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© Rune Midtgaard


Taxonomy of the family Cylindrophiidae
Bibliography of the genus Cylindrophis
Biodiversity of the genus Cylindrophis








Genus
Cylindrophis

Oriental Pipe Snakes, Asian Pipe Snakes, Cylinder Snakes

Walzenschlangen

Asiatiske Cylinderslanger

1828 Cylindrophis Wagler (type species: Cylindrophis resplendens Wagler 1828)
1829 Rouleau Guérin Méneville 1829-1838 (type species: Tortrix maculata Merrem = Anguis maculata Linnaeus 1758)
Contents: 15 species, of which 12 (80.0%) are endemic.
Endemism: 0% 100%
Remarks: Previously included in the family Aniliidae (e.g., Dowling 1959; Goin, Goin & Zug 1978) or the family Uropeltidae (e.g., Smith, Smith & Sawin 1977; Grazziotin, Zaher, Ferrarezzi, Klaczko, Bonatto & Wilkinson 2007). Now assigned to its own, separate family, Cylindrophiidae (e.g., Cundall, Wallach & Rossman 1993).
Distribution: Brunei, Cambodia, China (Fujian, Hainan), East Timor, Indonesia (Babar Islands, Bacan, Banggai Islands, Bangka, Belitung, Boano, Buton, Damar, Flores, Java, Kalimantan, Komodo, Lombok, Madura, Natuna Islands, Riau Islands, Sangir Islands, Sula Islands [incl. Sanana], Sulawesi, Sumatra, Sumbawa, Tanahjampea, Tanimbar Islands [incl. Yamdena], West Timor, Wetar), Laos, Malaysia (Sabah, Sarawak, West Malaysia [incl. Langkawi Archipelago, Penang Island]), Myanmar, Singapore, Sri Lanka, Thailand, Vietnam.

Cylindrophis aruensis

Damar Pipe Snake, (Aru Pipe Snake)

Damar-Walzenschlange, (Aru-Walzenschlange)

Damar-cylinderslange

1920 Cylindrophis aruensis Boulenger

Remarks: Presence in Aru Islands, the type locality, is questionable (Iskandar & Colijn 2001), although reported by some authors (e.g., Haas 1950; McDiarmid, Campbell & Touré 1999).
Distribution: Indonesia (Damar).


Cylindrophis boulengeri

Wetar Pipe Snake, (Boulenger's Pipe Snake)

Wetar-Walzenschlange

Wetar-cylinderslange

1911 Cylindrophis boulengeri Roux

Distribution: East Timor, Indonesia (Babar Islands, West Timor, Wetar).


Cylindrophis burmanus

Burmese Pipe Snake

Burmesische Walzenschlange

Burmesisk Cylinderslange

1943 Cylindrophis rufus burmanus Smith
Cylindrophis burmanus Iskandar & Colijn 2001

Remarks: Iskandar & Colijn (2001) listed burmanus as a separate species, but without discussion or references. Even after the separation of burmanus, they noted that ruffus is composite. Validity confirmed by Amarasinghe, Campbell, Hallermann, Sidik, Supriatna & Ineich (2015).
Distribution: Myanmar.


Cylindrophis engkariensis

Engkari Pipe Snake

Engkari-Walzenschlange

Engkari-cylinderslange

1994 Cylindrophis engkariensis Stuebing

Distribution: Malaysia (Sarawak).


Cylindrophis isolepis

Tanahjampea Pipe Snake

Tanajampea-Walzenschlange

Tanajampea-cylinderslange

1896 Cylindrophis isolepis Boulenger

Distribution: Indonesia (Tanahjampea).


Cylindrophis jodiae

Annam Pipe Snake, (Jodi's Pipe Snake)

Annam-Walzenschlange

Annam-cylinderslange

2015 Cylindrophis jodiae Amarasinghe, Ineich, Campbell & Hallermann in Amarasinghe, Campbell, Hallermann, Sidik, Supriatna & Ineich

Distribution: Malaysia (West Malaysia [incl. Penang Island]), Myanmar, Thailand, Vietnam.


Cylindrophis lineatus

Lined Pipe Snake, Striped Pipe Snake, (Blanford's Pipe Snake)

Gestreifte Walzenschlange

Stribet Cylinderslange

1880 Cylindrophis lineatus Dennys

Remarks: The type locality was originally "Singapore", but Figueroa & Low (2021) suggested to change this to "Borneo". These authors also demonstrated that the authorship should correctly be Dennys 1880, not Blanford 1881, as previously thought. They also questioned the presence of the species in Kalimantan, although reported by Das & Iskandar (2012) and Bernstein, Bauer, McGuire, Arida, Kaiser, Kieckbusch & Mecke (2020).
Distribution: Indonesia (Kalimantan), Malaysia (Sarawak).


Cylindrophis maculatus

Sri Lankan Pipe Snake, Blotched Pipe Snake

Srilankanische Walzenschlange, Ceylon-Walzenschlange

Srilankansk Cylinderslange

1758 Anguis maculata Linnaeus
Tortrix maculatus Oppel 1811
Ilysia maculata Lichtenstein 1823
Cylindrophis maculata Gray 1842
1768 Anguis decussata Laurenti (McDiarmid, Campbell & Touré 1999)
1768 Anguis hepatica Laurenti (McDiarmid, Campbell & Touré 1999)
1768 Anguis tessellata Laurenti (McDiarmid, Campbell & Touré 1999)

Distribution: Sri Lanka.


Cylindrophis melanotus

Black Pipe Snake, Sulawesi Pipe Snake

Schwarze Walzenschlange

Sort Cylinderslange

1830 Cylindrophis melanotus Wagler
Tortrix melanota Gray 1849
Cylindrophis rufa var. melanota Jan & Sordelli 1865
1844 Tortrix rufa var. Celebica Schlegel (McDiarmid, Campbell & Touré 1999)
1849 Tortrix rufa var. Celebensis Gray [unjustified emendation of Tortix rufa var. Celebica Schlegel 1844] (McDiarmid, Campbell & Touré 1999)
Tortrix celebensis Gray 1849
1927 Cylindrophis celebensis Smith [not Tortrix rufa var. Celebensis Gray 1849] (McDowell 1975; McDiarmid, Campbell & Touré 1999)
1935 Cylindrophis heinrichi Ahl (McDowell 1975; McDiarmid, Campbell & Touré 1999)

Remarks: Although reported from Halmahera (e.g., Ahl 1935), presence of melanotus in this island is questionable (Iskandar & Colijn 2001). Records from Boano refer to osheai (Kieckbusch, Mader, Kaiser & Mecke 2018).
Distribution: Indonesia (Bacan, Banggai Islands, Buton, Sangir Islands, Sula Islands [Sanana], Sulawesi).


Cylindrophis opisthorhodus

Lesser Sundas Pipe Snake, Fawn Sunda Pipe Snake

Sunda-Walzenschlange

Sunda-cylinderslange

1897 Cylindrophis opisthorhodus Boulenger

Distribution: Indonesia (Flores, Komodo, Lombok, Sumbawa).


Cylindrophis osheai

Boano Pipe Snake

Boano-Walzenschlange

Boano-cylinderslange

2018 Cylindrophis osheai Kieckbusch, Mader, Kaiser & Mecke

Distribution: Indonesia (Boano).





Cylindrophis ruffus

Red-tailed Pipe Snake, Red Pipe Snake, Common Pipe Snake

Rotschwanz-Walzenschlange, Rote Walzenschlange

Rød Cylinderslange, Malajisk Cylinderslange

1768 Anguis ruffa Laurenti
Eryx rufus Daudin 1803
Tortrix rufa Merrem 1820
Ilysia rufa Lichtenstein 1823
Cylindrophis rufa Gray 1842
1788 Anguis striatus Gmelin (Kieckbusch, Mecke, Hartmann, Ehrmantraut, O'Shea & Kaiser 2016)
1828 Cylindrophis resplendens Wagler (Kieckbusch, Mecke, Hartmann, Ehrmantraut, O'Shea & Kaiser 2016)
1849 Anguis rufa javanica Gray (status uncertain; Kieckbusch, Mecke, Hartmann, Ehrmantraut, O'Shea & Kaiser 2016)
2015 Cylindrophis mirzae Amarasinghe, Ineich, Campbell & Hallermann in Amarasinghe, Campbell, Hallermann, Sidik, Supriatna & Ineich (Kieckbusch, Mecke, Hartmann, Ehrmantraut, O'Shea & Kaiser 2016)

Remarks: Previously included burmanus and populations now assigned to jodiae. It is unclear whether certain populations of ruffus represent jodiae and/or unnamed species. Specimens north of Peninsular Malaysia belong either to burmanus or jodiae (Kieckbusch, Mecke, Hartmann, Ehrmantraut, O'Shea & Kaiser 2016), but are retained here as ruffus, pending further studies. Presence in Hong Kong needs confirmation, since there is only a single, 19th century record (Karsen, Lau & Bogadek 1998). Records from Boano refer to osheai (Kieckbusch, Mader, Kaiser & Mecke 2018).
Distribution: Brunei, Cambodia, China (Fujian, Hainan), Indonesia (Bacan, Bangka, Belitung, Buton, Java, Kalimantan, Madura, Natuna Islands, Riau Islands, Sangir Islands, Sula Islands [incl. Sanana], Sulawesi, Sumatra), Laos, Malaysia, (Sabah, Sarawak, West Malaysia [incl. Langkawi Archipelago]), Singapore, Thailand, Vietnam.


Cylindrophis ruffus
© Rune Midtgaard

Cylindrophis slowinskii

Kachin Pipe Snake, (Slowinski's Pipe Snake)

Kachin-Walzenschlange

Kachin-cylinderslange

2020 Cylindrophis slowinskii Bernstein, Bauer, McGuire, Arida, Kaiser, Kieckbusch & Mecke

Distribution: Myanmar.





Cylindrophis subocularis

Grabag Pipe Snake

Grabag-Walzenschlange

Grabag-cylinderslange

2016 Cylindrophis subocularis Kieckbusch, Mecke, Hartmann, Ehrmantraut, O'Shea & Kaiser

Distribution: Indonesia (Java).


Cylindrophis yamdena

Yamdena Pipe Snake

Yamdena-Walzenschlange

Yamdena-cylinderslange

1998 Cylindrophis yamdena Smith & Sidik

Distribution: Indonesia (Tanimbar Islands: Yamdena).