Reptiles of
Sumatra [Indonesia]

Related checklists:  Indonesia

Bibliography of the reptiles of Sumatra Taxonomic history of the reptiles of Sumatra Introduced species
Home - Taxonomy - Geography - Biodiversity - Literature - Purchase RepFocus Recent updates
Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist
General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial.
Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:
Crocodilians (Crocodylia)
= endemicEX = extinct or assumed extinct

Species
References
 Family Crocodylidae (Crocodiles)
Genus Crocodylus (Typical Crocodiles)
 
Crocodylus porosus

Saltwater Crocodile

Ladiges 1939 Rooij 1915 Wermuth & Mertens 1977
 Family Gavialidae (Gharials)
Genus Tomistoma (Malayan Gharial)
 
Tomistoma schlegelii

Malayan Gharial

Bezuijen, Shwedick, Sommerlad, Stevenson & Stuebing 2010 Bezuijen, Webb, Hartoyo & Samedi 2001 Ladiges 1939 Rooij 1915 Stuebing, Bezuijen, Auliya & Voris 2006 Teynie, David & Ohler 2010 Wermuth & Mertens 1977

Lizards (Sauria)
= endemicEX = extinct or assumed extinct

Species
References
 Family Agamidae (Agamas and Dragons)
Genus Acanthosaura (Pricklenapes)
 
Acanthosaura armata

Peninsular Pricklenape

Cox, Dijk, Nabhitabhata & Thirakhupt 1998 Grismer 2011 Manthey 2008 Teynie, David & Ohler 2010
Genus Aphaniotis (Earless Agamas)
 
Aphaniotis acutirostris

Long-snouted Earless Agama

Loennberg & Rendahl 1925 Malkmus & Kunkel 1992 Manthey 2008 Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010 Wermuth 1967
 
Aphaniotis fusca

Blue-mouthed Earless Agama

Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Rooij 1915 Teynie, David & Ohler 2010 Wermuth 1967
Genus Bronchocela (Long-tailed Agamas)
 
Bronchocela cristatella

Malayan Green Crested Agama

Brongersma in Dammerman 1929 Hallermann 2005 Ladiges 1939 Loennberg & Rendahl 1925 Malkmus & Kunkel 1992 Manthey 2008 Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010
  *
Bronchocela hayeki

Sumatran Long-tailed Agama

Amarasinghe, Kamsi, Riyanto, Putra, Hallermann, Andayani, Abinawanto & Supriatna 2022 Hallermann 2005 Manthey 2008 Teynie, David & Ohler 2010 Wermuth 1967
 
Bronchocela jubata

Maned Long-tailed Agama

Amarasinghe, Ineich, Riyanto, Hallermann, Andayani, Abinawanto & Supriatna 2022 Hallermann 2005 Manthey 2008
Genus Dendragama (Sumatran Tree Agamas)
  *
Dendragama australis

Southern Sumatran Tree Agama

Harvey, Shaney, Sidik, Kurniawan & Smith 2017 Shaney, Harvey, Hamidy, Kurniawan & Smith 2020
  *
Dendragama boulengeri

Central Sumatran Tree Agama

Harvey, Shaney, Sidik, Kurniawan & Smith 2017 Malkmus & Kunkel 1992 Manthey 2008 Rooij 1915 Shaney, Harvey, Hamidy, Kurniawan & Smith 2020 Teynie, David & Ohler 2010 Wermuth 1967
  *
Dendragama dioidema

Northern Sumatran Tree Agama

Harvey, Shaney, Sidik, Kurniawan & Smith 2017 Shaney, Harvey, Hamidy, Kurniawan & Smith 2020
  *
Dendragama schneideri

Simbolon Tree Agama

Ahl 1926 Shaney, Harvey, Hamidy, Kurniawan & Smith 2020
Genus Draco (Gliding Dragons)
 
Draco abbreviatus

Orange-bearded Gliding Dragon

Brongersma in Dammerman 1929 [as D. fimbriatus] Grismer 2011 Inger 1983 [as D. fimbriatus] Loennberg & Rendahl 1925 [as D. fimbriatus] McGuire, Cotoras, O'Connell, Lawalata, Wang-Claypool, Stubbs, Huang, Wogan, Hykin, Reilly, Bi, Riyanto, Arida, Smith, Milne, Streicher & Iskandar 2018 [as D. fimbriatus] Teynie, David & Ohler 2010 [as D. fimbriatus]
 
Draco cornutus

Horned Gliding Dragon

Musters 1983 Teynie, David & Ohler 2010 Wermuth 1967
 
Draco fimbriatus

Fringed Gliding Dragon

McGuire, Cotoras, O'Connell, Lawalata, Wang-Claypool, Stubbs, Huang, Wogan, Hykin, Reilly, Bi, Riyanto, Arida, Smith, Milne, Streicher & Iskandar 2018
 
Draco haematopogon

Red-bearded Gliding Dragon

Brongersma in Dammerman 1929 Inger 1983 Musters 1983 Teynie, David & Ohler 2010 Wermuth 1967
 
Draco maximus

Giant Gliding Dragon

Musters 1983 Teynie, David & Ohler 2010 Wermuth 1967
 
Draco melanopogon

Black-bearded Gliding Dragon

Manthey 2008 Musters 1983 Teynie, David & Ohler 2010 Wermuth 1967
 
Draco quinquefasciatus

Five-banded Gliding Dragon

Inger 1983 Musters 1983 Teynie, David & Ohler 2010
 
Draco sumatranus

Sumatran Gliding Dragon

Malkmus & Kunkel 1992 McGuire & Kiew 2001 Musters 1983
Genus Gonocephalus (Western Anglehead Agamas)
  *
Gonocephalus beyschlagi

Northern Sumatra Anglehead Agama

Malkmus & Kunkel 1992 Manthey & Denzer 1992 Manthey 2010 Teynie, David & Ohler 2010 Wermuth 1967
 
Gonocephalus chamaeleontinus

Chameleon Anglehead Agama

Brongersma in Dammerman 1929 Ladiges 1939 Loennberg & Rendahl 1925 Manthey & Denzer 1993 Manthey 2010 Rooij 1915 Teynie, David & Ohler 2010 Wermuth 1967
 
Gonocephalus doriae

Sarawak Anglehead Agama

Harvey, Sarker, Sidik, Kurniawan & Smith 2023
 
Gonocephalus grandis

Greater Anglehead Agama

Cox, Dijk, Nabhitabhata & Thirakhupt 1998 Ladiges 1939 Malkmus & Kunkel 1992 Manthey & Denzer 1991 Manthey 2010 Rooij 1915 Teynie, David & Ohler 2010 Wermuth 1967
  *
Gonocephalus inauris

Bengkulu Anglehead Agama

Harvey, Sarker, Sidik, Kurniawan & Smith 2023
  *
Gonocephalus klossi

Western Sumatra Anglehead Agama

Harvey, Rech, Riyanto, Kurniawan & Smith 2021 Manthey & Denzer 1991 Manthey 2010 Teynie, David & Ohler 2010 Wermuth 1967
 
Gonocephalus kuhlii

Java Anglehead Agama

Manthey & Denzer 1993 Manthey 2010
  *
Gonocephalus lacunosus

Sikulikap Anglehead Agama

Harvey, Rech, Riyanto, Kurniawan & Smith 2021 Manthey & Denzer 1991 Manthey 2010 Teynie, David & Ohler 2010
 
Gonocephalus liogaster

Blue-eyed Anglehead Agama

Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Manthey & Denzer 1992 Manthey 2010 Rooij 1915 Teynie, David & Ohler 2010 Wermuth 1967
  *
Gonocephalus megalepis

Sumatran Large-scaled Anglehead Agama

Harvey, Rech, Riyanto, Kurniawan & Smith 2021 Manthey & Denzer 1991 Manthey 2010 Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010 Wermuth 1967
  *
Gonocephalus pyrius

Barisan Range Anglehead Agama

Harvey, Rech, Riyanto, Kurniawan & Smith 2021
Genus Harpesaurus (Sunda Horned Agamas)
  *
Harpesaurus beccarii

West Sumatran Horned Agama

Manthey 2010 Rooij 1915 Teynie, David & Ohler 2010 Wermuth 1967
  *
Harpesaurus modigliani

East Sumatran Horned Agama

Manthey 2010 Putra, Amarasinghe, Hikmatullah, Scali, Brinkman, Manthey & Ineich 2020 Teynie, David & Ohler 2010 Wermuth 1967
Genus Lophocalotes (Sumatra Crested Agamas)
  *
Lophocalotes achlios

White-throated Crested Agama

Harvey, Scrivani, Shaney, Hamidy, Kurniawan & Smith 2018
  *
Lophocalotes ludekingi

Northern Crested Agama

Manthey 2010 Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010 Wermuth 1967
Genus Phoxophrys (Sumatran Shrub Agama)
  *
Phoxophrys tuberculata

Sumatran Shrub Agama

Harvey, Larson, Jacobs, Shaney, Streicher, Hamidy, Kurniawan & Smith 2020 Manthey 2010 Rooij 1915 Teynie, David & Ohler 2010 Wermuth 1967
Genus Pseudocalotes (Long-headed Agamas)
  *
Pseudocalotes baliomus

Barisan Range Long-headed Agama

Harvey, Shaney, Hamidy, Kurniawan & Smith 2017
  *
Pseudocalotes cybelidermus

Purple-throated Long-headed Agama

Harvey, Hamidy, Wan, Shaney & Smith 2014
  *
Pseudocalotes guttalineatus

Dash-lined Long-headed Agama

Harvey, Hamidy, Wan, Shaney & Smith 2014
  *
Pseudocalotes rhammanotus

Stitch-backed Long-headed Agama

Harvey, Hamidy, Wan, Shaney & Smith 2014
 
Pseudocalotes tympanistriga

Javanese Long-headed Agama

Brongersma in Dammerman 1929 Dring 1979 Hallermann & Boehme 2000 Ladiges 1939 Manthey 2010 Rooij 1915 Teynie, David & Ohler 2010
Genus Thaumatorhynchus (Bengkulu Horned Agama)
  *
Thaumatorhynchus brooksi

Bengkulu Horned Agama

Manthey 2010 Teynie, David & Ohler 2010 Wermuth 1967
 Family Anguidae (Slowworms, etc.)
Genus Dopasia (Oriental Glass Lizards)
  *
Dopasia wegneri

Sumatra Glass Lizard

Teynie, David & Ohler 2010 Wermuth 1969
 Family Dibamidae (Blind Skinks)
Genus Dibamus (Oriental Blind Skinks)
 
Dibamus leucurus

White-tailed Blind Skink

Greer 1985 Milto & Bezman-Moseyko 2021 Teynie, David & Ohler 2010
 
Dibamus novaeguineae

Indo-Australian Blind Skink

Milto & Bezman-Moseyko 2021
 Family Gekkonidae (Typical Geckos)
Genus Cnemaspis (Asian Rock and Forest Geckos)
  *
Cnemaspis aceh

Aceh Forest Gecko

Iskandar, McGuire & Amarasinghe 2017
  *
Cnemaspis andalas

West Sumatran Forest Gecko

Iskandar, McGuire & Amarasinghe 2017
  *
Cnemaspis calderana

Maninjau Rock Gecko

Milto & Bezman-Moseyko 2021
  *
Cnemaspis minang

Minang Forest Gecko

Iskandar, McGuire & Amarasinghe 2017 Khandekar, Thackeray & Agarwal 2021
  *
Cnemaspis rajabasa

Mount Rajabasa Rock Gecko

Amarasinghe, Harvey, Riyanto & Smith 2015
  *
Cnemaspis tapanuli

Tapanuli Forest Gecko

Iskandar, McGuire & Amarasinghe 2017
Genus Cyrtodactylus (Australasian Bent-toed Geckos)
  *
Cyrtodactylus agamensis

Agam Bent-toed Gecko

Mecke, Kieckbusch, Hartmann & Kaiser 2016 Milto & Bezman-Moseyko 2021 Roesler, Richards & Gunther 2007
  *
Cyrtodactylus lateralis

Sumatran Bent-toed Gecko

Harvey, O'Connell, Wostl, Riyanto, Kurniawan, Smith & Grismer 2016 Malkmus & Kunkel 1992 Rooij 1915 Teynie, David & Ohler 2010 Wermuth 1965
 
Cyrtodactylus majulah

Singapore Bent-toed Gecko

Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020
 
Cyrtodactylus marmoratus

Marbled Bent-toed Gecko

Brongersma in Dammerman 1929 Malkmus & Kunkel 1992 Rooij 1915 Teynie, David & Ohler 2010
  *
Cyrtodactylus psarops

Speckle-faced Bent-toed Gecko

Harvey, O'Connell, Barraza, Riyanto, Kurniawan & Smith 2015 Mecke, Kieckbusch, Hartmann & Kaiser 2016
 
Cyrtodactylus quadrivirgatus

Four-striped Bent-toed Gecko

Teynie, David & Ohler 2010
  *
Cyrtodactylus semicinctus

Kerinci Bent-toed Gecko

Harvey, O'Connell, Barraza, Riyanto, Kurniawan & Smith 2015
Genus Gehyra (Four-clawed Geckos)
 
Gehyra mutilata

Common Four-clawed Gecko

Brongersma in Dammerman 1929 Gaulke, Abel & Erdelen 1997 Gaulke, Abel, Erdelen & Fritz 1998 Ladiges 1939 Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010
Genus Gekko (Tokays)
 
Gekko albomaculatus

White-spotted Tokay

Brongersma in Dammerman 1929 [as stentor] Grismer, Pinto, Quah, Anuar, Cota, McGuire, Iskandar, Wood & Grismer 2022 Loennberg & Rendahl 1925 [as stentor] Malkmus & Kunkel 1992 [as G. smithii] Rooij 1915 [as stentor] Teynie, David & Ohler 2010 [as G. smithii]
  *
Gekko brooksii

Sumatra Flap-legged Gecko

Brown, Supriatna & Ota 2000 Teynie, David & Ohler 2010 Wermuth 1965
 
Gekko gecko

Tokay

Brongersma in Dammerman 1929 [as verticillatus] Ladiges 1939 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Rooij 1915 [as verticillatus] Teynie, David & Ohler 2010
 
Gekko horsfieldii

Narrow-lobed Gliding Gecko

Milto & Bezman-Moseyko 2021 Smith 1935 [as Ptychozoon horsfieldii]
 
Gekko kuhli

Wide-lobed Gliding Gecko

Brongersma in Dammerman 1929 [as Ptychozoon homalocephalum] Ladiges 1939 [as Ptychozoon kuhli] Loennberg & Rendahl 1925 [as Ptychozoon homalocephalum] Malkmus & Kunkel 1992 [as Ptychozoon kuhli] Rooij 1915 [as Ptychozoon homalocephalum] Teynie, David & Ohler 2010 [as Ptychozoon kuhli]
 
Gekko monarchus

Spotted House Gecko

Brongersma in Dammerman 1929 Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010
Genus Hemidactylus (Half-toed Geckos)
 
Hemidactylus frenatus

Asian House Gecko

Brongersma in Dammerman 1929 Gaulke, Abel & Erdelen 1997 Gaulke, Abel, Erdelen & Fritz 1998 Ladiges 1939 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Loennberg & Rendahl 1925 Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010
 
Hemidactylus garnotii

Fox Gecko

Brongersma in Dammerman 1929 Mertens 1930 Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010
 
Hemidactylus platyurus

Flat-tailed Gecko

Brongersma in Dammerman 1929 Gaulke, Abel & Erdelen 1997 Gaulke, Abel, Erdelen & Fritz 1998 Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010
Genus Hemiphyllodactylus (Slender Geckos)
  *
Hemiphyllodactylus margarethae

Sumatran Slender Gecko

Grismer, Wood, Anuar, Muin, Quah, McGuire, Brown, Ngo & Pham 2013 Zug 2010
 
Hemiphyllodactylus typus

Indo-Pacific Slender Gecko

Brongersma in Dammerman 1929 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Mertens 1930 Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010 Zug 2010
 Family Lacertidae (Typical Lizards)
Genus Takydromus (Oriental Grass Lizards)
 
Takydromus sexlineatus

Six-lined Grass Lizard

Arnold 1997 Bischoff 1992 Brongersma in Dammerman 1929 Rooij 1915 Teynie, David & Ohler 2010
 Family Scincidae (Skinks)
Genus Dasia (Indo-Malayan Tree Skinks)
 
Dasia grisea

Grey Tree Skink

Brown & Alcala 1980 Inger & Brown 1980
 
Dasia olivacea

Olive Tree Skink

Brongersma in Dammerman 1929 Inger & Brown 1980 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Loennberg & Rendahl 1925 Malkmus & Kunkel 1992 Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010
Genus Eutropis (South Asian Sun Skinks)
 
Eutropis multifasciata

Many-lined Sun Skink

Brongersma in Dammerman 1929 Gaulke, Abel & Erdelen 1997 Gaulke, Abel, Erdelen & Fritz 1998 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Loennberg & Rendahl 1925 Malkmus & Kunkel 1992 Milto & Bezman-Moseyko 2021 O'Shea, Kusuma & Kaiser 2018 Rooij 1915 Teynie, David & Ohler 2010
 
Eutropis quinquecarinata

Five-keeled Sun Skink

Brongersma in Dammerman 1929 Rooij 1915
 
Eutropis rudis

Three-keeled Sun Skink

Amarasinghe, Chandramouli, Deuti, Campbell, Henkanaththegedara & Karunarathna 2020 Rooij 1915 Teynie, David & Ohler 2010 Voris 1977
 
Eutropis rugifera

Rough-scaled Sun Skink

Brongersma 1934 Smith 1935 Taylor 1963 Teynie, David & Ohler 2010
Genus Larutia (Malay Slender Forest Skinks)
  *
Larutia sumatrensis

Sumatran Two-toed Skink

Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010
Genus Lygosoma (Southeast Asian Writhing Skinks)
  *
Lygosoma quadrupes

Short-limbed Writhing Skink

Milto & Bezman-Moseyko 2021
Genus Riopa (Asian Gracile Skinks)
  *
Riopa opisthorhoda

Indragiri Gracile Skink

Rooij 1915 Teynie, David & Ohler 2010
  *
Riopa schneideri

Djapura Gracile Skink

Grismer, Quah, Duzulkafly & Yambun 2018 Rooij 1915 [as bampfyldei] Teynie, David & Ohler 2010 [as bampfyldei] Werner 1900
Genus Sphenomorphus (Indo-Pacific Forest Skinks)
 
Sphenomorphus anomalopus

Penang Forest Skink

Ladiges 1939 Loennberg & Rendahl 1925 Rooij 1915 Teynie, David & Ohler 2010
 
Sphenomorphus cyanolaemus

Blue-throated Forest Skink

Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Teynie, David & Ohler 2010
  *
Sphenomorphus malayanus

Sumatra Forest Skink

Rooij 1915 Teynie, David & Ohler 2010
 
Sphenomorphus sanctus

Yellow-lined Forest Skink

Brongersma in Dammerman 1929 Rooij 1915 Teynie, David & Ohler 2010
 
Sphenomorphus scotophilus

Selangor Forest Skink

Cox, Dijk, Nabhitabhata & Thirakhupt 1998 Milto & Bezman-Moseyko 2021
Genus Subdoluseps (Asian Agile Skinks)
 
Subdoluseps samajaya

Kuching Agile Skink

Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020
Genus Tytthoscincus (Dwarf Leaf-litter Skinks)
 
Tytthoscincus temminckii

Sunda Leaf-litter Skink

Brongersma 1942 Brongersma in Dammerman 1929 Mertens 1930 Rooij 1915 Teynie, David & Ohler 2010
 Family Varanidae (Monitor Lizards)
Genus Varanus (Monitor Lizards)
 
Varanus dumerilii

Harlequin Monitor

Brongersma in Dammerman 1929 Ladiges 1939 Mertens 1963 Rooij 1915 Teynie, David & Ohler 2010
 
Varanus rudicollis

Rough-necked Monitor

Boehme 2003 Gaulke, Abel, Erdelen & Fritz 1998 Ladiges 1939 Malkmus & Kunkel 1992 Mertens 1963 Rooij 1915 Teynie, David & Ohler 2010
 
Varanus salvator

Asian Water Monitor

Brongersma in Dammerman 1929 Gaulke, Abel & Erdelen 1997 Gaulke, Abel, Erdelen & Fritz 1998 Koch, Auliya, Schmitz, Kuch & Boehme 2007 Ladiges 1939 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Mertens 1930 Mertens 1963 Milto & Bezman-Moseyko 2021 Rooij 1915 Teynie, David & Ohler 2010

Snakes (Serpentes)
= endemicEX = extinct or assumed extinct

Species
References
 Family Acrochordidae (Australasian File Snakes)
Genus Acrochordus (Australasian File Snakes)
 
Acrochordus granulatus

Lesser File Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Wallach, Williams & Boundy 2014
 
Acrochordus javanicus

Java File Snake

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 McDiarmid, Campbell & Toure 1999 Rooij 1917 Teynie, David & Ohler 2010 Wallach, Williams & Boundy 2014
 Family Anomochilidae (Dwarf Pipe Snakes)
Genus Anomochilus (Dwarf Pipe Snakes)
 
Anomochilus weberi

Sumatran Dwarf Pipe Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 McDiarmid, Campbell & Toure 1999 Rooij 1917 Teynie, David & Ohler 2010 Wallach, Williams & Boundy 2014
 Family Colubridae (Typical Snakes or Colubrids)
Genus Ahaetulla (Oriental Whip Snakes)
 
Ahaetulla fasciolata

Speckle-headed Whip Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Wallach, Williams & Boundy 2014
 
Ahaetulla mycterizans

Big-eyed Green Whip Snake

Miralles & David 2010 Wallach, Williams & Boundy 2014
 
Ahaetulla prasina

Short-nosed Whip Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Lang 2013 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Malkmus & Kunkel 1992 Milto & Bezman-Moseyko 2021 Rooij 1917 Teynie, David & Ohler 2010 Wallach, Williams & Boundy 2014
Genus Anoplohydrus (Sumatran Keelback)
  *
Anoplohydrus aemulans

Sumatran Keelback

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Murphy 2007 Rooij 1917 Teynie, David & Ohler 2010 Wallach, Williams & Boundy 2014
Genus Boiga (Australasian Cat Snakes)
 
Boiga bengkuluensis

Bengkulu Cat Snake

Orlov, Kudryavtzev, Ryabov & Shumakov 2003 Teynie, David & Ohler 2010
 
Boiga cynodon

Dog-toothed Cat Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Mertens 1930 Milto & Bezman-Moseyko 2021 Orlov & Ryabov 2002 Ramadhan, Iskandar & Subasri 2010 Rooij 1917 Teynie, David & Ohler 2010
 
Boiga drapiezii

White-spotted Cat Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Milto & Bezman-Moseyko 2021 Orlov & Ryabov 2002 Ramadhan, Iskandar & Subasri 2010 Rooij 1917 Teynie, David & Ohler 2010
 
Boiga jaspidea

Jasper Cat Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Orlov & Ryabov 2002 Ramadhan, Iskandar & Subasri 2010 Rooij 1917 Teynie, David & Ohler 2010
 
Boiga melanota

Western Mangrove Cat Snake

Brongersma 1934 [as B. dendrophila occidentalis] David & Vogel 1996 [as B. dendrophila occidentalis] Gaulke, Abel, Erdelen & Fritz 1998 Haas 1950 [as B. dendrophila occidentalis] Iskandar & Colijn 2001 [as B. dendrophila occidentalis] Ladiges 1939 [as B. dendrophila] Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 [as B. dendrophila] Malkmus & Kunkel 1992 Orlov & Ryabov 2002 [as B. dendrophila occidentalis] Rooij 1917 [as B. dendrophila] Teynie, David & Ohler 2010
 
Boiga multomaculata

Marbled Cat Snake

Bosch 1985 David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Orlov & Ryabov 2002 Rooij 1917 Teynie, David & Ohler 2010
 
Boiga nigriceps

Dark-headed Cat Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Orlov & Ryabov 2002 Ramadhan, Iskandar & Subasri 2010 Rooij 1917 Teynie, David & Ohler 2010
Genus Calamaria (Oriental Reed Snakes)
 
Calamaria abstrusa

Padang Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Teynie, David & Ohler 2010
 
Calamaria albiventer

White-bellied Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Rooij 1917 [as indragirica] Teynie, David & Ohler 2010
  *
Calamaria alidae

Bengkulu Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Teynie, David & Ohler 2010
  *
Calamaria crassa

Thick Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Teynie, David & Ohler 2010
  *
Calamaria doederleini

Langkat Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
  *
Calamaria eiselti

West Sumatran Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Teynie, David & Ohler 2010
 
Calamaria forcarti

Deli Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Teynie, David & Ohler 2010
 
Calamaria leucogaster

Collared Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
 
Calamaria lumbricoidea

Variable Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Rooij 1917 [as stahlknechti] Teynie, David & Ohler 2010
  *
Calamaria margaritophora

Stripe-necked Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
  *
Calamaria mecheli

Riau Lowland Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
 
Calamaria modesta

Yellow-spotted Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Teynie, David & Ohler 2010
 
Calamaria schlegeli

Pink-headed Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Milto & Bezman-Moseyko 2021 Rooij 1917 [as agamensis] Teynie, David & Ohler 2010
  *
Calamaria sumatrana

Sumatran Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
  *
Calamaria ulmeri

North Sumatra Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Teynie, David & Ohler 2010
 
Calamaria virgulata

Blunt-tailed Reed Snake

David & Vogel 1996 Inger & Marx 1965 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
Genus Calamorhabdium (Indonesian Iridescent Snakes)
 
Calamorhabdium sp.

(Indonesian Iridescent Snake)

David & Vogel 1996
Genus Ceratophallus (Indonesian Striped Keelback)
 
Ceratophallus vittatus

Indonesian Striped Keelback

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
Genus Chrysopelea (Flying Tree Snakes)
 
Chrysopelea paradisi

Paradise Flying Tree Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Mertens 1968 Teynie, David & Ohler 2010
 
Chrysopelea pelias

Twin-barred Flying Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Mertens 1968 Rooij 1917 [as chrysochlora] Teynie, David & Ohler 2010
Genus Coelognathus (Oriental Rat Snakes)
 
Coelognathus flavolineatus

Yellow-striped Rat Snake

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Haas 1950 Iskandar & Colijn 2001 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Milto & Bezman-Moseyko 2021 Rooij 1917 [as melanurus] Schulz 1996 Teynie, David & Ohler 2010
 
Coelognathus radiatus

Copper-headed Rat Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Milto & Bezman-Moseyko 2021 Rooij 1917 Schulz 1996 Teynie, David & Ohler 2010
Genus Dendrelaphis (Bronzebacks)
 
Dendrelaphis caudolineatus

Malayan Stripe-tailed Bronzeback

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Milto & Bezman-Moseyko 2021 Rooij 1917 Rooijen & Vogel 2012 Teynie, David & Ohler 2010
 
Dendrelaphis formosus

Elegant Bronzeback

Iskandar & Colijn 2001 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Rooij 1917 Teynie, David & Ohler 2010 Vogel & Rooijen 2007
 
Dendrelaphis haasi

Greater Sundas Bronzeback

Rooijen & Vogel 2008 Stuebing, Inger & Lardner 2014 Teynie, David & Ohler 2010
 
Dendrelaphis kopsteini

Red-necked Bronzeback

Teynie, David & Ohler 2010 Vogel & Rooijen 2007
 
Dendrelaphis pictus

Southern Painted Bronzeback

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Haas 1950 Iskandar & Colijn 2001 Lang 2011 Milto & Bezman-Moseyko 2021 Rooij 1917 Rooijen & Vogel 2008 Teynie, David & Ohler 2010 Vogel & Rooijen 2008 Vogel & Rooijen 2011
 
Dendrelaphis striatus

Banded Bronzeback

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Teynie, David & Ohler 2010
Genus Dryophiops (Malayan Vine Snakes)
 
Dryophiops rubescens

Keel-bellied Vine Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
Genus Elaphe (Palearctic-Oriental Rat Snakes)
 
Elaphe taeniura

Stripe-tailed Rat Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Rooij 1917 Schulz 1996 Teynie, David & Ohler 2010
Genus Elapoidis (Sunda Mountain Snakes)
  *
Elapoidis sumatranus

Sumatra Mountain Snake

Bleeker 1860 David & Vogel 1996 [as fuscus] Haas 1950 [as fuscus] Iskandar & Colijn 2001 [as fuscus] Rooij 1917 [as fuscus] Teynie, David & Ohler 2010 [as fuscus] Vogel, David & Reza 2018
Genus Etheridgeum (Sumatran Burrowing Snake)
  *
Etheridgeum pulchrum

Sumatran Burrowing Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Teynie, David & Ohler 2010
Genus Fowlea (Fishing Keelbacks)
 
Fowlea melanzosta

Southern Keelback

David & Vogel 1996 Iskandar & Colijn 2001 Rooij 1917 [as piscator] Teynie, David & Ohler 2010 Vogel & David 2012
Genus Gongylosoma (Southeast Asian Ground Snakes)
 
Gongylosoma baliodeira

Orange-bellied Ground Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
 
Gongylosoma longicaudum

Striped Ground Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)
 
Gonyosoma oxycephalum

Red-tailed Green Rat Snake

David & Vogel 1996 Dowling 1958 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Rooij 1917 Schulz 1996 Teynie, David & Ohler 2010
Genus Hebius (Oriental Equatorial Keelbacks)
  *
Hebius kerinciensis

Kerinchi Keelback

David & Das 2003 David, Vogel & Rooijen 2013 Purkayastha & David 2019 Teynie, David & Ohler 2010 Wallach, Williams & Boundy 2014
 
Hebius petersii

Sarawak Spotted Keelback

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Wallach, Williams & Boundy 2014
  *
Hebius viperinus

Viper-like Keelback

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Wallach, Williams & Boundy 2014
Genus Iguanognathus (Iguana-jawed Snake)
  *
Iguanognathus werneri

Iguana-jawed Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
Genus Liopeltis (Ringneck Snakes)
 
Liopeltis tricolor

Malayan Ringneck Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
Genus Lycodon (Oriental Wolf Snakes)
 
Lycodon albofuscus

Slender Wolf Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
 
Lycodon capucinus

Island Wolf Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Lang 2013 Mertens 1930 O'Shea, Kusuma & Kaiser 2018 Rooij 1917 [as aulicus] Teynie, David & Ohler 2010
 
Lycodon effraenis

Scarce Wolf Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
  *
Lycodon sidiki

Aceh Wolf Snake

Wostl, Hamidy, Kurniawan & Smith 2017
 
Lycodon subannulatus

Malayan Bridle Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Rooij 1917 Teynie, David & Ohler 2010
 
Lycodon subcinctus

Malayan Banded Wolf Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Mertens 1930 Milto & Bezman-Moseyko 2021 Rooij 1917 Teynie, David & Ohler 2010
Genus Oligodon (Kukri Snakes)
 
Oligodon bitorquatus

Javanese Mountain Kukri Snake

David & Vogel 2012
 
Oligodon octolineatus

Eight-lined Kukri Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Tillack & Gunther 2009
  *
Oligodon petronellae

Sumatra Ornate Kukri Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Tillack & Gunther 2009
  *
Oligodon pulcherrimus

Padang Kukri Snake

David & Vogel 1996 Haas 1950 [as durheimi] Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Tillack & Gunther 2009
 
Oligodon purpurascens

Purple Kukri Snake

David & Vogel 1996 David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Tillack & Gunther 2009
 
Oligodon signatus

Barred Kukri Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Tillack & Gunther 2009
 
Oligodon trilineatus

Three-lined Kukri Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Tillack & Gunther 2009
Genus Opisthotropis (Mountain Stream Keelbacks)
  *
Opisthotropis rugosa

Sumatran Mountain Stream Keelback

David & Vogel 1996 David, Pauwels, Nguyen & Vogel 2015 Haas 1950 Iskandar & Colijn 2001 Iskandar & Kamsi 2009 Rooij 1917 Teynie, David & Ohler 2010
Genus Oreocryptophis (Red Mountain Rat Snake)
 
Oreocryptophis porphyraceus

Red Mountain Rat Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Schulz & Helfenberger 1998 Schulz 1996 Teynie, David & Ohler 2010
Genus Pseudorabdion (Dwarf Reed Snakes)
 
Pseudorabdion eiselti

Sumatra Dwarf Reed Snake

David & Vogel 1996 Iskandar & Colijn 2001 Milto & Bezman-Moseyko 2021 Teynie, David & Ohler 2010
 
Pseudorabdion longiceps

Long-headed Dwarf Reed Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Leviton & Brown 1959 Rooij 1917 Teynie, David & Ohler 2010
  *
Pseudorabdion modiglianii

Padang Dwarf Reed Snake

Doria & Petri 2010
  *
Pseudorabdion sirambense

Sirambi Dwarf Reed Snake

Doria & Petri 2010
Genus Pseudoxenodon (Mock Cobras)
  *
Pseudoxenodon jacobsonii

Sumatran Mock Cobra

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Teynie, David & Ohler 2010 [as inornatus]
Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)
 
Ptyas carinata

Malayan Keeled Rat Snake

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Haas 1950 Iskandar & Colijn 2001 Milto & Bezman-Moseyko 2021 Rooij 1917 Teynie, David & Ohler 2010
 
Ptyas fusca

White-bellied Rat Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Rooij 1917 Teynie, David & Ohler 2010
 
Ptyas korros

Indochinese Rat Snake

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Haas 1950 Iskandar & Colijn 2001 Milto & Bezman-Moseyko 2021 Rooij 1917 Teynie, David & Ohler 2010
 
Ptyas mucosa

Common Oriental Rat Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Rooij 1917 Sindaco, Venchi & Grieco 2013 Teynie, David & Ohler 2010
Genus Rhabdophis (Asian Keelbacks)
  *
Rhabdophis akraios

Mount Singalang Keelback

Doria, Petri, Bellati, Tiso & Pistarino 2013
 
Rhabdophis chrysargos

Speckled-bellied Keelback

David & Vogel 1996 Doria, Petri, Bellati, Tiso & Pistarino 2013 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Malkmus & Kunkel 1992 Milto & Bezman-Moseyko 2021 Rooij 1917 Teynie, David & Ohler 2010
 
Rhabdophis conspicillatus

Red-bellied Keelback

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
 
Rhabdophis flaviceps

Orange-necked Keelback

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Rooij 1917 Teynie, David & Ohler 2010
 
Rhabdophis rhodomelas

Blue-necked Keelback

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Rooij 1917 Teynie, David & Ohler 2010
 
Rhabdophis subminiatus

Southern Red-necked Keelback

David & Vogel 1996 David & Vogel 2021 Haas 1950 Iskandar & Colijn 2001 Teynie, David & Ohler 2010
Genus Sibynophis (Oriental Black-headed Snakes)
 
Sibynophis geminatus

Striped Black-headed Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Mertens 1930 Milto & Bezman-Moseyko 2021 Rooij 1917 Teynie, David & Ohler 2010
 
Sibynophis melanocephalus

White-lipped Black-headed Snake

David & Vogel 1996 Iskandar & Colijn 2001 Teynie, David & Ohler 2010
Genus Xenelaphis (Asian Brown Snakes)
 
Xenelaphis ellipsifer

Ocellated Brown Snake

Auliya & Shepherd 2009 David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Teynie, David & Ohler 2010
 
Xenelaphis hexagonotus

Sunda Brown Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
Genus Xenochrophis (Oriental Painted Keelbacks)
 
Xenochrophis maculatus

Malayan Spotted Keelback

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Malkmus & Kunkel 1992 Rooij 1917 Teynie, David & Ohler 2010
 
Xenochrophis trianguligerus

South Asian Triangle Keelback

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Milto & Bezman-Moseyko 2021 Rooij 1917 Teynie, David & Ohler 2010
 Family Cylindrophiidae (Asian Pipe Snakes)
Genus Cylindrophis (Oriental Pipe Snakes)
 
Cylindrophis ruffus

Red-tailed Pipe Snake

Bernstein, Bauer, McGuire, Arida, Kaiser, Kieckbusch & Mecke 2020 David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 McDiarmid, Campbell & Toure 1999 Rooij 1917 Teynie, David & Ohler 2010
 Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.)
Genus Bungarus (Kraits)
 
Bungarus candidus

Malayan Krait

David & Vogel 1996 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
 
Bungarus fasciatus

Banded Krait

David & Vogel 1996 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
 
Bungarus flaviceps

Red-headed Krait

David & Vogel 1996 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
Genus Calliophis (South Asian Coral Snakes)
 
Calliophis bivirgatus

Red-bellied Coral Snake

David & Vogel 1996 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Milto & Bezman-Moseyko 2021 Rooij 1917 Teynie, David & Ohler 2010
 
Calliophis gracilis

Graceful Coral Snake

David & Vogel 1996 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Vogel 2006
 
Calliophis intestinalis

Malayan Striped Coral Snake

David & Vogel 1996 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Haas 1950 Iskandar & Colijn 2001 Leviton 1963 Milto & Bezman-Moseyko 2021 Rooij 1917 Teynie, David & Ohler 2010 Vogel 2006
 
Calliophis nigrotaeniatus

Black-striped Coral Snake

Fukuyama, Vogel, Matsui, Eto, Munir, Hossman, Hamidy & Nishikawa 2020 Haas 1950 [as Maticora intestinalis sumatranus]
Genus Naja (Typical Cobras)
 
Naja sumatrana

Sunda Spitting Cobra

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Haas 1950 Iskandar & Colijn 2001 Milto & Bezman-Moseyko 2021 Rooij 1917 [as N. tripudians] Teynie, David & Ohler 2010 Wuster & Thorpe 1990
Genus Ophiophagus (King Cobra)
 
Ophiophagus hannah

King Cobra

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 [as bungarus] Teynie, David & Ohler 2010
 Family Elapidae [part 2: marine species] (Sea Snakes)
Genus Hydrophis (Typical Sea Snakes)
 
Hydrophis atriceps

Black-headed Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 [as H. fasciatus] Haas 1950 Iskandar & Colijn 2001 [as H. fasciatus] Rooij 1917 [as H. fasciatus]
 
Hydrophis brookii

Bornean Slender-necked Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Iskandar & Colijn 2001
 
Hydrophis caerulescens

Bluish Slender-necked Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Iskandar & Colijn 2001
 
Hydrophis curtus

Short Sea Snake

Barme in Bucherl, Buckley & Deulofeu (eds.) 1968 [as Lapemis hardwickii] Iskandar & Colijn 2001 [as Lapemis hardwickii] Smith 1926 [as Lapemis hardwickii]
 
Hydrophis cyanocinctus

Blue-banded Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Iskandar & Colijn 2001
 
Hydrophis jerdonii

Cone-nosed Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Iskandar & Colijn 2001
 
Hydrophis klossi

Selangor Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Iskandar & Colijn 2001
 
Hydrophis melanosoma

Black-banded Robust Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Haas 1950 Iskandar & Colijn 2001 Smith 1926
 
Hydrophis ornatus

Reef Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Iskandar & Colijn 2001
 
Hydrophis platurus

Yellow-bellied Sea Snake

Haas 1950 Rooij 1917
 
Hydrophis schistosus

Beaked Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 [as H. valakadyn]
 
Hydrophis spiralis

Narrow-banded Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Iskandar & Colijn 2001 Rooij 1917
 
Hydrophis torquatus

Grey Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Haas 1950 Iskandar & Colijn 2001 Smith 1926
 
Hydrophis viperinus

Viperine Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Iskandar & Colijn 2001
Genus Laticauda (Sea Kraits)
 
Laticauda colubrina

Yellow-lipped Sea Krait

Bucherl, Buckley & Deulofeu (eds.) 1968 Heatwole, Busack & Cogger 2005 Haas 1950 Iskandar & Colijn 2001 Kharin, Roedel & Hallermann 2010 McCarthy 1986 Rooij 1917
 
Laticauda laticaudata

Blue-lipped Sea Krait

Bucherl, Buckley & Deulofeu (eds.) 1968 Haas 1950 Iskandar & Colijn 2001 Kharin, Roedel & Hallermann 2010 McCarthy 1986 Rooij 1917 Smith 1926
Genus Microcephalophis (Small-headed Sea Snakes)
 
Microcephalophis gracilis

Graceful Small-headed Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Haas 1950 Iskandar & Colijn 2001
Genus Thalassophis (Scale-snouted Sea Snake)
 
Thalassophis anomalus

Scale-snouted Sea Snake

Bucherl, Buckley & Deulofeu (eds.) 1968 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Smith 1926
 Family Gerrhopilidae (Indo-Malayan Blind Snakes)
Genus Gerrhopilus (Indo-Malayan Blind Snakes)
  *
Gerrhopilus sumatranus

Sumatra Blind Snake

Wynn 2021
 Family Homalopsidae (Australasian Mud Snakes)
Genus Brachyorrhos (Moluccan Short-tailed Snakes)
 
Brachyorrhos albus

Ambon Short-tailed Snake

David & Vogel 1996 Milto & Bezman-Moseyko 2021
Genus Cerberus (Dog-faced Water Snakes)
 
Cerberus schneiderii

South East Asian Dog-faced Water Snake

David & Vogel 1996 [as C. rynchops] Iskandar & Colijn 2001 [as C. rynchops] Lang 2013 Rooij 1917 [as C. rynchops] Teynie, David & Ohler 2010 [as C. rynchops]
Genus Enhydris (Asian Mud Snakes)
 
Enhydris enhydris

Striped Mud Snake

David & Vogel 1996 Gyi 1970 Iskandar & Colijn 2001 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Rooij 1917 Teynie, David & Ohler 2010
Genus Fordonia (White-bellied Mangrove Snake)
 
Fordonia leucobalia

White-bellied Mangrove Snake

David & Vogel 1996 Iskandar & Colijn 2001 Lang 2013 Rooij 1917 Teynie, David & Ohler 2010
Genus Homalopsis (Masked Water Snakes)
 
Homalopsis buccata

Malayan Masked Water Snake

David & Vogel 1996 Gyi 1970 Iskandar & Colijn 2001 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Murphy & Voris 2014 Murphy 2007 Murphy, Voris, Murthy, Traub & Cumberbatch 2012 Rooij 1917 Teynie, David & Ohler 2010
Genus Hypsiscopus (Orange-bellied Mud Snakes and Matanna Mud Snake)
 
Hypsiscopus plumbea

Southern Orange-bellied Mud Snake

David & Vogel 1996 Gyi 1970 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
Genus Karnsophis (Sumatra Short-tailed Snake)
  *
Karnsophis siantaris

Sumatra Short-tailed Snake

Murphy & Voris 2013 Murphy & Voris 2014
Genus Miralia (Greater Sunda Mud Snake)
 
Miralia alternans

Greater Sunda Mud Snake

David & Vogel 1996 Gyi 1970 Haas 1950 Iskandar & Colijn 2001 Murphy & Voris 2014 Murphy 2007 Rooij 1917 Teynie, David & Ohler 2010
Genus Phytolopsis (Blackwater Mud Snake)
 
Phytolopsis punctata

Blackwater Mud Snake

David & Vogel 1996 Gyi 1970 Iskandar & Colijn 2001 Murphy & Voris 2014 Murphy 2007 Rooij 1917 Teynie, David & Ohler 2010
 Family Lamprophiidae (House Snakes, etc.)
Genus Psammodynastes (Asian Mock Vipers)
 
Psammodynastes pictus

Painted Mock Viper

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
 
Psammodynastes pulverulentus

Dusky Mock Viper

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Lang 2013 Rooij 1917 Teynie, David & Ohler 2010
 Family Pareidae (Oriental Slug Eaters)
Genus Aplopeltura (Blunt-headed Slug Eater)
 
Aplopeltura boa

Blunt-headed Slug Eater

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Poyarkov, Nguyen, Pawangkhanant, Yushchenko, Brakels, Nguyen, Nguyen, Suwannapoom, Orlov & Vogel 2022 Rooij 1917 Teynie, David & Ohler 2010 Wallach, Williams & Boundy 2014
Genus Asthenodipsas (Malayan Slug Eaters)
 
Asthenodipsas laevis

Smooth Slug Eater

David & Vogel 1996 Grossmann & Tillack 2003 Haas 1950 Iskandar & Colijn 2001 Milto & Bezman-Moseyko 2021 Quah, Grismer, Lim, Anuar & Imbun 2019 Rooij 1917 Teynie, David & Ohler 2010 Wallach, Williams & Boundy 2014
 
Asthenodipsas malaccana

Western Dark-necked Slug Eater

David & Vogel 1996 Grossmann & Tillack 2003 Haas 1950 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Wallach, Williams & Boundy 2014
  *
Asthenodipsas tropidonotus

Sumatra Slug Eater

David & Vogel 1996 Grossmann & Tillack 2003 [as vertebralis] Haas 1950 Iskandar & Colijn 2001 Loredo, Wood, Quah, Anuar, Greer, Ahmad & Grismer 2013 Teynie, David & Ohler 2010 [as vertebralis]
Genus Pareas (Typical Oriental Slug Eaters)
 
Pareas carinatus

Keeled Slug Eater

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Malkmus & Kunkel 1992 Mertens 1930 Poyarkov, Nguyen, Pawangkhanant, Yushchenko, Brakels, Nguyen, Nguyen, Suwannapoom, Orlov & Vogel 2022 Rooij 1917 Teynie, David & Ohler 2010
 
Pareas margaritophorus

White-spotted Slug Eater

Nguyen, Nguyen, Agung & Poyarkov 2020 Vogel, Nguyen, Lalremsanga, Biakzuala, Hrima & Poyarkov 2020
 
Pareas nuchalis

Barred Slug Eater

Poyarkov, Nguyen, Pawangkhanant, Yushchenko, Brakels, Nguyen, Nguyen, Suwannapoom, Orlov & Vogel 2022
 Family Pythonidae (Pythons)
Genus Malayopython (Malayan Pythons)
 
Malayopython reticulatus

Reticulated Python

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Iskandar & Colijn 2001 Ladiges 1939 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 McDiarmid, Campbell & Toure 1999 Milto & Bezman-Moseyko 2021 Rooij 1917 Stimson 1969 Teynie, David & Ohler 2010
Genus Python (Typical Pythons)
 
Python brongersmai

Blood Python

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Iskandar & Colijn 2001 Keogh, Barker & Shine 2001 Stimson 1969 Teynie, David & Ohler 2010
 
Python curtus

Sumatra Short-tailed Python

David & Vogel 1996 Gaulke, Abel, Erdelen & Fritz 1998 Iskandar & Colijn 2001 Keogh, Barker & Shine 2001 Stimson 1969 Teynie, David & Ohler 2010
 Family Typhlopidae (Typical Blind Snakes)
Genus Argyrophis (Southeast Asian Blind Snakes)
  *
Argyrophis hypsobothrius

Pit-snouted Blind Snake

David & Vogel 1996 Hahn 1980 Iskandar & Colijn 2001 McDiarmid, Campbell & Toure 1999 Teynie, David & Ohler 2010
 
Argyrophis muelleri

Malayan Bicoloured Blind Snake

Brongersma 1934 David & Vogel 1996 Hahn 1980 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010
Genus Ramphotyphlops (Western Pacific Blind Snakes)
 
Ramphotyphlops lineatus

Striped Blind Snake

David & Vogel 1996 Hahn 1980 Iskandar & Colijn 2001 McDiarmid, Campbell & Toure 1999 Rooij 1917 Teynie, David & Ohler 2010
Genus Virgotyphlops (Brahminy Blind Snake)
 
Virgotyphlops braminus

Brahminy Blind Snake

David & Vogel 1996 Hahn 1980 Iskandar & Colijn 2001 Ladiges 1939 Lang 2013 McDiarmid, Campbell & Toure 1999 Rooij 1917 Teynie, David & Ohler 2010 Wallach 2009
 Family Viperidae (Vipers)
Genus Craspedocephalus (Ridge-nosed Tree Pitvipers)
  *
Craspedocephalus andalasensis

Sumatran Palm Pitviper

David, Vogel, Vijayakumar & Vidal 2006 Teynie, David & Ohler 2010 Vogel 2006
 
Craspedocephalus puniceus

Ashy Pitviper

David & Vogel 1996 David, Vogel, Vijayakumar & Vidal 2006 Iskandar & Colijn 2001 Rooij 1917 Teynie, David & Ohler 2010 Vogel 2006
Genus Ovophis (Asian Mountain Pitvipers)
 
Ovophis convictus

Indo-Malayan Mountain Pitviper

David & Vogel 1996 Gumprecht & Issem 2013 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Iskandar & Colijn 2001 McDiarmid, Campbell & Toure 1999 [as monticola] Rooij 1917 [as monticola] Teynie, David & Ohler 2010 [as monticola]
Genus Parias (Malay Archipelago Pitvipers)
  *
Parias gunaleni

Green-eyed Sumatran Pitviper

Vogel, David & Sidik 2014 Vogel, David & Sidik 2022
 
Parias hageni

Bangka Green Pitviper

David & Vogel 1996 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Haas 1950 Iskandar & Colijn 2001 McDiarmid, Campbell & Toure 1999 Teynie, David & Ohler 2010 Vogel 2006 Vogel, David & Sidik 2022
 
Parias sumatranus

Grey-eyed Sumatran Pitviper

David & Vogel 1996 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Haas 1950 Iskandar & Colijn 2001 McDiarmid, Campbell & Toure 1999 Regenass & Kramer 1981 Rooij 1917 Teynie, David & Ohler 2010 Vogel 2006 Vogel, David & Sidik 2014 Vogel, David & Sidik 2022
Genus Popeia (South Asian Bamboo Pitvipers)
 
Popeia sabahi

Sunda Bamboo Pitviper

David & Vogel 1996 [as barati] David, Petri, Vogel & Doria 2009 [as toba] Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 [as barati] Iskandar & Colijn 2001 [as barati] Milto & Bezman-Moseyko 2021 Regenass & Kramer 1981 [as barati] Teynie, David & Ohler 2010 [as barati] Vogel 2006 [as barati] Vogel, David & Pauwels 2004 [as barati]
Genus Trimeresurus (Oriental Pitvipers)
 
Trimeresurus albolabris

Southeast Asian White-lipped Tree Pitviper

David & Vogel 1996 David & Vogel 2000 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Haas 1950 Iskandar & Colijn 2001 Regenass & Kramer 1981 Teynie, David & Ohler 2010
 
Trimeresurus purpureomaculatus

Mangrove Pitviper

David & Vogel 1996 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Haas 1950 Iskandar & Colijn 2001 McDiarmid, Campbell & Toure 1999 Regenass & Kramer 1981 Rooij 1917 Teynie, David & Ohler 2010
Genus Tropidolaemus (Temple Pitvipers)
 
Tropidolaemus wagleri

Speckled Pitviper

David & Vogel 1996 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Milto & Bezman-Moseyko 2021 Rooij 1917 Teynie, David & Ohler 2010 Vogel, David, Lutz, Rooijen & Vidal 2007
 Family Xenodermidae (Asian Litter Snakes)
Genus Xenodermus (Rough-backed Litter Snake)
 
Xenodermus javanicus

Rough-backed Litter Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Milto & Bezman-Moseyko 2021 Rooij 1917 Teynie, David & Ohler 2010
 Family Xenopeltidae (Asian Sunbeam Snakes)
Genus Xenopeltis (Asian Sunbeam Snakes)
 
Xenopeltis unicolor

Common Asian Sunbeam Snake

David & Vogel 1996 Haas 1950 Iskandar & Colijn 2001 Ladiges 1939 McDiarmid, Campbell & Toure 1999 Milto & Bezman-Moseyko 2021 Orlov 2000 Rooij 1917 Teynie, David & Ohler 2010
 Family Xenophidiidae (Spiny-jawed Snakes)
Genus Xenophidion (Spiny-jawed Snakes)
 
Xenophidion sp.

(Spiny-jawed Snake)

Quah, Grismer, Jetten, Wood, Miralles, Sah, Guek & Brady 2018

Turtles (Testudines)
= endemicEX = extinct or assumed extinct

Species
References
 Family Cheloniidae (Hard-shelled Sea Turtles)
Genus Chelonia (Green Sea Turtle)
 
Chelonia mydas

Green Sea Turtle

Njoman & Nuitja 1984 Polunin & Nuitja in Bjorndal (ed.) 1982 Pritchard 1979 Rooij 1915 Stringell, Bangkaru, Steeman & Bateman 2000
Genus Eretmochelys (Hawksbill Sea Turtle)
 
Eretmochelys imbricata

Hawksbill Sea Turtle

Njoman & Nuitja 1984 Nuitja & Lazell 1982 Polunin & Nuitja in Bjorndal (ed.) 1982 Pritchard 1979 Rooij 1915
Genus Lepidochelys (Ridley Sea Turtles)
 
Lepidochelys olivacea

Olive Ridley Sea Turtle

Njoman & Nuitja 1984
 Family Dermochelyidae (Leatherback Sea Turtle)
Genus Dermochelys (Leatherback Sea Turtle)
 
Dermochelys coriacea

Leatherback Sea Turtle

Njoman & Nuitja 1984 Nuitja & Lazell 1982 Polunin & Nuitja in Bjorndal (ed.) 1982 Rooij 1915
 Family Geoemydidae (Eastern Pond Turtles)
Genus Batagur (River Terrapins)
 
Batagur affinis

Southern River Terrapin

Fritz & Gaulke 1997 [as baska] Gaulke, Abel, Erdelen & Fritz 1998 [as baska] Moll, Platt, Chan, Horne, Platt, Praschag, Chen & Dijk 2015 Praschag, Sommer, McCarthy, Gemel & Fritz 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Teynie, David & Ohler 2010 [as baska]
 
Batagur borneoensis

Red-crowned Terrapin

Fritz & Havas 2006 Iverson 1992 Ladiges 1939 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Teynie, David & Ohler 2010 Wermuth & Mertens 1977
Genus Cuora (Asian Box Turtles)
 
Cuora amboinensis

Southeast Asian Box Turtle

Gaulke, Abel & Erdelen 1997 Gaulke, Abel, Erdelen & Fritz 1998 Iverson 1992 Ladiges 1939 Leo, Suherman, Permatasari, Suganda, Zulamri & Winarni 2020 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Rooij 1915 Schoppe & Das 2011 Teynie, David & Ohler 2010 Wermuth & Mertens 1977
Genus Cyclemys (Asian Leaf Turtles)
 
Cyclemys dentata

Common Leaf Turtle

Fritz, Guicking, Auer, Sommer, Wink & Hundsdoerfer 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Teynie, David & Ohler 2010
 
Cyclemys enigmatica

Sunda Leaf Turtle

Fritz, Guicking, Auer, Sommer, Wink & Hundsdoerfer 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021
Genus Heosemys (Forest Turtles)
 
Heosemys spinosa

Spiny Turtle

Fritz & Gaulke 1997 Fritz & Havas 2006 Gaulke, Abel, Erdelen & Fritz 1998 Iverson 1992 Ladiges 1939 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Rooij 1915 Teynie, David & Ohler 2010 Wermuth & Mertens 1977
Genus Notochelys (Malayan Flat-shelled Turtle)
 
Notochelys platynota

Malayan Flat-shelled Turtle

Fritz & Gaulke 1997 Gaulke, Abel, Erdelen & Fritz 1998 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Rooij 1915 Teynie, David & Ohler 2010 Wermuth & Mertens 1977
Genus Orlitia (Malaysian Giant Turtle)
 
Orlitia borneensis

Malaysian Giant Turtle

Fritz & Gaulke 1997 Gaulke, Abel, Erdelen & Fritz 1998 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Rooij 1915 Teynie, David & Ohler 2010 Wermuth & Mertens 1977
Genus Siebenrockiella (Black Marsh Turtle and Philippine Pond Turtle)
 
Siebenrockiella crassicollis

Black Marsh Turtle

Fritz & Gaulke 1997 Gaulke, Abel, Erdelen & Fritz 1998 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Rooij 1915 Teynie, David & Ohler 2010 Wermuth & Mertens 1977
 Family Testudinidae (Tortoises)
Genus Manouria (Indochinese Tortoises)
 
Manouria emys

Asian Giant Tortoise

Fritz & Gaulke 1997 Gaulke, Abel, Erdelen & Fritz 1998 Iverson 1992 Ladiges 1939 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Rooij 1915 Stanford, Wanchai, Schaffer, Schaffer & Thirakhupt 2015 Teynie, David & Ohler 2010 Wermuth & Mertens 1977
 Family Trionychidae (Softshell Turtles)
Genus Amyda (Southeast Asian Softshell Turtles)
 
Amyda cartilaginea

Sunda Softshell Turtle

Fritz & Gaulke 1997 Fritz, Gemel, Kehlmaier, Vamberger & Praschag 2014 Gaulke, Abel, Erdelen & Fritz 1998 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Rooij 1915 Teynie, David & Ohler 2010 Wermuth & Mertens 1977
Genus Chitra (Narrow-headed Softshell Turtles)
 
Chitra chitra

Southeast Asian Narrow-headed Softshell Turtle

Bonin, Devaux & Dupre 2006 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Teynie, David & Ohler 2010
Genus Dogania (Malayan Softshell Turtle)
 
Dogania subplana

Malayan Softshell Turtle

Fritz & Gaulke 1997 Fritz & Havas 2006 Gaulke, Abel, Erdelen & Fritz 1998 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Rooij 1915 Teynie, David & Ohler 2010 Wermuth & Mertens 1977
Genus Pelochelys (Indomalayan Giant Softshell Turtles)
 
Pelochelys cantorii

Asian Giant Softshell Turtle

Das 2008 Fritz & Gaulke 1997 Gaulke, Abel, Erdelen & Fritz 1998 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Rooij 1915 Teynie, David & Ohler 2010 Wermuth & Mertens 1977 [as bibroni]