinensis Chinese Short-Legged Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lazell, Kolby, Lin, Zhuang & Lu 1999 Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xu 2007 Zhao & Adler 1993 Genus Eutropis (South Asian Sun Skinks)   Eutropis longicaudata Long-tailed Sun Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Zhao & Adler 1993   Eutropis multifasciata Many-lined Sun Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 David, Pauwels, Lays & Lenglet 2006 Genus Plestiodon (Blue-tailed Skinks)   Plestiodon chinensis Eastern Chinese Long Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lazell, Kolby, Lin, Zhuang & Lu 1999 Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xu 2007 Zhao & Adler 1993   Plestiodon elegans Elegant Five-lined Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xu 2007 Zhao & Adler 1993   Plestiodon liui Jiangsu Blue-tailed Skink Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015   Plestiodon quadrilineatus Four-striped Long Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell, Lau & Lu 1997 Li, Xiao, Qing, Lu & Lazell 2009 Xu 2007 Zhao & Adler 1993 Genus Scincella (South Asian and American Ground Skinks)   Scincella modesta Slender Ground Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell, Lau & Lu 1997 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015   Scincella reevesii Speckled Ground Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Li, Xiao, Qing, Lu & Lazell 2009 Xu 2007 Zhao & Adler 1993 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus incognitus Koshun Forest Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Li, Xiao, Qing, Lu & Lazell 2009   Sphenomorphus indicus Common Indian Forest Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lazell 1999 Lazell, Kolby, Lin, Zhuang & Lu 1999 Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xu 2007 Zhao & Adler 1993   Sphenomorphus tonkinensis Tonkin Forest Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Wang, Yang & Liu 2013 Genus Tropidophorus (Stream Skinks)   Tropidophorus hainanus Hainan Stream Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Guo, Shu, Wu, Chen, Hou, Shi & Deng 2021 Rao, Pan, Jiang, Hu & Zhang 2011   Tropidophorus sinicus South Chinese Stream Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Guo, Shu, Wu, Chen, Hou, Shi & Deng 2021 Li, Xiao, Qing, Lu & Lazell 2009 Zhao & Adler 1993  Family Shinisauridae (Chinese Crocodile Lizard) Genus Shinisaurus (Crocodile Lizard)   Shinisaurus crocodilurus Crocodile Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Li & Xiao 2002 Schingen, Ihlow, Nguyen, Ziegler, Bonkowski, Wu & Roedder 2014 Xu 2007 Ziegler, Le, Vu, Hendrix & Boehme 2008  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus salvator Asian Water Monitor Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Xu 2007

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake Li, Xiao, Qing, Lu & Lazell 2009 Wallach, Williams & Boundy 2014 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Zhao & Adler 1993 Genus Amphiesma (South Asian Keelbacks)   Amphiesma stolatum Buff-striped Keelback Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Wallach, Williams & Boundy 2014 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Zhao & Adler 1993 Genus Archelaphe (Venus Rat Snake)   Archelaphe bella Venus Rat Snake Schulz, Boehme & Tillack 2011 Wallach, Williams & Boundy 2014 Genus Boiga (Australasian Cat Snakes)   Boiga kraepelini Kelung Cat Snake Orlov & Ryabov 2002   Boiga multomaculata Marbled Cat Snake Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xu 2007 Zhao & Adler 1993 Genus Calamaria (Oriental Reed Snakes)   * Calamaria arcana Lingnan Reed Snake Yeung, Lau & Yang 2022   Calamaria pavimentata Brown Reed Snake Inger & Marx 1965 Xu 2007 Zhao & Adler 1993   Calamaria septentrionalis Northern Reed Snake Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xu 2007 Zhao & Adler 1993 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea ornata Ornate Flying Tree Snake Deuve 1970 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus radiatus Copper-headed Rat Snake Lazell, Kolby, Lin, Zhuang & Lu 1999 Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Schulz 1996 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis proarchos Northern Painted Bronzeback Xie, Liu, Qiu, Zheng, Nie & Ma 1998 [as pictus] Zhao & Adler 1993 [as pictus] Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe carinata East Asian Keeled Rat Snake Schulz 1996 Xu 2007 Zhao & Adler 1993   Elaphe moellendorffi Red-headed Rat Snake Schulz 1996 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993   Elaphe taeniura Stripe-tailed Rat Snake Li, Xiao, Qing, Lu & Lazell 2009 Qi, Shi, Ma, Gao, Bu, Grismer, Li & Wang 2021 Schulz 1996 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993 Genus Euprepiophis (East Asian Rat Snakes)   Euprepiophis mandarinus Mandarin Rat Snake Schulz 1996 Xu 2007 Zhao & Adler 1993 Genus Fowlea (Fishing Keelbacks)   Fowlea flavipunctata Yellow-spotted Keelback Lazell, Kolby, Lin, Zhuang & Lu 1999 [as Natrix piscator] Li, Xiao, Qing, Lu & Lazell 2009 [as X. piscator] Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Vogel & David 2012 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 [as piscator] Xu 2007 [as piscator] Zhao & Adler 1993 [as piscator] Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma frenatum Khasi Hills Rat Snake Schulz 1996 Zhao & Adler 1993 Genus Hebius (Oriental Equatorial Keelbacks)   Hebius atemporalis Tonkin Keelback Lalremsanga, Bal, Vogel & Biakzuala 2022 Wallach, Williams & Boundy 2014 Zhao & Adler 1993   Hebius bitaeniatus Two-striped Keelback Gong, He & Yang 2009 Wallach, Williams & Boundy 2014 Xie, Liu, Qiu, Zheng, Nie & Ma 1998   Hebius boulengeri White-browed Keelback David, Bain, Nguyen, Orlov, Vogel, Vu & Ziegler 2007 [as "cf. boulengeri"] David, Vogel & Rooijen 2013 Lalremsanga, Bal, Vogel & Biakzuala 2022 Purkayastha & David 2019 Wallach, Williams & Boundy 2014 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Zhao & Adler 1993 Zhou, Sun, Qi, Lu, Lyu, Wang, Li & Ma 2019   Hebius craspedogaster Kuatun Keelback Malnate 1962 Wallach, Williams & Boundy 2014 Zhao & Adler 1993   Hebius optatus Mount Omei Keelback Xie, Liu, Qiu, Zheng, Nie & Ma 1998   Hebius popei Chinese Mountain Keelback Malnate 1962 Wallach, Williams & Boundy 2014 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Zhao & Adler 1993   Hebius sauteri Kosempo Keelback Malnate 1962 Wallach, Williams & Boundy 2014 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993 Genus Lycodon (Oriental Wolf Snakes)   Lycodon capucinus Island Wolf Snake O'Shea, Kusuma & Kaiser 2018 Zhao & Adler 1993 [as Ophites aulicus]   Lycodon flavozonatus Yellow Big-toothed Snake Wang, Qi, Lyu, Zeng & Wang 2020 Xu 2007 Zhao & Adler 1993   Lycodon futsingensis Fujian Wolf Snake Wang, Qi, Lyu, Zeng & Wang 2020 Zhang, Jiang, Hu, Yu, Peng, Tang & Hu 2011 Zhang, Jiang, Vogel & Rao 2011   Lycodon liuchengchaoi Tangjiahe Wolf Snake Li, Luo, Wei, Yu & Zhang 2012 [as L. fasciatus] Wang, Qi, Lyu, Zeng & Wang 2020   Lycodon meridionalis Vietnamese Big-toothed Snake Wang, Qi, Lyu, Zeng & Wang 2020   Lycodon rufozonatus Red-banded Big-toothed Snake Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xu 2007 Zhao & Adler 1993   Lycodon ruhstrati Taiwanese Wolf Snake Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler 2009 Xu 2007 Zhang, Jiang, Vogel & Rao 2011   Lycodon subcinctus Malayan Banded Wolf Snake Wang, Qi, Lyu, Zeng & Wang 2020 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993 Genus Oligodon (Kukri Snakes)   Oligodon catenatus Chain-banded Kukri Snake Gong, Auer, Zhang, Zhong & Zeng 2007 Qian, Qi, Shi, Lu, Jenkins, Mo & Li 2021   Oligodon chinensis Chinese Kukri Snake Qian, Qi, Shi, Lu, Jenkins, Mo & Li 2021 Xu 2007 Zhao & Adler 1993   Oligodon cinereus Black-barred Kukri Snake David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012 Li, Xiao, Qing, Lu & Lazell 2009 Zhao & Adler 1993   Oligodon formosanus Taiwanese Kukri Snake David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012 Lazell, Kolby, Lin, Zhuang & Lu 1999 Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xu 2007 Zhao & Adler 1993   Oligodon ornatus Chinese Ornate Kukri Snake Gong & He 2008 Genus Oocatochus (Red-backed Rat Snake)   Oocatochus rufodorsatus Red-backed Rat Snake Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Schulz 1996 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Genus Opisthotropis (Mountain Stream Keelbacks)   Opisthotropis andersonii Hong Kong Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019 Ren, Wang, Jiang, Guo & Li 2017 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Xu 2007   Opisthotropis cheni Guangdong Mountain Stream Keelback Guo, Peng & Gu 2008 Li, Liu & Guo 2010 Ren, Wang, Guo, Wang, Nguyen & Li 2019 Ren, Wang, Jiang, Guo & Li 2017 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Yang, Wang, Zhang, Lau & Chou 2011   Opisthotropis guangxiensis Guangxi Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017   Opisthotropis hungtai Heishiding Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019 [as O. maculosa (part)] Ren, Wang, Jiang, Guo & Li 2017 [as O. maculosa] Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 [as O. maculosa] Wang, Lyu, Zeng, Lin, Yang, Nguyen, Le, Ziegler & Wang 2020 Yang, Wang, Zhang, Lau & Chou 2011 [as O. maculosa]   Opisthotropis kuatunensis Striped Mountain Stream Keelback David, Pauwels, Nguyen & Vogel 2015 Ren, Wang, Guo, Wang, Nguyen & Li 2019 Ren, Wang, Jiang, Guo & Li 2017 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Yang, Wang, Zhang, Lau & Chou 2011   Opisthotropis lateralis Bicoloured Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019 Ren, Wang, Jiang, Guo & Li 2017 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Yang, Wang, Zhang, Lau & Chou 2011   Opisthotropis latouchii South Chinese Mountain Stream Keelback David, Pauwels, Nguyen & Vogel 2015 Ren, Wang, Guo, Wang, Nguyen & Li 2019 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Yang, Wang, Zhang, Lau & Chou 2011 Zhao & Adler 1993   * Opisthotropis laui Gudou Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Wang, Lyu, Zeng, Liu & Wang 2017 Yang, Sung & Chan 2013   Opisthotropis maxwelli Fujian Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019 Ren, Wang, Jiang, Guo & Li 2017 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Xu 2007 Zhao & Adler 1993   * Opisthotropis shenzhenensis Shenzhen Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019 Ren, Wang, Jiang, Guo & Li 2017 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Genus Oreocryptophis (Red Mountain Rat Snake)   Oreocryptophis porphyraceus Red Mountain Rat Snake Li, Xiao, Qing, Lu & Lazell 2009 Schulz & Helfenberger 1998 Schulz 1996 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993 Genus Pseudagkistrodon (False Habu)   Pseudagkistrodon rudis False Habu Xu 2007 Zhao & Adler 1993 Genus Pseudoxenodon (Mock Cobras)   Pseudoxenodon bambusicola Bamboo Mock Cobra Xu 2007 Zhao & Adler 1993   Pseudoxenodon karlschmidti Common Chinese Mock Cobra Xu 2007 Zhao & Adler 1993   Pseudoxenodon macrops Common Large-eyed Mock Cobra Xu 2007 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas dhumnades Black-striped Rat Snake Xu 2007 Zhao & Adler 1993   Ptyas korros Indochinese Rat Snake Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993   Ptyas major Greater Green Snake Li, Xiao, Qing, Lu & Lazell 2009 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993   Ptyas mucosa Common Oriental Rat Snake Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993   Ptyas multicincta Many-banded Green Snake Hauser 2018 Xiao, Li & Qing 2010 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis callichroma Bavi Keelback Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993   * Rhabdophis guangdongensis Guangdong Keelback Zhu, Wang, Takeuchi & Zhao 2014   Rhabdophis helleri Northern Red-necked Keelback David & Vogel 2021 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Li, Xiao, Qing, Lu & Lazell 2009 [as subminiatus] Xie, Liu, Qiu, Zheng, Nie & Ma 1998 [as subminiatus] Xu 2007 [as subminiatus] Zhao & Adler 1993 Zhu, Wang, Takeuchi & Zhao 2014 [as subminiatus]   Rhabdophis nigrocinctus Black-banded Keelback Xu 2007 Genus Sibynophis (Oriental Black-headed Snakes)   Sibynophis chinensis Chinese Black-headed Snake Li, Xiao, Qing, Lu & Lazell 2009 Xu 2007 Zhao & Adler 1993 Genus Trimerodytes (Asian Water Snakes)   Trimerodytes aequifasciatus Equal-banded Water Snake Guo, Zhu, Liu, Wang, Che & Nguyen 2020 Li, Xiao, Qing, Lu & Lazell 2009 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993   Trimerodytes annularis Asian Banded Water Snake Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993   Trimerodytes balteatus Banded Mountain Stream Keelback Guo, Zhu, Liu, Wang, Che & Nguyen 2020 Li, Xiao, Qing, Lu & Lazell 2009 Ren, Wang, Guo, Wang, Nguyen & Li 2019 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Zhao & Adler 1993   Trimerodytes percarinatus Chinese Keelback Guo, Zhu, Liu, Wang, Che & Nguyen 2020 Li, Xiao, Qing, Lu & Lazell 2009 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Bungarus (Kraits)   Bungarus candidus Malayan Krait Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020   Bungarus fasciatus Banded Krait Li, Xiao, Qing, Lu & Lazell 2009 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993   Bungarus multicinctus Many-banded Krait Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993 Genus Naja (Typical Cobras)   Naja atra Chinese Spitting Cobra Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Shi, Vogel, Ding, Rao, Liu, Zhang, Wu & Chen 2022 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993 Genus Sinomicrurus (East Asian Coral Snakes)   Sinomicrurus annularis Hong Kong Coral Snake Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021   Sinomicrurus kelloggi Indochinese Coral Snake Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021 Xu 2007  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis atriceps Black-headed Sea Snake Zhao & Adler 1993 [as H. fasciatus]   Hydrophis caerulescens Bluish Slender-necked Sea Snake Zhao & Adler 1993   Hydrophis curtus Short Sea Snake Zhao & Adler 1993   Hydrophis cyanocinctus Blue-banded Sea Snake Zhao & Adler 1993   Hydrophis melanocephalus Black-headed Slender-necked Sea Snake Zhao & Adler 1993   Hydrophis ornatus Reef Sea Snake Zhao & Adler 1993   Hydrophis peronii Horned Sea Snake Wallach, Williams & Boundy 2014 Zhao & Adler 1993   Hydrophis platurus Yellow-bellied Sea Snake Zhao & Adler 1993   Hydrophis viperinus Viperine Sea Snake Zhao & Adler 1993 Genus Microcephalophis (Small-headed Sea Snakes)   Microcephalophis gracilis Graceful Small-headed Sea Snake Zhao & Adler 1993  Family Homalopsidae (Australasian Mud Snakes) Genus Hypsiscopus (Orange-bellied Mud Snakes and Matanna Mud Snake)   Hypsiscopus murphyi Northern Orange-bellied Mud Snake Gyi 1970 [as plumbea] Li, Xiao, Qing, Lu & Lazell 2009 [as plumbea] Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 [as plumbea] Xie, Liu, Qiu, Zheng, Nie & Ma 1998 [as plumbea] Xu 2007 [as plumbea] Zhao & Adler 1993 [as plumbea] Genus Myrrophis (Indochinese Mud Snakes)   Myrrophis bennetti Mangrove Mud Snake Zhao & Adler 1993   Myrrophis chinensis Chinese Mud Snake Gyi 1970 Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xu 2007 Zhao & Adler 1993  Family Pareidae (Oriental Slug Eaters) Genus Pareas (Typical Oriental Slug Eaters)   Pareas boulengeri Guizhou Slug Eater Zhao & Adler 1993   Pareas chinensis Chinese Slug Eater Zhao & Adler 1993   Pareas formosensis Taiwanese Red-eyed Slug Eater Ding, Chen, Suwannapoom, Nguyen, Pogarkov & Vogel 2020 Le, Tran, Hoang & Stuart 2021   Pareas macularius Black-spotted Slug Eater Xu 2007 Zhao & Adler 1993   Pareas margaritophorus White-spotted Slug Eater Deuve 1970 Li, Xiao, Qing, Lu & Lazell 2009 Vogel, Nguyen, Lalremsanga, Biakzuala, Hrima & Poyarkov 2020   Pareas monticola Assam Slug Eater Deuve 1970  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python bivittatus Burmese Python Barker & Barker 2010 Li, Xiao, Qing, Lu & Lazell 2009 [as P. molurus] Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 [as molurus] Zhao & Adler 1993  Family Typhlopidae (Typical Blind Snakes) Genus Virgotyphlops (Brahminy Blind Snake)   Virgotyphlops braminus Brahminy Blind Snake Lazell, Kolby, Lin, Zhuang & Lu 1999 Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Xu 2007 Zhao & Adler 1993  Family Viperidae (Vipers) Genus Azemiops (Asian Mole Vipers)   Azemiops kharini White-headed Mole Viper Zhang, Xiao, Yu & Luo 2012 [as feae] Genus Daboia (Daboias)   Daboia siamensis Siamese Chain Viper Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Zhao & Adler 1993 Genus Deinagkistrodon (Sharp-nosed Pitviper)   Deinagkistrodon acutus Sharp-nosed Pitviper Gloyd & Conant 1990 Xu 2007 Zhao & Adler 1993 Genus Gloydius (Mamushis)   Gloydius brevicaudus Short-tailed Mamushi Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 Watkins-Colwell & Leenders 2002 [as G. blomhofii brevicaudus] Genus Ovophis (Asian Mountain Pitvipers)   Ovophis makazayazaya Eastern Mountain Pitviper Xu 2007 [as monticola] Zhao & Adler 1993 [as monticola]   Ovophis tonkinensis Tonkin Pitviper Malhotra, Dawson, Guo & Thorpe 2011 Zhang, Wei, Jiang, Yu, Tang, Dang & Hu 2011 Genus Protobothrops (Habus and allies)   Protobothrops cornutus Horned Tree Pitviper David, Tong, Vogel & Tian 2008 Gong, Wang, Yang, Yu, Pan & Wang 2010 Vogel 2006 [as shenlii]   Protobothrops mucrosquamatus Brown-spotted Pitviper Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Hoge & Romano Hoge 1981 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Zhao & Adler 1993 Genus Trimeresurus (Oriental Pitvipers)   Trimeresurus albolabris Southeast Asian White-lipped Tree Pitviper Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Lazell, Kolby, Lin, Zhuang & Lu 1999 [as "cf. albolabris"] Li, Xiao, Qing, Lu & Lazell 2009 Qing, Xiao, Watkins-Colwell, Hou, Lu, Lazell & Sun 2015 [as "cf. albolabris"] Regenass & Kramer 1981 Xu 2007 Zhao & Adler 1993 Genus Viridovipera (Green Tree Pitvipers)   Viridovipera stejnegeri Chinese Green Tree Pitviper David, Vidal & Pauwels 2001 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Li, Xiao, Qing, Lu & Lazell 2009 Vogel 2006 Xie, Liu, Qiu, Zheng, Nie & Ma 1998 Xu 2007 Zhao & Adler 1993 Genus Zhaoermia (Mangshan Pitviper)   Zhaoermia mangshanensis Mangshan Pitviper Gong, Yang, Chen, Lau & Wang 2013 Guo, Liu, Li, Cao & Wang 2016  Family Xenodermidae (Asian Litter Snakes) Genus Achalinus (East Asian Burrowing Snakes)   Achalinus rufescens Rufous Burrowing Snake Li, Yu, Wu, Liao, Tang, Liu & Guo 2020 Wallach, Williams & Boundy 2014 Zhao & Adler 1993   Achalinus yunkaiensis Yunkai Mountains Burrowing Snake Li, Yu, Wu, Liao, Tang, Liu & Guo 2020 Wang, Li, Zeng, Lyu, Sung, Li, Lin & Wang 2019  Family Xenopeltidae (Asian Sunbeam Snakes) Genus Xenopeltis (Asian Sunbeam Snakes)   Xenopeltis hainanensis Hainan Sunbeam Snake Orlov 2000 Zhao 1997   Xenopeltis unicolor Common Asian Sunbeam Snake McDiarmid, Campbell & Toure 1999 Zhao & Adler 1993

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Huang in Bjorndal (ed.) 1982 Zhao & Adler 1993 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Huang in Bjorndal (ed.) 1982 Zhao & Adler 1993 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Huang in Bjorndal (ed.) 1982 Zhao & Adler 1993 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Zhao & Adler 1993  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Huang in Bjorndal (ed.) 1982 Zhao & Adler 1993  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora galbinifrons Northern Indochinese Box Turtle McCord & Joseph-Ouni 2002   Cuora trifasciata Northern Three-striped Box Turtle Fritz & Havas 2006 Iverson 1992 Lau & Shi 2000 Li, Xiao, Qing, Lu & Lazell 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Xu 2007 Zhao & Adler 1993 Genus Geoemyda (Black-breasted Leaf Turtles)   Geoemyda spengleri Common Black-breasted Leaf Turtle Dawson, Gaillard, Gong, Lin, McCormack, Nanthavong, Nguyen, Nguyen, Pham & Shi 2022 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Xu 2007 Yasukawa & Ota 2010 Zhao & Adler 1993 Genus Mauremys (Stripe-necked Turtles)   Mauremys mutica Yellow Pond Turtle Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Xu 2007 Zhao & Adler 1993   Mauremys nigricans Guangdong River Turtle Lau & Shi 2000 Li, Xiao, Qing, Lu & Lazell 2009 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Xu 2007 Zhao & Adler 1993   Mauremys reevesii Chinese Three-keeled Pond Turtle Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Xu 2007 Zhao & Adler 1993   Mauremys sinensis Chinese Stripe-necked Turtle Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Xu 2007 Zhao & Adler 1993 Genus Sacalia (Eastern Eyed Turtles)   Sacalia bealei Eastern Chinese Eyed Turtle Iverson 1992 Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Xu 2007 Zhao & Adler 1993   Sacalia quadriocellata Four-eyed Turtle Iverson 1992 Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Xu 2007 Zhao & Adler 1993  Family Platysternidae (Big-headed Turtle) Genus Platysternon (Big-headed Turtle)   Platysternon megacephalum Big-headed Turtle Lau & Shi 2000 Li, Xiao, Qing, Lu & Lazell 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Xu 2007 Zhao & Adler 1993  Family Trionychidae (Softshell Turtles) Genus Palea (Wattle-necked Softshell Turtle)   Palea steindachneri Wattle-necked Softshell Turtle Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Xu 2007 Zhao & Adler 1993 Genus Pelochelys (Indomalayan Giant Softshell Turtles)   Pelochelys cantorii Asian Giant Softshell Turtle Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Xu 2007 Zhao & Adler 1993 [as bibroni] Genus Pelodiscus (East Asian Softshell Turtles)   Pelodiscus axenaria Hunan Softshell Turtle Gong, Fritz, Vamberger, Gao & Farkas 2022 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelodiscus sinensis Chinese Softshell Turtle Li, Xiao, Qing, Lu & Lazell 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Xu 2007 Zhao & Adler 1993