Reptiles of Penang Island  [Malaysia: West Malaysia] Related checklists:  Malaysia - West Malaysia Bibliography of the reptiles of Penang Island Taxonomic history of the reptiles of Penang Island Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthosaura (Pricklenapes)   Acanthosaura armata Peninsular Pricklenape Grismer 2011 Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama Grismer 2011 Genus Calotes (South Asian Forest Agamas)   Calotes versicolor South Indian Crested Agama Grismer 2011 Jaafar, Shahrudin, Ibrahim & Ahmad 2015 Genus Draco (Gliding Dragons)   Draco abbreviatus Orange-bearded Gliding Dragon Grismer 2011   Draco formosus Malay Peninsula Dusky Gliding Dragon Grismer 2011   Draco melanopogon Black-bearded Gliding Dragon Grismer 2011   Draco quinquefasciatus Five-banded Gliding Dragon Grismer 2011   Draco sumatranus Sumatran Gliding Dragon Grismer 2011 Genus Gonocephalus (Western Anglehead Agamas)   Gonocephalus bellii Blue-necked Anglehead Agama Grismer 2011   Gonocephalus grandis Greater Anglehead Agama Grismer 2011  Family Gekkonidae (Typical Geckos) Genus Cnemaspis (Asian Rock and Forest Geckos)   * Cnemaspis affinis Penang Island Rock Gecko Grismer & Chan 2010 Grismer 2011 Grismer, Grismer, Wood & Chan 2008 Grismer, Wood, Anuar, Riyanto, Ahmad, Muin, Sumontha, Grismer, Chan, Quah & Pauwels 2014 Quah, Wood, Anuar, Muin & Grismer 2020   Cnemaspis shahruli Telok Bahang Rock Gecko Grismer 2011 Grismer, Chan, Quah, Muin, Savage, Grismer, Ahmad, Greer & Remegio 2010 Grismer, Wood, Anuar, Riyanto, Ahmad, Muin, Sumontha, Grismer, Chan, Quah & Pauwels 2014 Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus consobrinus Giant Bent-toed Gecko Grismer 2011   * Cyrtodactylus pulchellus Penang Island Bent-toed Gecko Grismer 2011 Grismer, Wood, Anuar, Quah, Muin, Mohamed, Chan, Sumarli, Loredo & Heinz 2014 Grismer, Wood, Quah, Anuar, Muin, Sumontha, Ahmad, Bauer, Wangkulangkul, Grismer & Pauwels 2012   Cyrtodactylus quadrivirgatus Four-striped Bent-toed Gecko Grismer 2011 Jaafar, Shahrudin, Ibrahim & Ahmad 2015 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Grismer 2011 Genus Gekko (Tokays)   Gekko gecko Tokay Grismer 2011   Gekko horsfieldii Narrow-lobed Gliding Gecko Grismer 2011 [as Ptychozoon horsfieldii]   Gekko kuhli Wide-lobed Gliding Gecko Grismer 2011 [as Ptychozoon kuhli]   Gekko monarchus Spotted House Gecko Grismer 2011   Gekko smithii Greater Green-eyed Gecko Grismer 2011 Grismer, Pinto, Quah, Anuar, Cota, McGuire, Iskandar, Wood & Grismer 2022 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus craspedotus Malayan Frilled Gecko Grismer 2011   Hemidactylus frenatus Asian House Gecko Grismer 2011 Jaafar, Shahrudin, Ibrahim & Ahmad 2015 Lajmi, Bansal, Giri & Karanth 2019 Lajmi, Giri & Karanth 2016   Hemidactylus murrayi Pimpri Half-toed Gecko Agarwal, Bauer, Giri & Khandekar 2019 Chaitanya, Agarwal, Lajmi & Khandekar 2019 Grismer 2011 [as brookii] Lajmi, Bansal, Giri & Karanth 2019 Lajmi, Giri & Karanth 2016   Hemidactylus platyurus Flat-tailed Gecko Grismer 2011 Jaafar, Shahrudin, Ibrahim & Ahmad 2015   Hemidactylus tenkatei Rote Half-toed Gecko Kathriner, O'Shea & Kaiser 2014 Genus Hemiphyllodactylus (Slender Geckos)   Hemiphyllodactylus cicak Penang Slender Gecko Cobos, Grismer, Wood, Quah, Anuar & Muin 2016 Grismer & Quah 2019 Genus Lepidodactylus (Scaly-toed Geckos)   Lepidodactylus lugubris Common Mourning Gecko Grismer 2011  Family Lacertidae (Typical Lizards) Genus Takydromus (Oriental Grass Lizards)   Takydromus sexlineatus Six-lined Grass Lizard Grismer 2011  Family Scincidae (Skinks) Genus Dasia (Indo-Malayan Tree Skinks)   Dasia grisea Grey Tree Skink Grismer 2011   Dasia olivacea Olive Tree Skink Grismer 2011 Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Grismer 2011 Genus Eutropis (South Asian Sun Skinks)   Eutropis multifasciata Many-lined Sun Skink Grismer 2011 Jaafar, Shahrudin, Ibrahim & Ahmad 2015 Genus Larutia (Malay Slender Forest Skinks)   * Larutia penangensis Penang Island Limbless Skink Grismer 2011 Grismer, Huat, Siler, Chan, Wood, Grismer, Sah & Ahmad 2011 Genus Lipinia (Indo-Pacific Striped Skinks)   Lipinia vittigera Sipora Striped Skink Grismer 2011 Poyarkov, Geissler, Gorin, Dunayev, Hartmann & Suwannapoom 2019 Genus Lygosoma (Southeast Asian Writhing Skinks)   Lygosoma siamense Siamese Writhing Skink Grismer 2011 [as L. quadrupes] Siler, Heitz, Davis, Freitas, Aowphol, Termprayoon & Grismer 2018 Genus Riopa (Asian Gracile Skinks)   Riopa albopunctata White-spotted Gracile Skink Grismer 2011 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus anomalopus Penang Forest Skink Grismer 2011   Sphenomorphus scotophilus Selangor Forest Skink Grismer 2011 Genus Subdoluseps (Asian Agile Skinks)   Subdoluseps bowringii Short-legged Agile Skink Grismer 2011 Wood, Grismer & Muin 2008   Subdoluseps herberti Thai Peninsular Agile Skink Grismer & Quah 2019 Genus Toenayar (Nine-keeled Sun Skink)   Toenayar novemcarinata Nine-keeled Sun Skink Grismer & Quah 2019 Grismer 2011 Genus Tytthoscincus (Dwarf Leaf-litter Skinks)   Tytthoscincus butleri Bintang Leaf-litter Skink Grismer & Quah 2019 Grismer, Wood, Quah, Anuar, Ngadi, Mohd-Izam & Ahmad 2017   Tytthoscincus panchorensis Mount Panchor Leaf-litter Skink Grismer & Quah 2019  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus dumerilii Harlequin Monitor Grismer 2011   Varanus nebulosus Clouded Monitor Grismer 2011   Varanus salvator Asian Water Monitor Grismer 2011 Jaafar, Shahrudin, Ibrahim & Ahmad 2015

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake Jaafar, Shahrudin, Ibrahim & Ahmad 2015 Genus Calamaria (Oriental Reed Snakes)   Calamaria albiventer White-bellied Reed Snake Flower 1899 Quah, Lim, Leong & Anuar 2018   Calamaria lumbricoidea Variable Reed Snake Quah, Lim, Leong & Anuar 2018   Calamaria pavimentata Brown Reed Snake Quah, Lim, Leong & Anuar 2018   Calamaria schlegeli Pink-headed Reed Snake Quah, Lim, Leong & Anuar 2018 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis vogeli Yunnan Bronzeback Biakzuala, Lalremsanga, Tariang, Vabeiryureilai, Muansanga, Hrima, Kumar, Kundu, Purkayastha & Vogel 2022 Genus Liopeltis (Ringneck Snakes)   Liopeltis tricolor Malayan Ringneck Snake Som, Grismer, Wood, Quah, Brown, Diesmos, Weinell & Stuart 2020 Genus Pseudorabdion (Dwarf Reed Snakes)   Pseudorabdion longiceps Long-headed Dwarf Reed Snake Quah, Lim, Leong & Anuar 2018  Family Cylindrophiidae (Asian Pipe Snakes) Genus Cylindrophis (Oriental Pipe Snakes)   Cylindrophis jodiae Annam Pipe Snake Bernstein, Bauer, McGuire, Arida, Kaiser, Kieckbusch & Mecke 2020  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis spiralis Narrow-banded Sea Snake Smith 1926  Family Homalopsidae (Australasian Mud Snakes) Genus Homalopsis (Masked Water Snakes)   Homalopsis buccata Malayan Masked Water Snake Jaafar, Shahrudin, Ibrahim & Ahmad 2015  Family Pareidae (Oriental Slug Eaters) Genus Asthenodipsas (Malayan Slug Eaters)   Asthenodipsas laevis Smooth Slug Eater Quah, Grismer, Lim, Anuar & Imbun 2019  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Jaafar, Shahrudin, Ibrahim & Ahmad 2015  Family Viperidae (Vipers) Genus Ovophis (Asian Mountain Pitvipers)   Ovophis convictus Indo-Malayan Mountain Pitviper Gumprecht & Issem 2013 Genus Trimeresurus (Oriental Pitvipers)   Trimeresurus purpureomaculatus Mangrove Pitviper Jaafar, Shahrudin, Ibrahim & Ahmad 2015 Genus Tropidolaemus (Temple Pitvipers)   Tropidolaemus wagleri Speckled Pitviper Jaafar, Shahrudin, Ibrahim & Ahmad 2015

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Geoemydidae (Eastern Pond Turtles) Genus Heosemys (Forest Turtles)   Heosemys spinosa Spiny Turtle Jaafar, Shahrudin, Ibrahim & Ahmad 2015  Family Testudinidae (Tortoises) Genus Indotestudo (Asian Tortoises)   Indotestudo elongata Elongated Tortoise Fritz & Havas 2006 Ihlow, Dawson, Hartmann & Som 2016