Reptiles of Brunei (Brunei Darussalam) Related checklists:  Borneo Bibliography of the reptiles of Brunei Taxonomic history of the reptiles of Brunei Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus porosus Saltwater Crocodile Das 2007   Crocodylus raninus Bornean Swamp Crocodile Das 2007  Family Gavialidae (Gharials) Genus Tomistoma (Malayan Gharial)   Tomistoma schlegelii Malayan Gharial Bezuijen, Shwedick, Sommerlad, Stevenson & Stuebing 2010 Stuebing, Bezuijen, Auliya & Voris 2006

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Aphaniotis (Earless Agamas)   Aphaniotis ornata Ornate Earless Agama Das 2007 Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama Das 1995 Das 2007 Genus Draco (Gliding Dragons)   Draco cornutus Horned Gliding Dragon Das 2007 Manthey 2008   Draco cristatellus Crested Gliding Dragon Das 1995   Draco melanopogon Black-bearded Gliding Dragon Das 1995 Das 2007   Draco quinquefasciatus Five-banded Gliding Dragon Das 1995 Das 2007   Draco sumatranus Sumatran Gliding Dragon Das 2007 Genus Gonocephalus (Western Anglehead Agamas)   Gonocephalus borneensis Bornean Anglehead Agama Das 1995 Das 2007 Manthey 2010   Gonocephalus grandis Greater Anglehead Agama Das 2007 Manthey 2010 Sah, Grafe, Dornburg, McLeod, Bauer, Wahab, Grismer & Watkins-Colwell 2016   Gonocephalus liogaster Blue-eyed Anglehead Agama Das 2007  Family Gekkonidae (Typical Geckos) Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus baluensis Kinabalu Bent-toed Gecko Das 1995 Das 2007 Sah, Grafe, Dornburg, McLeod, Bauer, Wahab, Grismer & Watkins-Colwell 2016   Cyrtodactylus hutan Gunung Mulu Giant Bent-toed Gecko Das 1995 [as consobrinus] Das 2007 [as consobrinus] Sah, Grafe, Dornburg, McLeod, Bauer, Wahab, Grismer & Watkins-Colwell 2016 [as consobrinus]   Cyrtodactylus ingeri North Bornean Bent-toed Gecko Das 1995 Das 2007   Cyrtodactylus malayanus Malayan Bent-toed Gecko Das 2007 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Das 1995 Das 2007 Genus Gekko (Tokays)   Gekko albofasciolatus White-banded Narrow-disked Gecko Das 2007 [as G. smithii] Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 [as G. smithii]   Gekko horsfieldii Narrow-lobed Gliding Gecko Das 1995 [as Ptychozoon horsfieldii] Das 2007 [as Ptychozoon horsfieldii]   Gekko kuhli Wide-lobed Gliding Gecko Das 2007 [as Ptychozoon kuhli] Shabrani & Grafe 2019   Gekko monarchus Spotted House Gecko Das 2007 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus frenatus Asian House Gecko Das 1995 Das 2007   Hemidactylus platyurus Flat-tailed Gecko Das 2007 Genus Hemiphyllodactylus (Slender Geckos)   Hemiphyllodactylus typus Indo-Pacific Slender Gecko Das 1995 Das 2007  Family Scincidae (Skinks) Genus Apterygodon (Bornean Striped Tree Skink)   Apterygodon vittatum Bornean Striped Tree Skink Das 1995 Das 2007 Genus Brachymeles (Indo-Malayan Slender Skinks)   Brachymeles sp. (Slender Skink) Das 2007 Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Das 2007 Genus Eutropis (South Asian Sun Skinks)   Eutropis multifasciata Many-lined Sun Skink Das 2007 O'Shea, Kusuma & Kaiser 2018 Sah, Grafe, Dornburg, McLeod, Bauer, Wahab, Grismer & Watkins-Colwell 2016   Eutropis rudis Three-keeled Sun Skink Das 1995 Das 2007   Eutropis rugifera Rough-scaled Sun Skink Das 2007 Genus Lamprolepis (Malay Archipelago Tree Skinks)   Lamprolepis nieuwenhuisii Kalimantan Tree Skink Abdulla & Grafe 2019 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus cyanolaemus Blue-throated Forest Skink Das 1995 Das 2007 Genus Subdoluseps (Asian Agile Skinks)   Subdoluseps bowringii Short-legged Agile Skink Das 2007 Genus Tropidophorus (Stream Skinks)   Tropidophorus beccarii Sarawak Stream Skink Das 1995 Das 2007   Tropidophorus brookei Bornean Stream Skink Das 1995 Das 2007 Sah, Grafe, Dornburg, McLeod, Bauer, Wahab, Grismer & Watkins-Colwell 2016 Genus Tytthoscincus (Dwarf Leaf-litter Skinks)   Tytthoscincus hallieri Bornean Leaf-litter Skink Das 2007  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus dumerilii Harlequin Monitor Das 2007   Varanus salvator Asian Water Monitor Das 1995 Das 2007

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Acrochordidae (Australasian File Snakes) Genus Acrochordus (Australasian File Snakes)   Acrochordus granulatus Lesser File Snake Das 2007 Wallach, Williams & Boundy 2014   Acrochordus javanicus Java File Snake Das 2007 Wallach, Williams & Boundy 2014  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla fasciolata Speckle-headed Whip Snake Das 1995 Das 2007 Iskandar & Colijn 2001 Wallach, Williams & Boundy 2014   Ahaetulla prasina Short-nosed Whip Snake Das 1995 Das 2007 Stuebing, Inger & Lardner 2014 Wallach, Williams & Boundy 2014 Genus Boiga (Australasian Cat Snakes)   Boiga cynodon Dog-toothed Cat Snake Das 2007   Boiga dendrophila Eastern Mangrove Cat Snake Das 2007 Iskandar & Colijn 2001   Boiga drapiezii White-spotted Cat Snake Das 1995 Das 2007   Boiga jaspidea Jasper Cat Snake Das 2007   Boiga nigriceps Dark-headed Cat Snake Das 2007 Stuebing, Inger & Lardner 2014 Genus Calamaria (Oriental Reed Snakes)   Calamaria grabowskyi Kalimantan Spotted Reed Snake Das 2007   Calamaria griswoldi Yellow-lined Reed Snake Das 2007   Calamaria leucogaster Collared Reed Snake Das 2006   Calamaria lovii Rejang River Reed Snake Das 2007   Calamaria lumbricoidea Variable Reed Snake Das 2007 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea paradisi Paradise Flying Tree Snake Das 2007 Iskandar & Colijn 2001   Chrysopelea pelias Twin-barred Flying Snake Das 1995 Das 2007 Iskandar & Colijn 2001 Stuebing, Inger & Lardner 2014 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus flavolineatus Yellow-striped Rat Snake Das 2007 Iskandar & Colijn 2001   Coelognathus radiatus Copper-headed Rat Snake Das 2007 Stuebing, Inger & Lardner 2014 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis caudolineatus Malayan Stripe-tailed Bronzeback Das 1995 Das 2007 Iskandar & Colijn 2001   Dendrelaphis kopsteini Red-necked Bronzeback Das 2007   Dendrelaphis pictus Southern Painted Bronzeback Das 2007 Iskandar & Colijn 2001 Genus Dryophiops (Malayan Vine Snakes)   Dryophiops rubescens Keel-bellied Vine Snake Das 2007 Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe taeniura Stripe-tailed Rat Snake Das 2007 Stuebing, Inger & Lardner 2014 Genus Gongylosoma (Southeast Asian Ground Snakes)   Gongylosoma baliodeira Orange-bellied Ground Snake Das 1995 Das 2007 Iskandar & Colijn 2001 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma oxycephalum Red-tailed Green Rat Snake Das 2007 Genus Hebius (Oriental Equatorial Keelbacks)   Hebius flavifrons White-fronted Keelback Das 1995 Das 2007 Iskandar & Colijn 2001 Sah, Grafe, Dornburg, McLeod, Bauer, Wahab, Grismer & Watkins-Colwell 2016 Stuebing, Inger & Lardner 2014 Wallach, Williams & Boundy 2014   Hebius sarawacensis Sarawak Mountain Keelback Das 2006 Stuebing, Inger & Lardner 2014 Wallach, Williams & Boundy 2014 Genus Hydrablabes (Bornean Water Snakes)   Hydrablabes periops Spectacled Water Snake Das 1995 Das 2007 Iskandar & Colijn 2001 Stuebing, Inger & Lardner 2014 Genus Liopeltis (Ringneck Snakes)   Liopeltis tricolor Malayan Ringneck Snake Das 1995 Das 2007 Iskandar & Colijn 2001 Stuebing, Inger & Lardner 2014 Genus Lycodon (Oriental Wolf Snakes)   Lycodon albofuscus Slender Wolf Snake Grismer & Chan-Ard 2012   Lycodon effraenis Scarce Wolf Snake Das 2007   Lycodon sealei Palawan Wolf Snake Iskandar & Colijn 2001   Lycodon subannulatus Malayan Bridle Snake Das 2007 Stuebing, Inger & Lardner 2014   Lycodon subcinctus Malayan Banded Wolf Snake Das 1995 Das 2007   Lycodon tristrigatus Three-banded Bridle Snake Das 2007 Dehling & Das 2006 Genus Oligodon (Kukri Snakes)   Oligodon annulifer Indonesian Spotted Kukri Snake Das 1995 Das 2007 Iskandar & Colijn 2001 Stuebing, Inger & Lardner 2014   Oligodon octolineatus Eight-lined Kukri Snake Das 2007 Iskandar & Colijn 2001 Qian, Qi, Shi, Lu, Jenkins, Mo & Li 2021 Stuebing, Inger & Lardner 2014   Oligodon purpurascens Purple Kukri Snake Das 2007   Oligodon signatus Barred Kukri Snake Chau & Grafe 2019 Genus Opisthotropis (Mountain Stream Keelbacks)   Opisthotropis typica Dwarf Mountain Stream Keelback Das 1995 Das 2007 Iskandar & Colijn 2001 Iskandar & Kamsi 2009 Stuebing, Inger & Lardner 2014 Genus Pseudorabdion (Dwarf Reed Snakes)   Pseudorabdion albonuchalis Ring-necked Dwarf Reed Snake Das 1995 Das 2007 Iskandar & Colijn 2001   Pseudorabdion longiceps Long-headed Dwarf Reed Snake Das 1995 Das 2007 Iskandar & Colijn 2001 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas carinata Malayan Keeled Rat Snake Das 2007 Stuebing, Inger & Lardner 2014   Ptyas fusca White-bellied Rat Snake Das 2007 Iskandar & Colijn 2001 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis chrysargos Speckled-bellied Keelback Das 2007 Iskandar & Colijn 2001 Stuebing, Inger & Lardner 2014 Genus Sibynophis (Oriental Black-headed Snakes)   Sibynophis melanocephalus White-lipped Black-headed Snake Das 2007 Dehling 2006 Genus Xenelaphis (Asian Brown Snakes)   Xenelaphis hexagonotus Sunda Brown Snake Das 2007 Stuebing, Inger & Lardner 2014 Genus Xenochrophis (Oriental Painted Keelbacks)   Xenochrophis maculatus Malayan Spotted Keelback Das 2007 Iskandar & Colijn 2001 Stuebing, Inger & Lardner 2014   Xenochrophis trianguligerus South Asian Triangle Keelback Das 2007 Iskandar & Colijn 2001  Family Cylindrophiidae (Asian Pipe Snakes) Genus Cylindrophis (Oriental Pipe Snakes)   Cylindrophis ruffus Red-tailed Pipe Snake Das 2007 Stuebing, Inger & Lardner 2014  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Bungarus (Kraits)   Bungarus fasciatus Banded Krait Das 2007   Bungarus flaviceps Red-headed Krait Das 2007 Genus Calliophis (South Asian Coral Snakes)   Calliophis bivirgatus Red-bellied Coral Snake Das 1995 Das 2007 Iskandar & Colijn 2001   Calliophis intestinalis Malayan Striped Coral Snake Das 1995 Das 2007 Iskandar & Colijn 2001 Genus Naja (Typical Cobras)   Naja sumatrana Sunda Spitting Cobra Das 2007 Iskandar & Colijn 2001 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Das 2007 Iskandar & Colijn 2001  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis curtus Short Sea Snake Das 2007   Hydrophis cyanocinctus Blue-banded Sea Snake Das 2007 Stuebing, Inger & Lardner 2014   Hydrophis jerdonii Cone-nosed Sea Snake Das 2007 Stuebing, Inger & Lardner 2014   Hydrophis melanosoma Black-banded Robust Sea Snake Stuebing, Inger & Lardner 2014   Hydrophis ornatus Reef Sea Snake Stuebing, Inger & Lardner 2014   Hydrophis platurus Yellow-bellied Sea Snake Das 1992 Das 2007 Stuebing, Inger & Lardner 2014 Genus Kolpophis (Granular Sea Snake)   Kolpophis annandalei Granular Sea Snake Das 1993 Das 2007 Iskandar & Colijn 2001 Stuebing 1994 Stuebing, Inger & Lardner 2014 Genus Laticauda (Sea Kraits)   Laticauda colubrina Yellow-lipped Sea Krait Das 1992 Das 2007  Family Homalopsidae (Australasian Mud Snakes) Genus Cerberus (Dog-faced Water Snakes)   Cerberus schneiderii South East Asian Dog-faced Water Snake Das 2007 [as C. rynchops] Genus Enhydris (Asian Mud Snakes)   Enhydris enhydris Striped Mud Snake Das 2007 Genus Homalophis (Bornean Mud Snakes)   Homalophis doriae Sarawak Mud Snake Das 2007 Genus Hypsiscopus (Orange-bellied Mud Snakes and Matanna Mud Snake)   Hypsiscopus plumbea Southern Orange-bellied Mud Snake Das 2007 Genus Phytolopsis (Blackwater Mud Snake)   Phytolopsis punctata Blackwater Mud Snake Das 1995 Das 2007 Iskandar & Colijn 2001  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pictus Painted Mock Viper Das 1995 Das 2007 Iskandar & Colijn 2001   Psammodynastes pulverulentus Dusky Mock Viper Das 2007  Family Pareidae (Oriental Slug Eaters) Genus Aplopeltura (Blunt-headed Slug Eater)   Aplopeltura boa Blunt-headed Slug Eater Das 1995 Das 2007 Iskandar & Colijn 2001 Wallach, Williams & Boundy 2014 Genus Asthenodipsas (Malayan Slug Eaters)   Asthenodipsas borneensis Bornean Dark-necked Slug Eater Das 2007 [as vertebralis] Genus Pareas (Typical Oriental Slug Eaters)   Pareas carinatus Keeled Slug Eater Das 2007 Poyarkov, Nguyen, Pawangkhanant, Yushchenko, Brakels, Nguyen, Nguyen, Suwannapoom, Orlov & Vogel 2022   Pareas nuchalis Barred Slug Eater Ding, Chen, Suwannapoom, Nguyen, Pogarkov & Vogel 2020 Poyarkov, Nguyen, Pawangkhanant, Yushchenko, Brakels, Nguyen, Nguyen, Suwannapoom, Orlov & Vogel 2022  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Das 1995 Das 2007 McDiarmid, Campbell & Toure 1999 Sah, Grafe, Dornburg, McLeod, Bauer, Wahab, Grismer & Watkins-Colwell 2016 Genus Python (Typical Pythons)   Python breitensteini Borneo Short-tailed Python Das 2007  Family Typhlopidae (Typical Blind Snakes) Genus Virgotyphlops (Brahminy Blind Snake)   Virgotyphlops braminus Brahminy Blind Snake Das 2007 Iskandar & Colijn 2001 Wallach 2009  Family Viperidae (Vipers) Genus Craspedocephalus (Ridge-nosed Tree Pitvipers)   Craspedocephalus borneensis Bornean Flat-nosed Pitviper Das 1995 Das 2007 David, Vogel, Vijayakumar & Vidal 2006 Vogel 2006 Genus Parias (Malay Archipelago Pitvipers)   Parias sumatranus Grey-eyed Sumatran Pitviper Keller 2008 Genus Popeia (South Asian Bamboo Pitvipers)   Popeia sabahi Sunda Bamboo Pitviper Das 2007 Genus Tropidolaemus (Temple Pitvipers)   Tropidolaemus subannulatus North Philippine Temple Pitviper Das 1995 [as wagleri] Das 2007 [as wagleri] Sah, Grafe, Dornburg, McLeod, Bauer, Wahab, Grismer & Watkins-Colwell 2016 Vogel, David, Lutz, Rooijen & Vidal 2007  Family Xenodermidae (Asian Litter Snakes) Genus Xenodermus (Rough-backed Litter Snake)   Xenodermus javanicus Rough-backed Litter Snake Keller 2008  Family Xenopeltidae (Asian Sunbeam Snakes) Genus Xenopeltis (Asian Sunbeam Snakes)   Xenopeltis unicolor Common Asian Sunbeam Snake Das 2007

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Das 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Das 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Das 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Das 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Geoemydidae (Eastern Pond Turtles) Genus Batagur (River Terrapins)   Batagur borneoensis Red-crowned Terrapin Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle Das 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Schoppe & Das 2011 Genus Cyclemys (Asian Leaf Turtles)   Cyclemys dentata Common Leaf Turtle Das 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Heosemys (Forest Turtles)   Heosemys spinosa Spiny Turtle Charles 2000 Das 1995 Das 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Notochelys (Malayan Flat-shelled Turtle)   Notochelys platynota Malayan Flat-shelled Turtle Das 1995 Das 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Testudinidae (Tortoises) Genus Manouria (Indochinese Tortoises)   Manouria emys Asian Giant Tortoise Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Trionychidae (Softshell Turtles) Genus Amyda (Southeast Asian Softshell Turtles)   Amyda cartilaginea Sunda Softshell Turtle Auliya, Dijk, Moll & Meylan 2016 Das 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Dogania (Malayan Softshell Turtle)   Dogania subplana Malayan Softshell Turtle Das 1995 Das 2007 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sah, Grafe, Dornburg, McLeod, Bauer, Wahab, Grismer & Watkins-Colwell 2016 Genus Pelochelys (Indomalayan Giant Softshell Turtles)   Pelochelys cantorii Asian Giant Softshell Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021