Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk).
Latest update: November 9th, 2023.
 
|
Genus
Gloydius
 |
Mamushis, Asian Moccasins, (Asian Copperheads)
|
 |
Mamushis, Halysottern
|
 |
Mamushier
|
|
1822 |  |
Trigonocephalus Kuhl & van Hasselt [not Trigonocephalus Oppel 1811 (Serpentes)] (type species: Coluber halys Pallas 1776) |
|
1826 | |
Trigonocephalus Boie [not Trigonocephalus Oppel 1811 (Serpentes); not Trigonocephalus Kuhl & van Hasselt 1822 (Serpentes)] (type species: Coluber Halys Pallas 1776) |
|
1849 | |
Halys Gray [not Halys Fabricius 1803 (Insecta)] (type species: Coluber halys Pallas 1776) |
|
1981 | |
Gloydius Hoge & Romano Hoge [substitute name for Halys Gray 1849] (type species: Coluber halys Pallas 1776) |
Contents:
27 species, of which 18 (66.7%) are endemic.
Endemism: 0%  100%
Remarks:
Previously included in
Agkistrodon (e.g., Gloyd & Conant 1990). Validity as a separate genus confirmed by several subsequent works (Knight, Densmore & Rael 1992; Parkinson, Moody & Ahlquist 1996; Cullings, Morafka, Hernandez & Roberts 1997; Vidal, Lecointre, Vie & Gasc 1997). Validity of several nominal species is still unresolved, and various authorities have suggested different arrangements.
Distribution:
SW., C., and E. Asia.
Reported from:
Afghanistan,
Azerbaijan,
Bhutan,
China
(Anhui,
Beijing,
Chongqing,
Fujian,
Gansu,
Guangdong,
Guizhou,
Hebei,
Heilongjiang,
Henan,
Hubei,
Hunan,
Inner Mongolia,
Jiangsu,
Jiangxi,
Jilin,
Liaoning,
Ningxia,
Qinghai,
Shandong
[incl.
Daheishan Islands],
Shaanxi,
Shanxi,
Sichuan,
Tibet,
Xinjiang,
Yunnan,
Zhejiang),
India
(Haryana,
Himachal Pradesh,
Jammu & Kashmir,
Punjab,
Uttar Pradesh,
Uttarakhand,
West Bengal),
Iran,
Japan
(Hokkaido,
Honshu,
Kyushu,
Shikoku,
Tsushima Islands,
Izu Islands,
Ryukyu Islands
[incl.
Satsunan Islands]),
Kazakhstan,
Kyrgyzstan,
Mongolia,
Nepal,
North Korea,
Pakistan,
Russia
(incl.
Kuril Islands),
South Korea,
Tajikistan,
Turkmenistan,
Uzbekistan.
| |
|
Gloydius angusticeps
|
Narrow-headed Mamushi, Zoige Pitviper
|
|
Schmalkopf-Mamushi
|
|
Smalhovedet Mamushi
|
|
2018 | |
Gloydius angusticeps Shi, Yang, Zhang, Peng, Orlov, Jiang, Ding, Hou, Huang, Huang & Li |
Distribution:
China
(Gansu,
Qinghai,
Sichuan).
|
|
|
Gloydius blomhoffii
|
Japanese Mamushi, Common Mamushi
|
|
Japanische Mamushi
|
|
Japansk Mamushi
|
|
1826 | |
Trigonocephalus Blomhoffii Boie |
|
| |
Halys blomhoffii Peters 1862 |
|
| |
Ancistrodon blomhoffii Boulenger 1896 |
|
| |
Agkistrodon blomhoffii Stejneger 1907 |
|
| |
Ancistrodon halys blomhoffii Nikolsky 1916 |
|
| |
Gloydius blomhoffii Hoge & Romano Hoge 1981 |
|
1849 | |
Trigonocephalus affinis Gray (status uncertain, syn. Gloyd & Conant 1990) |
|
| |
Agkistrodon blomhoffii ? affinis Stejneger 1907 |
|
| |
Agkistrodon halys affinis Mell 1929 |
|
| |
Agkistrodon affinis Gloyd & Conant 1990 |
|
1860 | |
Trigonocephalus blomhoffii var. megaspilus Cope (Golay & al. 1993) |
Remarks:
See remarks under halys and brevicaudus.
Older records from Tsushima Island (Japan) refer to tsushimaensis (Gumprecht, Tillack, Orlov, Captain & Ryabov 2004).
Erroneously recorded from Sakhalin Island, Russia (e.g., Gloyd & Conant 1990; David & Ineich 1999) and Taiwan (e.g., Zhao, Jiang & Huang 1979)
(Zhao & Adler 1993; Gumprecht, Tillack, Orlov, Captain & Ryabov 2004).
Distribution:
Japan
(Hokkaido,
Honshu,
Kyushu,
Shikoku,
Izu Islands,
Ryukyu Islands
[incl.
Satsunan Islands]),
Russia
(Kuril Islands).
|
|
|
Gloydius brevicaudus
|
Short-tailed Mamushi, Korean Mamushi
|
|
Kurzschwanz-Mamushi
|
|
Korthalet Mamushi
|
|
1907 | |
Agkistrodon blomhoffii brevicaudus Stejneger |
|
| |
Ancistrodon blomhoffii brevicaudus Nikolsky 1914 |
|
| |
Gloydius blomhoffi brevicaudus Hoge & Romano Hoge 1981 |
|
| |
Gloydius brevicaudus Zhou, Zhang, Huang, Chen, Chen & Yao 2001 |
|
1977 | |
Agkistrodon blomhoffii dubitatus Gloyd (Zhao & Adler 1993; see remarks) |
|
| |
Gloydius blomhoffi dubitatus Hoge & Romano-Hoge 1981 |
|
1977 | |
Agkistrodon blomhoffii siniticus Gloyd (Wallach, Williams & Boundy 2014) |
|
| |
Gloydius blomhoffii siniticus Hoge & Romano Hoge 1981 |
Other common names:
dubitatus: Tung Ling Mamushi, (Northern Chinese Mamushi)
siniticus: Yangtze Mamushi, (Chinese Mamushi)
Remarks:
Previously included cognatus and caraganus.
Formerly regarded as a synonym or subspecies of blomhoffii by some authors (e.g., McDiarmid, Campbell & Touré 1999; Watkins-Colwell & Leenders 2002)
or as a valid species by others (e.g., Vogel 2006; Wallach, Williams & Boundy 2014).
Some authors regard dubitatus as a synonym of blomhoffii (e.g., Wallach, Williams & Boundy 2014).
Herein, dubitatus is assigned to brevicaudus, following Zhao & Adler (1993), thus restricting blomhoffii to Japan.
Gumprecht, Tillack, Orlov, Captain & Ryabov (2004) did not mention dubitatus in the synonymy of blomhoffii.
Distribution:
China
(Anhui,
Beijing,
Chongqing,
Fujian,
Guangdong
[incl.
Nan Ao Island],
Guizhou,
Hebei,
Heilongjiang,
Henan,
Hubei,
Hunan,
Inner Mongolia,
Jiangsu,
Jiangxi,
Jilin,
Liaoning,
Shandong,
Shaanxi,
Sichuan,
Yunnan,
Zhejiang),
North Korea,
South Korea.
|
|
|
Gloydius caraganus
|
Karaganda Mamushi, Karaganda Pitviper, (Siberian Pitviper)
|
|
Karaganda-Mamushi
|
|
Karaganda-mamushi
|
|
1831 | |
Trigonocephalus caraganus Eichwald |
|
| |
Agkistrodon halys caraganus Chernov 1934 |
|
| |
Ancistrodon halys caraganus Terentjev & Chernov 1949 |
|
| |
Gloydius halys caraganus Hoge & Romano Hoge 1981 |
|
| |
Gloydius caraganus Shi, Yang, Zhang, Qi, Li & Ding 2016 |
|
1931 | |
Ancistrodon halys paramonovi Nikolsky (Golay & al. 1993) |
Remarks:
Revalidated as a separate species by Shi, Yang, Zhang, Qi, Li & Ding (2016).
Validity supported by Shi, Wang, Chen, Fang, Ding, Huang, Hou, Liu & Li (2017).
See remarks under halys regarding taxonomic instability of the names caraganus and halys.
Distribution:
Kazakhstan,
Kyrgyzstan,
Mongolia,
Russia,
Tajikistan,
Turkmenistan,
Uzbekistan.
|
|
|
Gloydius caucasicus
|
Caucasian Mamushi, Caucasian Pitviper
|
|
Kaukasische Mamushi
|
|
Kaukasisk Mamushi
|
|
1916 | |
Ancistrodon halys caucasicus Nikolsky |
|
| |
Agkistrodon halys caucasicus Chernov 1934 |
|
| |
Agkistrodon intermedius caucasicus Gloyd & Conant 1982 |
|
| |
Gloydius halys caucasicus Hoge & Romano Hoge 1981 |
|
| |
Gloydius caucasicus Asadi, Montgelard, Nazarizadeh, Moghaddasi, Fatemizadeh, Simonov, Kami & Kaboli 2019 |
Remarks:
Previously regarded as a subspecies of halys (e.g., David & Vogel 2010; Midtgaard 2019).
Distribution:
Afghanistan,
Azerbaijan,
Iran,
Turkmenistan.
|
|
|
Gloydius chambensis
|
Chamba Mamushi, Chamba Pitviper
|
|
Chamba-Mamushi
|
|
Chamba-mamushi
|
|
2022 | |
Gloydius chambensis Kuttalam, Santra, Owens, Selvan, Mukherjee, Graham, Togridou, Bharti, Shi, Shanker & Malhotra |
Distribution:
India
(Himachal Pradesh).
|
|
|
Gloydius changdaoensis
|
Jiaodong Mamushi
|
|
Jiaodong-Mamushi
|
|
Jiaodong-mamushi
|
|
1999 | |
Gloydius saxatilis changdaoensis Li |
|
| |
Gloydius changdaoensis Shi, Wang, Chen, Fang, Ding, Huang, Hou, Liu & Li 2017 |
Distribution:
China
(Jiangsu,
Shandong
[incl.
Daheishan Islands]).
|
|
|
Gloydius cognatus
|
Alashan Mamushi, Alashan Pitviper
|
|
Alashan-Mamushi
|
|
Alashan-mamushi
|
|
1977 | |
Agkistrodon halys cognatus Gloyd (David & Vogel in Visser 2015) |
|
| |
Gloydius halys cognatus Hoge & Romano Hoge 1981 |
|
| |
Gloydius cognatus Shi, Yang, Zhang, Qi, Li & Ding 2016 |
Remarks:
Previously regarded as a synonym of brevicaudus (e.g., David & Vogel in Visser 2015) or a subspecies of halys (e.g., Hoge & Romano Hoge 1981).
Revalidated as a separate species by Shi, Yang, Zhang, Qi, Li & Ding (2016).
Validity supported by Shi, Wang, Chen, Fang, Ding, Huang, Hou, Liu & Li (2017).
Distribution:
China
(Gansu,
Inner Mongolia,
Ningxia,
Qinghai,
Sichuan).
|
|
|
Gloydius halys
|
Central Asian Mamushi, Asiatic Pitviper, Asiatic Moccasin, (Halys' Pitviper, Pallas' Pitviper)
|
|
Halysschlange, Schitomordnik
|
|
Centralasiatisk Mamushi, Halysslange
|
|
1776 | |
Coluber Halys Pallas |
|
| |
Trigonocephalus halys Lichtenstein in Eversman & Lichtenstein 1823 |
|
| |
Vipera halys Lichtenstein in Eversman & Lichtenstein 1823 (as cited in the synonymy of T. halys) |
|
| |
Ancistrodon halys Boulenger 1896 |
|
| |
Agkistrodon halys Stejneger 1907 |
|
| |
Gloydius halys Hoge & Romano Hoge 1981 |
|
1820 | |
Vipera (Echidna) aspis pallasii Merrem (McDiarmid, Campbell & Touré 1999) |
|
1876 | |
Halys pallasii Blanford [not Vipera (Echidna) aspis pallasii Merrem 1820] (Golay & al. 1993) |
|
1933 | |
Ancistrodon halys persicus Rendahl (Wallach, Williams & Boundy 2014) |
|
| |
Agkistrodon halys persicus Golay & al. 1993 |
|
1983 | |
Agkistrodon halys boehmei Nilson |
|
1993 | |
Agkistrodon halys mogoi Bour (Ananjeva, Munkhbayar, Orlov, Orlova, Semenov & Terbisk 1997; Orlov & Barabanov 1999) |
|
| |
Gloydius halys mogoi David & Vogel 2010 |
|
2017 | |
Gloydius halys ubsunurensis Kropachev & Orlov |
Other common names:
boehmei: Afghan Mamushi, Afghan Pitviper, (Böhme’s Pitviper)
halys (incl. mogoi): Mongolian Mamushi, (Siberian Pitviper)
ubsunurensis: Ubsunur Mamushi, Ubsunur Pitviper
Remarks:
Previously included blomhoffi, caraganus, caucasicus, intermedius, and stejnegeri.
Confusion or lack of consensus still prevails regarding the use of the names halys and caraganus.
The latter was considered a distinct species by Shi, Yang, Zhang, Qi, Li & Ding (2016), as already suspected to be by
Gumprecht, Tillack, Orlov, Captain & Ryabov (2004), which is followed herein.
However, Bour (1993), followed by some other authors (e.g., David 2007; David & Vogel 2010), regarded halys as the correct name for the western species referred to
herein as caraganus (above), and he introduced the new name Agkistrodon halys mogoi for the eastern form, referred to herein as Gloydius h. halys.
This taxonomic instability is related to uncertainty regarding the type locality.
Orlov & Barabanov (2000) selected a neotype from Siberia, an action critisized by David (2007) and David & Vogel (2010) with reference to Bour (1993),
who presented evidence that the type was collected north of Astrakhan, near the border between Russia and Kazakhstan.
Pending the establishment of a consensus regard these taxonomic issues, the work of Orlov & Barabanoc (1999) is provisionally followed herein.
Distribution:
Afghanistan,
Azerbaijan,
China
(Inner Mongolia),
Iran,
Kazakhstan,
Kyrgyzstan,
Mongolia,
Russia,
Tajikistan,
Turkmenistan,
Uzbekistan.
|
|
|
Gloydius himalayanus
|
Himalayan Mamushi, Brown Himalayan Viper, (Himalayan Pitviper)
|
|
Himalaya-Mamushi
|
|
Himalaya-mamushi, (Himalayagrubeorm)
|
|
1864 | |
Halys himalayanus Günther |
|
| |
Trigonocephalus himalayanus Strauch 1873 |
|
| |
Crotalus halus himalayanus Higgins 1873 |
|
| |
Ancistrodon himalayanus Boulenger 1890 |
|
| |
Agkistrodon himalayanus Hatta 1928 |
|
| |
Gloydius himalayanus Hoge & Romano Hoge 1981 |
Remarks:
Presence in Sikkim, India, needs confirmation (Gumprecht, Tillack, Orlov, Captain & Ryabov 2004), although likely, since the species is found both to the east and
to the west of the state.
A single specimen has been reported from Afghanistan, but was later reidentified as Macrovipera lebetina (Wagner, Bauer, Leviton, Wilms & Böhme 2016).
Not mentioned for Punjab by Singh, Dutta & Singh (2021).
Records from Bangladesh (e.g., Sarker & Sarker 1985) are erroneous (Gumprecht, Tillack, Orlov, Captain & Ryabov 2004).
Distribution:
Bhutan,
India
(Haryana,
Himachal Pradesh,
Jammu & Kashmir,
Punjab,
Uttar Pradesh,
Uttarakhand,
West Bengal),
Nepal,
Pakistan.
|
|
|
Gloydius huangi
|
Lancang Mamushi, Lancang Plateau Viper
|
|
Lancang-Mamushi
|
|
Lancang-mamushi
|
|
2019 | |
Gloydius huangi Wang, Ren, Dong, Jiang, Shi, Siler & Che |
Distribution:
China
(Tibet).
|
|
|
Gloydius intermedius
|
Siberian Mamushi, (Central Asian Pitviper, Amur Pitviper)
|
|
Zentralasiatische Mamushi
|
|
Siberisk Mamushi
|
|
1868 | |
Trigonocephalus intermedius Strauch |
|
| |
Halys intermedius Peters 1877 |
|
| |
Ancistrodon intermedius Boulenger 1896 |
|
| |
Agkistrodon blomhoffii intermedius Stejneger 1907 |
|
| |
Ancistrodon blomhoffii intermedius Despax 1913 |
|
| |
Ancistrodon halys intermedius Nikolsky 1916 |
|
| |
Agkistrodon halys intermedius Stejneger 1925 |
|
| |
Agkistrodon intermedius Werner 1929 |
|
| |
Gloydius halys intermedius Hoge & Romano Hoge 1981 |
|
| |
Gloydius intermedius Kraus, Mink & Brown 1996 |
Remarks:
Previously included stejnegeri.
Formerly regarded as a subspecies of blomhoffii (e.g., Stejneger 1907), or of halys (e.g., Nikolsky 1916).
Some authors have considered saxatilis a synonym of intermedius.
Distribution:
China
(Gansu,
Heilongjiang,
Inner Mongolia,
Ningxia,
Shaanxi,
Shanxi,
Xinjiang),
Russia.
|
|
|
Gloydius lateralis
|
Jiuzhai Mamushi, Jiuzhai Pitviper
|
|
Jiuzhai-Mamushi
|
|
Jiuzhai-mamushi
|
|
2022 | |
Gloydius lateralis Zhang, Shi, Jiang & Shi in Zhang, Shi, Li, Yan, Wang, Ding, Du, Plenkovic-Moraj, Jiang & Shi |
Distribution:
China
(Sichuan).
|
|
|
Gloydius lijianlii
|
Shandong Mamushi
|
|
Shandong-Mamushi
|
|
Shandong-mamushi
|
|
2009 | |
Gloydius lijianlii Jiang & Zhao |
Distribution:
China
(Shandong:
Daheishan Islands).
|
|
|
Gloydius lipipengi
|
Nujiang Mamushi, Nujiang Pitviper
|
|
Nujiang-Mamushi
|
|
Nujiang-mamushi
|
|
2021 | |
Gloydius lipipengi Shi, Liu & Malhotra in Shi, Liu, Giri, Owens, Santra, Kuttalam, Selvan, Guo & Malhotra |
Distribution:
China
(Tibet).
|
|
|
Gloydius liupanensis
|
Liupan Mamushi
|
|
Liupan-Mamushi
|
|
Liupan-mamushi
|
|
1989 | |
Agkistrodon halys liupanensis Liu, Song & Luo |
|
| |
Gloydius liupanensis Xu, Liu, Myers, Wang, Huang, He, Peng & Guo 2012 |
Distribution:
China
(Gansu,
Ningxia).
|
|
|
Gloydius monticola
|
Likiang Mamushi, Likiang Pitviper, Mountain Mamushi, Yunnan Mamushi
|
|
Likiang-Mamushi
|
|
Likiang-mamushi
|
|
1922 | |
Ancistrodon blomhoffii monticola Werner |
|
| |
Agkistrodon halys monticola Gee 1929 |
|
| |
Agkistrodon monticola Mell 1929 |
|
| |
Ancistrodon halys monticola Rendahl 1933 |
|
| |
Agkistrodon blomhoffi monticola Mertens 1934 |
|
| |
Gloydius monticola Hoge & Romano Hoge 1981 |
|
| |
Gloydius strauchi monticola David & Ineich 1999 |
|
1931 | |
Agkistrodon strauchi popei Mell "1929" (Golay & al. 1993; Zhao & Adler 1993) |
Remarks:
Considered a synonym of strauchi by Zhao (1995), but as a valid species in most recent publications (e.g., McDiarmid, Campbell & Touré 1999, Orlov, Ananjeva, Barabanov, Ryabov & Kha-Likov 2002, Gumprecht, Tillack, Orlov, Captain & Ryabov 2004, Vogel 2006).
Distribution:
China
(Yunnan).
|
|
|
Gloydius qinlingensis
|
Shaanxi Mamushi
|
|
Shaanxi-Mamushi
|
|
Shaanxi-mamushi
|
|
1985 | |
Agkistrodon halys qinlingensis Song & Chen |
|
| |
Gloydius qinlingensis Guo, Zhang & Chen 1999 |
Distribution:
China
(Shaanxi).
|
|
|
Gloydius rickmersi
|
Kyrgyz Mamushi
|
|
Kirgisische Mamushi
|
|
Kirgisisk Mamushi
|
|
2015 | |
Gloydius rickmersi Wagner, Tiutenko, Borkin & Simonov |
Remarks:
Validity supported by Shi, Wang, Chen, Fang, Ding, Huang, Hou, Liu & Li (2017).
Distribution:
Kyrgyzstan.
|
|
|
Gloydius rubromaculatus
|
Red-spotted Mamushi, Red-spotted Alpine Pitviper, Tongtianhe Pitviper
|
|
Rotfleckige Mamushi
|
|
Rødplettet Mamushi
|
|
2017 | |
Gloydius rubromaculatus Shi, Wang, Chen, Fang, Ding, Huang, Hou, Liu & Li |
Distribution:
China
(Qinghai,
Sichuan,
Tibet).
|
|
|
Gloydius saxatilis
|
Rock Mamushi, Brown Mamushi
|
|
Felsen-Mamushi
|
|
Klippe-mamushi
|
|
1937 | |
Ancistrodon saxatilis Emelianov |
|
| |
Agkistrodon saxatilis Gloyd 1972 |
|
| |
Agkistrodon intermedius saxatilis Gloyd & Conant 1982 |
|
| |
Gloydius saxatilis Hoge & Romano Hoge 1981 |
|
1980 | |
Agkistrodon shedaoensis continentalis Zhao (Golay & al. 1993) |
Remarks:
Previously included changdaoensis.
Regarded by some as a synonym or subspecies of intermedius
(e.g., Gloyd & Conant 1982, 1990; Orlov & Barabanov 2000; Gumprecht, Tillack, Orlov, Captain & Ryabov 2004; David & Vogel in Visser 2015; Dunayev & Orlova 2023).
Treated herein as a valid species, following
(e.g., Zhao & Adler 1993; Golay & al. 1993; McDiarmid, Campbell & Touré 1999; Xu, Liu, Myers, Wang, Huang, He, Peng & Guo 2012; Wallach, Williams & Boundy 2014).
Not listed for China by Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che (2020).
Distribution:
China
(Heilongjiang,
Inner Mongolia,
Jiangsu,
Jilin,
Liaoning,
Shandong
[incl.
Daheishan Islands]).
North Korea,
Russia,
South Korea.
|
Gloydius saxatilis
© Rune Midtgaard
|
|
Gloydius shedaoensis
|
Shedao Island Mamushi, Shedao Island Pitviper
|
|
Shedao-Mamushi
|
|
Shedao-mamushi
|
|
1979 | |
Agkistrodon shedaoensis Zhao |
|
| |
Agkistrodon intermedius shedaoensis Welch 1994 |
|
| |
Gloydius shedaoensis McDiarmid, Campbell & Touré 1999 |
|
| |
Gloydius saxatilis shedaoensis Zhou, Zhang, Huang, Chen, Chen & Yao 2001 |
|
1999 | |
Gloydius shedaoensis qianshanensis Li |
Distribution:
China
(Liaoning [Shedao Island and mainland]).
|
|
|
Gloydius stejnegeri
|
Gobi Mamushi, Gobi Pitviper, (Northern Chinese Pitviper)
|
|
Gobi-Mamushi
|
|
Gobi-mamushi
|
|
1933 | |
Ancistrodon halys stejnegeri Rendahl |
|
| |
Agkistrodon intermedius stejnegeri Gloyd & Conant 1982 |
|
| |
Gloydius halys stejnegeri Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 |
|
| |
Gloydius stejnegeri Shi, Yang, Zhang, Qi, Li & Ding 2016 |
Remarks:
Previously regarded as a synonym of intermedius (e.g., David & Vogel 2010; David & Vogel in Visser 2015).
Revalidated as a separate species by Shi, Yang, Zhang, Qi, Li & Ding (2016).
Validity supported by Shi, Wang, Chen, Fang, Ding, Huang, Hou, Liu & Li (2017).
Distribution:
China
(Beijing,
Hebei,
Inner Mongolia,
Shaanxi,
Shanxi).
|
|
|
Gloydius strauchi
|
Sichuan Mamushi (Tibetan Mamushi, Strauch’s Pitviper, Strauch's Mamushi)
|
|
Sichuan-Mamushi, (Strauchs Grubenotter)
|
|
Sichuan-mamushi, (Tibetansk Mamushi)
|
|
1912 | |
Ancistrodon Strauchi Bedriaga |
|
| |
Agkistrodon strauchi Stejneger 1925 |
|
| |
Ancistrodon halys strauchi Pavlov 1932 |
|
| |
Gloydius strauchi Hoge & Romano Hoge 1981 |
|
1912 | |
Agkistrodon tibetanus Barbour (Golay & al. 1993; Zhao & Adler 1993) |
|
| |
Ancistrodon tibetanus Werner 1922 |
Remarks:
Currently considered restricted to Sichuan, China.
Previously included qinlingensis and liupanensis, as well as the populations now assigned to angusticeps and rubromaculatus
(Shi, Yang, Zhang, Peng, Orlov, Jiang, Ding, Hou, Huang, Huang & Li 2018).
Exact western limit of distribution is unclear, since some records may refer to rubromaculatus or angusticeps
(Shi, Wang, Chen, Fang, Ding, Huang, Hou, Liu & Li 2017; Shi, Yang, Zhang, Peng, Orlov, Jiang, Ding, Hou, Huang, Huang & Li 2018).
A single report from Yunnan (He & Zhou 2002) is regarded as doubtful.
Distribution:
China
(Sichuan).
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Gloydius swild
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Glacier Mamushi, Glacier Pitviper
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Gletcher-Mamushi
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Gletcher-mamushi
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2021 | |
Gloydius swild Shi & Malhotra in Shi, Liu, Giri, Owens, Santra, Kuttalam, Selvan, Guo & Malhotra |
Distribution:
China
(Sichuan).
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Gloydius tsushimaensis
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Tsushima Mamushi, Tsushima Island Pitviper
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Tsushima-Mamushi
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Tsushima-mamushi
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1994 | |
Agkistrodon tsushimaensis Isogawa, Moriya & Mitsui |
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Gloydius tsushimaensis Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 |
Distribution:
Japan
(Tsushima Islands).
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Gloydius ussuriensis
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Ussuri Mamushi, Ussurian Pitviper, (Amur Mamushi)
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Ussuri-Mamushi
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Ussuri-mamushi
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1929 | |
Ancistrodon blomhoffii ussuriensis Emelianov |
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Agkistrodon blomhoffii ussuriensis Chernov 1934 |
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Agkistrodon halys ussuriensis Klemmer 1963 |
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Gloydius halys ussuriensis Hoge & Romano Hoge 1981 |
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Agkistrodon ussuriensis Toriba 1986 |
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Gloydius ussuriensis Vogel 2006 |
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1972 | |
Agkistrodon caliginosus Gloyd (Toriba 1988; McDiarmid, Campbell & Touré 1999; Gumprecht, Tillack, Orlov, Captain & Ryabov 2004) |
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Gloydius caliginosus Hoge & Romano Hoge 1981 |
Other common names:
caliginosus: Dusky Mamushi, Caliginous Mamushi, Kankoku Mamushi
Distribution:
China
(Heilongjiang,
Inner Mongolia,
Jilin,
Liaoning),
Mongolia,
North Korea,
Russia,
South Korea.
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