Reptiles of Guangxi [China] Related checklists:  China Bibliography of the reptiles of Guangxi Taxonomic history of the reptiles of Guangxi Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthosaura (Pricklenapes)   Acanthosaura lepidogaster Brown Pricklenape Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993 Genus Calotes (South Asian Forest Agamas)   Calotes emma Forest Crested Agama Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021   Calotes versicolor South Indian Crested Agama Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993 Genus Diploderma (Eastern Mountain Agamas)   Diploderma fasciatum Banded Mountain Agama Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 [as Diploderma szechwanensis] Liu, Hou, Ananjeva & Rao 2023 Wang, Qi, Wang, Koehler, Lu, Lyu, Wang, Wang & Che 2022 Wang, Ren, Wu, Jiang, Jin, Hou, Zheng, Xie, Siler & Che 2021 Zhao & Adler 1993 [as Japalura szechwanensis] Genus Draco (Gliding Dragons)   Draco maculatus Black-spotted Gliding Dragon Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993 Genus Leiolepis (Butterfly Agamas)   Leiolepis reevesii Eastern Butterfly Agama Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993 Genus Physignathus (Indochinese Water Dragon)   Physignathus cocincinus Indochinese Water Dragon Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993 Genus Pseudocalotes (Long-headed Agamas)   Pseudocalotes brevipes Short-footed Long-headed Agama Ananjeva, Guo & Wang 2011 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993   Pseudocalotes microlepis Small-scaled Long-headed Agama Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021  Family Anguidae (Slowworms, etc.) Genus Dopasia (Oriental Glass Lizards)   Dopasia gracilis Burmese Glass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993   Dopasia harti South Chinese Glass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993  Family Dibamidae (Blind Skinks) Genus Dibamus (Oriental Blind Skinks)   Dibamus bourreti Indochinese Blind Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993  Family Eublepharidae (Eyelid Geckos) Genus Goniurosaurus (East Asian Leopard Geckos)   Goniurosaurus araneus Vietnamese Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chen, Meng, Jiang, Li, Wen, Lu, Lazell & Hou 2014 Zhu, Chen, Roman-Palacios, Li & He 2020   * Goniurosaurus chengzheng Chengzheng Leopard Gecko Zhu, Liu, Bai, Roman-Palacios, Li & He 2021   * Goniurosaurus gezhi Gezhi Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhu, Chen, Roman-Palacios, Li & He 2020   * Goniurosaurus kwangsiensis Guangxi Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Yang & Chan 2015 Zhu, Chen, Roman-Palacios, Li & He 2020   Goniurosaurus liboensis Libo Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021   Goniurosaurus lichtenfelderi Norway Islands Leopard Gecko Zhu, Chen, Roman-Palacios, Li & He 2020 Zhu, Shen, Liu, Chen, Li & He 2020   Goniurosaurus luii Pingxiang Leopard Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Grismer, Viets & Boyle 1999 Seufer, Kaverkin & Kirschner (eds.) 2005 Yang & Chan 2015 [as kadoorieorum] Zhu, Chen, Roman-Palacios, Li & He 2020 [as kadoorieorum]  Family Gekkonidae (Typical Geckos) Genus Gekko (Tokays)   Gekko adleri Cao Bang Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lyu, Lin, Ren, Jiang, Zhang, Qi & Wang 2021 Nguyen, Wang, Yang, Lehmann, Le, Ziegler & Bonkowski 2013   Gekko chinensis Chinese Narrow-disked Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Zhao & Adler 1993   Gekko gecko Tokay Lyu, Lin, Ren, Jiang, Zhang, Qi & Wang 2021 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Xu, Li, Zhang, Zhao, Miao & Tang 2009 Zhao & Adler 1993   Gekko japonicus Japanese Giant Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lyu, Lin, Ren, Jiang, Zhang, Qi & Wang 2021 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Zhao & Adler 1993   * Gekko kwangsiensis Wuming Rock Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lyu, Lin, Ren, Jiang, Zhang, Qi & Wang 2021 Yang 2015   Gekko liboensis Guizhou Narrow-disked Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021   Gekko palmatus Tonkin Palmated Gecko Lyu, Lin, Ren, Jiang, Zhang, Qi & Wang 2021   Gekko reevesii Chinese Black Tokay Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus bowringii Hong Kong House Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993   Hemidactylus frenatus Asian House Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhou & al. 2003   Hemidactylus garnotii Fox Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Jiang, Zhou, Lu, Han & Li 2006 Genus Hemiphyllodactylus (Slender Geckos)   Hemiphyllodactylus jinpingensis Jinping Slender Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhou, Liu & Yang 1981  Family Lacertidae (Typical Lizards) Genus Takydromus (Oriental Grass Lizards)   Takydromus intermedius Sichuan Grass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Wang, Lyu, Yang, Li & Wang 2019 Zhao & Adler 1993   Takydromus kuehnei Kanshirei Grass Lizard Bischoff 1992 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993   Takydromus septentrionalis Northern Grass Lizard Zhao & Adler 1993   Takydromus sexlineatus Six-lined Grass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Wang, Lyu, Yang, Li & Wang 2019 Zhao & Adler 1993  Family Scincidae (Skinks) Genus Ateuchosaurus (East Asian Short-legged Skinks)   Ateuchosaurus chinensis Chinese Short-Legged Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993 Genus Eutropis (South Asian Sun Skinks)   Eutropis longicaudata Long-tailed Sun Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021   Eutropis multifasciata Many-lined Sun Skink David, Pauwels, Lays & Lenglet 2006 Genus Plestiodon (Blue-tailed Skinks)   Plestiodon chinensis Eastern Chinese Long Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993   Plestiodon elegans Elegant Five-lined Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993   Plestiodon quadrilineatus Four-striped Long Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993 Genus Scincella (South Asian and American Ground Skinks)   Scincella doriae Burmese Ground Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021   Scincella modesta Slender Ground Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021   Scincella reevesii Speckled Ground Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus cryptotis Depressed-eared Forest Skink Qi, Lyu, Xie, Wan, Zhang, Liu, Mo & Wang 2022   Sphenomorphus incognitus Koshun Forest Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993   Sphenomorphus indicus Common Indian Forest Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993   Sphenomorphus tonkinensis Tonkin Forest Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Wang, Yang & Liu 2013 Genus Tropidophorus (Stream Skinks)   Tropidophorus guangxiensis Guangxi Stream Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Guo, Shu, Wu, Chen, Hou, Shi & Deng 2021 Wen 1992 Zhao & Adler 1993   Tropidophorus hainanus Hainan Stream Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Guo, Shu, Wu, Chen, Hou, Shi & Deng 2021 Zhao & Adler 1993   Tropidophorus sinicus South Chinese Stream Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Guo, Shu, Wu, Chen, Hou, Shi & Deng 2021 Zhao & Adler 1993  Family Shinisauridae (Chinese Crocodile Lizard) Genus Shinisaurus (Crocodile Lizard)   Shinisaurus crocodilurus Crocodile Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lu 2003 Schingen, Ihlow, Nguyen, Ziegler, Bonkowski, Wu & Roedder 2014 Zhao & Adler 1993  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus salvator Asian Water Monitor Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Zhou & Yu 2002 Genus Amphiesma (South Asian Keelbacks)   Amphiesma stolatum Buff-striped Keelback Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Zhou & Yu 2002 Genus Amphiesmoides (White-eyed Keelback)   Amphiesmoides ornaticeps White-eyed Keelback Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Genus Archelaphe (Venus Rat Snake)   Archelaphe bella Venus Rat Snake Jiang, Zhou, Liu & Wang 2006 Schulz, Boehme & Tillack 2011 Wallach, Williams & Boundy 2014 Genus Boiga (Australasian Cat Snakes)   Boiga guangxiensis Guangxi Cat Snake Orlov & Ryabov 2002 Wen 1998 Zhou & Yu 2002   Boiga kraepelini Kelung Cat Snake Ananjeva, Golynsky, Lin, Orlov & Tseng 2015 Orlov & Ryabov 2002 Zhao & Adler 1993   Boiga multomaculata Marbled Cat Snake Orlov & Ryabov 2002 Zhao & Adler 1993 Zhou & Yu 2002 Genus Calamaria (Oriental Reed Snakes)   Calamaria pavimentata Brown Reed Snake Zhao & Adler 1993   Calamaria septentrionalis Northern Reed Snake Inger & Marx 1965 Zhao & Adler 1993 Zhou & Yu 2002 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea ornata Ornate Flying Tree Snake Deuve 1970 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus radiatus Copper-headed Rat Snake Schulz 1996 Zhao & Adler 1993 Zhou & Yu 2002 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis proarchos Northern Painted Bronzeback Zhao & Adler 1993 [as pictus] Zhou & Yu 2002 [as pictus] Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe carinata East Asian Keeled Rat Snake Schulz 1996 Zhao & Adler 1993   Elaphe moellendorffi Red-headed Rat Snake Qi, Shi, Ma, Gao, Bu, Grismer, Li & Wang 2021 Schulz 1996 Zhao & Adler 1993 Zhou & Yu 2002   Elaphe taeniura Stripe-tailed Rat Snake Schulz 1996 Zhao & Adler 1993 Zhou & Yu 2002 Genus Euprepiophis (East Asian Rat Snakes)   Euprepiophis mandarinus Mandarin Rat Snake Schulz 1996 Zhou & Yu 2002 Genus Fowlea (Fishing Keelbacks)   Fowlea flavipunctata Yellow-spotted Keelback Vogel & David 2012 Zhao & Adler 1993 [as piscator] Zhou & Yu 2002 [as piscator]   Fowlea piscator Checkered Keelback Vogel & David 2006 Vogel & David 2012 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma boulengeri Vietnamese Rhinoceros Snake Zhao & Adler 1993 Zhou & Yu 2002   Gonyosoma coeruleum Blue-eyed Bush Rat Snake David, Campbell, Deuti, Hauser, Luu, Nguyen, Orlov, Pauwels, Scheinberg, Sethy, Smits, Teynie & Vogel 2022   Gonyosoma frenatum Khasi Hills Rat Snake Schulz 1996 Zhou & Yu 2002 Genus Hebius (Oriental Equatorial Keelbacks)   Hebius atemporalis Tonkin Keelback Wallach, Williams & Boundy 2014 Zhao & Adler 1993   Hebius bitaeniatus Two-striped Keelback David, Agarwal, Athreya, Mathew, Vogel & Mistry 2015 David, Bain, Nguyen, Orlov, Vogel, Vu & Ziegler 2007 David, Vogel & Pauwels 2005 Lalremsanga, Bal, Vogel & Biakzuala 2022 Purkayastha & David 2019 Wallach, Williams & Boundy 2014   Hebius boulengeri White-browed Keelback David, Bain, Nguyen, Orlov, Vogel, Vu & Ziegler 2007 [as "cf. boulengeri"] David, Vogel & Rooijen 2013 Purkayastha & David 2019 Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Zhou & Yu 2002   Hebius craspedogaster Kuatun Keelback Wallach, Williams & Boundy 2014 Zhao & Adler 1993   Hebius khasiensis Khasi Hills Keelback Wallach, Williams & Boundy 2014   Hebius octolineatus Eight-striped Keelback Mo, Zhou, Xie & Zhou 2002 Wallach, Williams & Boundy 2014   Hebius optatus Mount Omei Keelback Guo, Liu, Li, Zhong, Chen & Wang 2012 Lalremsanga, Bal, Vogel & Biakzuala 2022 Wallach, Williams & Boundy 2014 Zhao & Adler 1993   Hebius popei Chinese Mountain Keelback David, Bain, Nguyen, Orlov, Vogel, Vu & Ziegler 2007 David, Vogel & Rooijen 2013 Malnate 1962 Purkayastha & David 2019 Wallach, Williams & Boundy 2014 Zhao & Adler 1993   Hebius sauteri Kosempo Keelback Malnate 1962 Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Zhou & Yu 2002 Genus Lycodon (Oriental Wolf Snakes)   * Lycodon cathaya Huaping Wolf Snake Wang, Qi, Lyu, Zeng & Wang 2020   Lycodon fasciatus Yellow-banded Wolf Snake Zhao & Adler 1993 Zhou & Yu 2002   Lycodon flavozonatus Yellow Big-toothed Snake Zhao & Adler 1993 Zhou & Yu 2002   Lycodon laoensis Laotian Wolf Snake Zhou & Yu 2002   Lycodon meridionalis Vietnamese Big-toothed Snake Orlov & Ryabov 2004 Wang, Qi, Lyu, Zeng & Wang 2020   Lycodon pictus Painted Wolf Snake Janssen, Ren, Li, Zeng, Nguyen, Nguyen, Bui, Ngo, Le & Ziegler 2020   Lycodon rufozonatus Red-banded Big-toothed Snake Zhao & Adler 1993   Lycodon ruhstrati Taiwanese Wolf Snake Wang, Qi, Lyu, Zeng & Wang 2020 Zhao & Adler 1993 Zhou & Yu 2002   Lycodon subcinctus Malayan Banded Wolf Snake Zhou & Yu 2002 Genus Oligodon (Kukri Snakes)   Oligodon chinensis Chinese Kukri Snake Qian, Qi, Shi, Lu, Jenkins, Mo & Li 2021 Zhao & Adler 1993 Zhou & Yu 2002   Oligodon cinereus Black-barred Kukri Snake Zhao & Adler 1993 Zhou & Yu 2002   Oligodon eberhardti Tam Dao Kukri Snake Zhao & Adler 1993   Oligodon formosanus Taiwanese Kukri Snake David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012 Zhao & Adler 1993 Zhou & Yu 2002   Oligodon nagao Lang Son Kukri Snake David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012   Oligodon ornatus Chinese Ornate Kukri Snake Qian, Qi, Shi, Lu, Jenkins, Mo & Li 2021 Genus Oocatochus (Red-backed Rat Snake)   Oocatochus rufodorsatus Red-backed Rat Snake Schulz 1996 Zhou & Yu 2002 Genus Opisthotropis (Mountain Stream Keelbacks)   Opisthotropis cheni Guangdong Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019   Opisthotropis guangxiensis Guangxi Mountain Stream Keelback David, Pauwels, Nguyen & Vogel 2015 Guo, Liu, Li, Zhong, Chen & Wang 2012 Ren, Wang, Guo, Wang, Nguyen & Li 2019 Ren, Wang, Jiang, Guo & Li 2017 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Zhao & Adler 1993   Opisthotropis haihaensis Hai Ha Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019 [as O. maculosa (part)] Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 [as O. maculosa (part)] Yang, Wang, Zhang, Lau & Chou 2011 [as O. maculosa (part)]   Opisthotropis hungtai Heishiding Mountain Stream Keelback Wang, Lyu, Zeng, Lin, Yang, Nguyen, Le, Ziegler & Wang 2020 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 [as O. maculosa [part]] Yang, Wang, Zhang, Lau & Chou 2011 [as O. maculosa (part)]   Opisthotropis kuatunensis Striped Mountain Stream Keelback Zhao & Adler 1993   Opisthotropis lateralis Bicoloured Mountain Stream Keelback David, Pauwels, Nguyen & Vogel 2015 Ren, Wang, Guo, Wang, Nguyen & Li 2019 Ren, Wang, Jiang, Guo & Li 2017 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Zhao & Adler 1993   Opisthotropis latouchii South Chinese Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Zhao & Adler 1993 Zhou & Yu 2002   Opisthotropis maxwelli Fujian Mountain Stream Keelback Zhao & Adler 1993 Genus Oreocryptophis (Red Mountain Rat Snake)   Oreocryptophis porphyraceus Red Mountain Rat Snake Schulz & Helfenberger 1998 Schulz 1996 Zhao & Adler 1993 Zhou & Yu 2002 Genus Plagiopholis (Blotch-necked Snakes)   Plagiopholis styani Fujian Blotch-necked Snake Zhao & Adler 1993 Genus Pseudagkistrodon (False Habu)   Pseudagkistrodon rudis False Habu Zhao & Adler 1993 Genus Pseudoxenodon (Mock Cobras)   Pseudoxenodon bambusicola Bamboo Mock Cobra Zhao & Adler 1993 Zhou & Yu 2002   Pseudoxenodon karlschmidti Common Chinese Mock Cobra Zhao & Adler 1993   Pseudoxenodon macrops Common Large-eyed Mock Cobra Zhao & Adler 1993 Zhou & Yu 2002   Pseudoxenodon stejnegeri Chinese Large-eyed Mock Cobra Zhao & Adler 1993 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas dhumnades Black-striped Rat Snake Zhao & Adler 1993   Ptyas korros Indochinese Rat Snake Zhao & Adler 1993 Zhou & Yu 2002   Ptyas major Greater Green Snake Zhao & Adler 1993 Zhou & Yu 2002   Ptyas mucosa Common Oriental Rat Snake Zhao & Adler 1993 Zhou & Yu 2002   Ptyas multicincta Many-banded Green Snake Hauser 2018 Zhao & Adler 1993 Zhou & Yu 2002 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis helleri Northern Red-necked Keelback Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Zhao & Adler 1993 Zhou & Yu 2002 [as subminiatus]   Rhabdophis lateralis Chinese Tiger Keelback Zhao & Adler 1993 Zhu, Wang, Takeuchi & Zhao 2014 [as R. tigrinus]   Rhabdophis leonardi Burmese Mountain Keelback Zhao & Adler 1993   Rhabdophis nuchalis Collared Keelback Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Genus Sibynophis (Oriental Black-headed Snakes)   Sibynophis chinensis Chinese Black-headed Snake Zhao & Adler 1993 Zhou & Yu 2002 Genus Trimerodytes (Asian Water Snakes)   Trimerodytes aequifasciatus Equal-banded Water Snake Guo, Zhu, Liu, Wang, Che & Nguyen 2020 Zhao & Adler 1993 Zhou & Yu 2002   Trimerodytes annularis Asian Banded Water Snake Ren, Wang, Guo, Wang, Nguyen & Li 2019 Zhao & Adler 1993   Trimerodytes balteatus Banded Mountain Stream Keelback Guo, Zhu, Liu, Wang, Che & Nguyen 2020 Ren, Wang, Guo, Wang, Nguyen & Li 2019 Zhao & Adler 1993   Trimerodytes percarinatus Chinese Keelback Guo, Zhu, Liu, Wang, Che & Nguyen 2020 Zhao & Adler 1993 Zhou & Yu 2002  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Bungarus (Kraits)   Bungarus candidus Malayan Krait Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020   Bungarus fasciatus Banded Krait Zhao & Adler 1993 Zhou & Yu 2002   Bungarus multicinctus Many-banded Krait Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Zhao & Adler 1993 Zhou & Yu 2002 Genus Naja (Typical Cobras)   Naja atra Chinese Spitting Cobra Pope 1935 Zhao & Adler 1993 [as N. naja] Zhou & Yu 2002   Naja fuxi Brown Banded Cobra Shi, Vogel, Chen & Ding in Shi, Vogel, Ding, Rao, Liu, Zhang, Wu & Chen 2022 Wuster 1998 [as N. kaouthia] Zhao & Adler 1993 [as N. kaouthia] Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Zhao & Adler 1993 Zhou & Yu 2002 Genus Sinomicrurus (East Asian Coral Snakes)   Sinomicrurus kelloggi Indochinese Coral Snake Nguyen & Poyarkov 2020 [as houi] Nguyen, Nanthavong, Coudrat, Brakels & Poyarkov 2020 [as houi] Zhao & Adler 1993   Sinomicrurus peinani Guangxi Coral Snake Liu, Yan, Hou, Wang, Nguyen, Murphy, Che & Guo 2020 Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis atriceps Black-headed Sea Snake Zhao & Adler 1993 [as H. fasciatus] Zhou & Yu 2002 [as H. fasciatus]   Hydrophis curtus Short Sea Snake Zhao & Adler 1993 Zhou & Yu 2002   Hydrophis cyanocinctus Blue-banded Sea Snake Zhao & Adler 1993 Zhou & Yu 2002   Hydrophis melanocephalus Black-headed Slender-necked Sea Snake Zhou & Yu 2002   Hydrophis ornatus Reef Sea Snake Zhao & Adler 1993 Zhou & Yu 2002   Hydrophis platurus Yellow-bellied Sea Snake Zhou & Yu 2002   Hydrophis viperinus Viperine Sea Snake Zhao & Adler 1993 Zhou & Yu 2002 Genus Microcephalophis (Small-headed Sea Snakes)   Microcephalophis gracilis Graceful Small-headed Sea Snake Zhao & Adler 1993 Zhou & Yu 2002  Family Homalopsidae (Australasian Mud Snakes) Genus Hypsiscopus (Orange-bellied Mud Snakes and Matanna Mud Snake)   Hypsiscopus murphyi Northern Orange-bellied Mud Snake Zhao & Adler 1993 [as plumbea] Zhou & Yu 2002 [as plumbea] Genus Myrrophis (Indochinese Mud Snakes)   Myrrophis bennetti Mangrove Mud Snake Zhou & Yu 2002   Myrrophis chinensis Chinese Mud Snake Zhao & Adler 1993 Zhou & Yu 2002  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Zhao & Adler 1993 Zhou & Yu 2002  Family Pareidae (Oriental Slug Eaters) Genus Pareas (Typical Oriental Slug Eaters)   Pareas boulengeri Guizhou Slug Eater Zhao & Adler 1993   Pareas chinensis Chinese Slug Eater Zhao & Adler 1993   Pareas formosensis Taiwanese Red-eyed Slug Eater Le, Tran, Hoang & Stuart 2021   Pareas macularius Black-spotted Slug Eater Vogel, Nguyen, Lalremsanga, Biakzuala, Hrima & Poyarkov 2020 Zhao & Adler 1993 Zhou & Yu 2002   Pareas margaritophorus White-spotted Slug Eater Deuve 1970 Vogel, Nguyen, Lalremsanga, Biakzuala, Hrima & Poyarkov 2020 Zhou & Yu 2002   Pareas monticola Assam Slug Eater Deuve 1970   Pareas stanleyi Fujian Slug Eater Ding, Chen, Suwannapoom, Nguyen, Pogarkov & Vogel 2020 Vogel, Nguyen, Lalremsanga, Biakzuala, Hrima & Poyarkov 2020 Yang, Yeung, Huang & Yang 2021  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python bivittatus Burmese Python Barker & Barker 2010 Zhao & Adler 1993 Zhou & Yu 2002  Family Typhlopidae (Typical Blind Snakes) Genus Virgotyphlops (Brahminy Blind Snake)   Virgotyphlops braminus Brahminy Blind Snake Zhao & Adler 1993 Zhou & Yu 2002  Family Viperidae (Vipers) Genus Azemiops (Asian Mole Vipers)   Azemiops kharini White-headed Mole Viper Orlov, Ryabov & Nguyen 2013 Zhao & Adler 1993 [as feae] Zhou & Yu 2002 [as feae] Genus Daboia (Daboias)   Daboia siamensis Siamese Chain Viper Zhao & Adler 1993 Zhou & Yu 2002 [as russellii] Genus Deinagkistrodon (Sharp-nosed Pitviper)   Deinagkistrodon acutus Sharp-nosed Pitviper Gloyd & Conant 1990 Zhao & Adler 1993 Genus Ovophis (Asian Mountain Pitvipers)   Ovophis makazayazaya Eastern Mountain Pitviper Zhao & Adler 1993 [as monticola] Zhou & Yu 2002 [as monticola orientalis]   Ovophis tonkinensis Tonkin Pitviper Malhotra, Dawson, Guo & Thorpe 2011 Genus Protobothrops (Habus and allies)   Protobothrops jerdonii Spotted Mountain Pitviper Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Zhao & Adler 1993   Protobothrops maolanensis Maolan Pitviper Chen, Luo, Meng, Wen & Jiang 2013   Protobothrops mucrosquamatus Brown-spotted Pitviper Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Hoge & Romano Hoge 1981 Zhao & Adler 1993 Zhou & Yu 2002   Protobothrops trungkhanhensis Trung Khanh Pitviper Guo, Liu, Wen, Xiao, Fang, Zhong, Nguyen, Zhu, Jadin & Li 2016 Genus Trimeresurus (Oriental Pitvipers)   Trimeresurus albolabris Southeast Asian White-lipped Tree Pitviper Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Regenass & Kramer 1981 Zhao & Adler 1993 Zhou & Yu 2002 Genus Viridovipera (Green Tree Pitvipers)   Viridovipera stejnegeri Chinese Green Tree Pitviper David, Vidal & Pauwels 2001 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Zhao & Adler 1993 Zhou & Yu 2002  Family Xenodermidae (Asian Litter Snakes) Genus Achalinus (East Asian Burrowing Snakes)   Achalinus ater Chinese Black Burrowing Snake Li, Yu, Wu, Liao, Tang, Liu & Guo 2020 Wallach, Williams & Boundy 2014 Zhao & Adler 1993   Achalinus emilyae Quang Ninh Burrowing Snake Hou, Wang, Guo, Chen, Yuan & Che 2021   Achalinus rufescens Rufous Burrowing Snake Li, Yu, Wu, Liao, Tang, Liu & Guo 2020 Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Zhou & Yu 2002   Achalinus spinalis Grey Burrowing Snake Zhao & Adler 1993   Achalinus yunkaiensis Yunkai Mountains Burrowing Snake Li, Yu, Wu, Liao, Tang, Liu & Guo 2020  Family Xenopeltidae (Asian Sunbeam Snakes) Genus Xenopeltis (Asian Sunbeam Snakes)   Xenopeltis hainanensis Hainan Sunbeam Snake McDiarmid, Campbell & Toure 1999 Orlov 2000 Zhao & Adler 1993 Zhao 1997

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Huang in Bjorndal (ed.) 1982 Wen 1984 Zhao & Adler 1993 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Huang in Bjorndal (ed.) 1982 Wen 1984 Zhao & Adler 1993 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Huang in Bjorndal (ed.) 1982 Wen 1984 Zhao & Adler 1993 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Wen 1984 Zhao & Adler 1993  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Huang in Bjorndal (ed.) 1982 Zhao & Adler 1993  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora cyclornata Vietnamese Three-striped Box Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Cuora flavomarginata Chinese Yellow-margined Box Turtle Ota, Yasukawa, Fu & Chen 2009 Zhao & Adler 1993   Cuora galbinifrons Northern Indochinese Box Turtle Iverson 1992 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993   * Cuora mccordi Guangxi Box Turtle Bonin, Devaux & Dupre 2006 Iverson 1992 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993   Cuora mouhotii Keeled Box Turtle Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993   Cuora trifasciata Northern Three-striped Box Turtle Fritz & Havas 2006 Iverson 1992 Lau & Shi 2000 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993   Cuora zhoui Southern Chinese Box Turtle Iverson 1992 Lau & Shi 2000 Zhao & Adler 1993 Genus Geoemyda (Black-breasted Leaf Turtles)   Geoemyda spengleri Common Black-breasted Leaf Turtle Dawson, Gaillard, Gong, Lin, McCormack, Nanthavong, Nguyen, Nguyen, Pham & Shi 2022 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Yasukawa & Ota 2010 Zhao & Adler 1993 Genus Mauremys (Stripe-necked Turtles)   Mauremys mutica Yellow Pond Turtle Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993   Mauremys nigricans Guangdong River Turtle Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993   Mauremys reevesii Chinese Three-keeled Pond Turtle Lau & Shi 2000 McCord & Joseph-Ouni 2004 Zhao & Adler 1993 [as megalocephala]   Mauremys sinensis Chinese Stripe-necked Turtle Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 Genus Sacalia (Eastern Eyed Turtles)   Sacalia bealei Eastern Chinese Eyed Turtle Lau & Shi 2000 Zhao & Adler 1993   Sacalia quadriocellata Four-eyed Turtle Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993  Family Platysternidae (Big-headed Turtle) Genus Platysternon (Big-headed Turtle)   Platysternon megacephalum Big-headed Turtle Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993  Family Testudinidae (Tortoises) Genus Indotestudo (Asian Tortoises)   Indotestudo elongata Elongated Tortoise Ihlow, Dawson, Hartmann & Som 2016 Iverson 1992 Lau & Shi 2000 Zhao & Adler 1993  Family Trionychidae (Softshell Turtles) Genus Palea (Wattle-necked Softshell Turtle)   Palea steindachneri Wattle-necked Softshell Turtle Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 Genus Pelochelys (Indomalayan Giant Softshell Turtles)   Pelochelys cantorii Asian Giant Softshell Turtle Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 [as bibroni] Genus Pelodiscus (East Asian Softshell Turtles)   Pelodiscus axenaria Hunan Softshell Turtle Gong, Fritz, Vamberger, Gao & Farkas 2022 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Yang, Tang, Ding, Guo & Wang 2011   Pelodiscus parviformis Lesser Chinese Softshell Turtle Fritz & Havas 2006 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelodiscus sinensis Chinese Softshell Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993