Reptiles of the Ryukyu Islands  (=Nansei Islands) [Japan] Related checklists:  Japan - Amami Islands - Okinawa Islands - Osumi Islands - Sakishima Islands - Tokara Islands Bibliography of the reptiles of Ryukyu Islands Taxonomic history of the reptiles of Ryukyu Islands Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus porosus Saltwater Crocodile Spennemann 2021

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Diploderma (Eastern Mountain Agamas)   Diploderma polygonatum Japanese Mountain Agama Goris & Maeda 2005 Johnson 1969 Manthey 2010 Ota 1991 Ota 2003 Stejneger 1907 Wang, Gao, Wu, Dong, Feng, Shen, Jin, Shi, Qi, Siler & Che 2021 Wang, Ren, Wu, Jiang, Jin, Hou, Zheng, Xie, Siler & Che 2021 Wermuth 1967  Family Eublepharidae (Eyelid Geckos) Genus Goniurosaurus (East Asian Leopard Geckos)   * Goniurosaurus kuroiwae Okinawa Leopard Gecko Goris & Maeda 2005 Grismer, Viets & Boyle 1999 Nakamura, Takahashi & Ota 2014 Seufer, Kaverkin & Kirschner (eds.) 2005   * Goniurosaurus orientalis Tonaki Leopard Gecko Goris & Maeda 2005 Grismer, Viets & Boyle 1999 Honda & Ota 2017 Johnson 1969 Seufer, Kaverkin & Kirschner (eds.) 2005   * Goniurosaurus splendens Tokuno Leopard Gecko Goris & Maeda 2005 Grismer, Viets & Boyle 1999 Seufer, Kaverkin & Kirschner (eds.) 2005   * Goniurosaurus toyamai Iheya Leopard Gecko Goris & Maeda 2005 Grismer, Viets & Boyle 1999 Seufer, Kaverkin & Kirschner (eds.) 2005   * Goniurosaurus yamashinae Kume Leopard Gecko Goris & Maeda 2005 Grismer, Viets & Boyle 1999 Seufer, Kaverkin & Kirschner (eds.) 2005  Family Gekkonidae (Typical Geckos) Genus Gehyra (Four-clawed Geckos)   * Gehyra intermedia Ryukyu Stump-toed Gecko Brown 1902 Wermuth 1965   Gehyra mutilata Common Four-clawed Gecko Goris & Maeda 2005 Stejneger 1907 Zhao & Adler 1993 Genus Gekko (Tokays)   Gekko hokouensis Hokou Gecko Goris & Maeda 2005 Ota, Sakaguchi, Ikehara & Hikida 1993 Zhao & Adler 1993   Gekko japonicus Japanese Giant Gecko Goris & Maeda 2005 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Stejneger 1907 Wermuth 1965   * Gekko shibatai Tokara Narrow-disked Gecko Toda, Sengoku, Hikida & Ota 2008   * Gekko vertebralis Spine-patterned Gecko Toda, Sengoku, Hikida & Ota 2008   Gekko yakuensis Yaku Gecko Goris & Maeda 2005 Matsui & Okada 1968 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus bowringii Hong Kong House Gecko Goris & Maeda 2005 McMahan & Zug 2007 Wermuth 1965 Zhao & Adler 1993   * Hemidactylus marmoratus Riu Kiu Half-toed Gecko Stejneger 1907 Wermuth 1965 Genus Hemiphyllodactylus (Slender Geckos)   Hemiphyllodactylus typus Indo-Pacific Slender Gecko Goris & Maeda 2005 Ota 1990 Zug 2010 Genus Lepidodactylus (Scaly-toed Geckos)   Lepidodactylus lugubris Common Mourning Gecko Goris & Maeda 2005 Ota, Fischer, Ineich & Case 1995 Zhao & Adler 1993  Family Lacertidae (Typical Lizards) Genus Takydromus (Oriental Grass Lizards)   * Takydromus dorsalis Sakishima Grass Lizard Arnold 1997 Bischoff 1992 Goris & Maeda 2005 Stejneger 1907   * Takydromus smaragdinus Emerald Grass Lizard Arnold 1997 Bischoff 1992 Goris & Maeda 2005 Johnson 1969 Stejneger 1907 Takeda & Ota 1996   Takydromus tachydromoides Japanese Grass Lizard Goris & Maeda 2005   * Takydromus toyamai Miyako Grass Lizard Arnold 1997 Goris & Maeda 2005 Takeda & Ota 1996  Family Scincidae (Skinks) Genus Ateuchosaurus (East Asian Short-legged Skinks)   * Ateuchosaurus okinavensis Okinawa Short-legged Skink Goris & Maeda 2005 [as pellopleurus] Johnson 1969 [as p. pellopleurus] Makino, Nakano, Okamoto & Hikida 2023 Thompson 1912   * Ateuchosaurus pellopleurus Amami Short-legged Skink Goris & Maeda 2005 Johnson 1969 Makino, Nakano, Okamoto & Hikida 2023 Stejneger 1907 Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Brown 1991 Goris & Maeda 2005 Zhao & Adler 1993 Genus Plestiodon (Blue-tailed Skinks)   * Plestiodon barbouri Amami Islands Blue-tailed Skink Goris & Maeda 2005 Kato, Ota & Hikida 1994 Taylor 1935   Plestiodon elegans Elegant Five-lined Skink Kato, Ota & Hikida 1994 Ota, Sakaguchi, Ikehara & Hikida 1993 Zhao & Adler 1993   Plestiodon japonicus Japanese Five-lined Skink Goris & Maeda 2005 Kato, Ota & Hikida 1994 Okamoto & Hikida 2012   * Plestiodon kishinouyei Ryukyu Islands Giant Skink Goris & Maeda 2005 Stejneger 1907   * Plestiodon kuchinoshimensis Kuchinoshima Five-lined Skink Kurita & Hikida 2014   * Plestiodon marginatus Ryukyu Five-lined Skink Goris & Maeda 2005 Johnson 1969 Kato, Ota & Hikida 1994 Motokawa, Toyama & Hikida 2001 Taylor 1935   * Plestiodon stimpsonii Yaeyama Seven-lined Skink Goris & Maeda 2005 Kato, Ota & Hikida 1994 Taylor 1935   * Plestiodon takarai Senkaku Skink Goris & Maeda 2005 [as elegans] Kurita, Ota & Hikida 2017 Genus Scincella (South Asian and American Ground Skinks)   * Scincella boettgeri Sakishima Ground Skink Goris & Maeda 2005   * Scincella dunan Yonagunijima Ground Skink Koizumi, Ota & Hikida 2022

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Calamaria (Oriental Reed Snakes)   Calamaria pavimentata Brown Reed Snake Goris & Maeda 2005 Inger & Marx 1965 Zhao & Adler 1993   * Calamaria pfefferi Ryukyu Reed Snake Goris & Maeda 2005 Koch, Arida, McGuire, Iskandar & Boehme 2009 Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe carinata East Asian Keeled Rat Snake Goris & Maeda 2005 Ota, Sakaguchi, Ikehara & Hikida 1993 Schulz 1996   Elaphe climacophora Japanese Green Rat Snake Goris & Maeda 2005   Elaphe quadrivirgata Japanese Four-lined Rat Snake Goris & Maeda 2005   Elaphe taeniura Stripe-tailed Rat Snake Goris & Maeda 2005 Schulz 1996 Stejneger 1907 Genus Euprepiophis (East Asian Rat Snakes)   Euprepiophis conspicillatus Japanese Forest Rat Snake Goris & Maeda 2005 Genus Hebius (Oriental Equatorial Keelbacks)   * Hebius concelarus Miyako Islands Keelback Lalremsanga, Bal, Vogel & Biakzuala 2022 Malnate & Munsterman 1960 [as Natrix p. pryeri] Malnate 1963 Ota & Iwanaga 1997 Wallach, Williams & Boundy 2014   * Hebius ishigakiensis Yaeyama Water Snake Goris & Maeda 2005 Lalremsanga, Bal, Vogel & Biakzuala 2022 Malnate & Munsterman 1960 Malnate 1962 Ota & Iwanaga 1997 Wallach, Williams & Boundy 2014   * Hebius pryeri Ryukyu Keelback Goris & Maeda 2005 Johnson 1969 Malnate & Munsterman 1960 Malnate 1962 Ota & Iwanaga 1997 Wallach, Williams & Boundy 2014   Hebius vibakari Northeastern Forest Keelback Goris & Maeda 2005 Genus Lycodon (Oriental Wolf Snakes)   * Lycodon multifasciatus Ryukyu Mountain Wolf Snake Goris & Maeda 2005 Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler 2009 Vogel, Nguyen, Kingsada & Ziegler 2012   Lycodon orientalis East Asian Wolf Snake Goris & Maeda 2005   Lycodon rufozonatus Red-banded Big-toothed Snake Goris & Maeda 2005 Ota, Sakaguchi, Ikehara & Hikida 1993 Stejneger 1907 Zhao & Adler 1993   * Lycodon semicarinatus Tricolored Banded Snake Goris & Maeda 2005 Johnson 1969 Wang, Qi, Lyu, Zeng & Wang 2020 Genus Opisthotropis (Mountain Stream Keelbacks)   * Opisthotropis kikuzatoi Ryukyu Mountain Stream Keelback Goris & Maeda 2005 Ota 2004 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   * Ptyas herminae Sakishima Green Snake Goris & Maeda 2005 Kidera & Ota 2017 Stejneger 1907   * Ptyas semicarinata Ryukyu Green Snake Goris & Maeda 2005 Johnson 1969 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis tigrinus Japanese Tiger Keelback Goris & Maeda 2005  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Sinomicrurus (East Asian Coral Snakes)   * Sinomicrurus boettgeri Okinawa Coral Snake Goris & Maeda 2005 Ota, Ito & Lin 1999 Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021   * Sinomicrurus iwasakii Ishigaki Coral Snake Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Goris & Maeda 2005 Maki 1935 Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021 Zhao & Adler 1993 [as macclellandii]   * Sinomicrurus japonicus Ryukyu Coral Snake Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Goris & Maeda 2005 Ota, Ito & Lin 1999 Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021 Stejneger 1907  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Emydocephalus (Turtle-headed Sea Snakes)   Emydocephalus ijimae Japanese Turtle-headed Sea Snake Cogger in Dunson (ed.) 1975 Goris & Maeda 2005 Smith 1926 Stejneger 1907 Zhao & Adler 1993 Genus Hydrophis (Typical Sea Snakes)   Hydrophis cyanocinctus Blue-banded Sea Snake Goris & Maeda 2005 Stejneger 1907   Hydrophis melanocephalus Black-headed Slender-necked Sea Snake Goris & Maeda 2005 Smith 1926 Zhao & Adler 1993   Hydrophis ornatus Reef Sea Snake Goris & Maeda 2005 Haas 1950 Smith 1926 Toriba & Nakamoto 1991 Zhao & Adler 1993   Hydrophis platurus Yellow-bellied Sea Snake Goris & Maeda 2005 Ineich 1988 Smith 1926   Hydrophis stokesii Large-headed Sea Snake Sasai, Yamamoto, Oka & Toda 2021 Genus Laticauda (Sea Kraits)   Laticauda colubrina Yellow-lipped Sea Krait Goris & Maeda 2005 Heatwole, Busack & Cogger 2005 Kharin, Roedel & Hallermann 2010 McCarthy 1986 Ota, Takahashi & Kamezaki 1985 Zhao & Adler 1993   Laticauda laticaudata Blue-lipped Sea Krait Goris & Maeda 2005 McCarthy 1986 Smith 1926 Stejneger 1907   Laticauda semifasciata Erabu Sea Krait Goris & Maeda 2005 Kharin, Roedel & Hallermann 2010 McCarthy 1986 Smith 1926 Stejneger 1907  Family Pareidae (Oriental Slug Eaters) Genus Pareas (Typical Oriental Slug Eaters)   * Pareas iwasakii Ryukyu Slug Eater Ding, Chen, Suwannapoom, Nguyen, Pogarkov & Vogel 2020 Goris & Maeda 2005 Vogel, Nguyen, Lalremsanga, Biakzuala, Hrima & Poyarkov 2020 Yang, Yeung, Huang & Yang 2021  Family Viperidae (Vipers) Genus Gloydius (Mamushis)   Gloydius blomhoffii Japanese Mamushi Gloyd & Conant 1990 Goris & Maeda 2005 Genus Ovophis (Asian Mountain Pitvipers)   * Ovophis okinavensis Kufah Goris & Maeda 2005 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Johnson 1969 Stejneger 1907 Genus Protobothrops (Habus and allies)   * Protobothrops elegans Elegant Pitviper Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Goris & Maeda 2005 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 McDiarmid, Campbell & Toure 1999 Stejneger 1907   * Protobothrops flavoviridis Common Habu Goris & Maeda 2005 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Stejneger 1907   * Protobothrops tokarensis Tokara Habu Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Vogel 2006  Family Xenodermidae (Asian Litter Snakes) Genus Achalinus (East Asian Burrowing Snakes)   Achalinus formosanus Taiwanese Burrowing Snake Goris & Maeda 2005 Ota & Toyama 1989 Wallach, Williams & Boundy 2014 Zhao & Adler 1993   * Achalinus werneri Amami Burrowing Snake Goris & Maeda 2005 Johnson 1969 Wallach, Williams & Boundy 2014

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Goris & Maeda 2005 Uchida & Nishiwaki in Bjorndal (ed.) 1982 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Goris & Maeda 2005 Uchida & Nishiwaki in Bjorndal (ed.) 1982 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Goris & Maeda 2005 Pritchard 1979 Uchida & Nishiwaki in Bjorndal (ed.) 1982  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   * Cuora evelynae Riu Kiu Yellow-margined Box Turtle Ernst & Lovich 1990 Goris & Maeda 2005 Iverson 1992 Ota, Yasukawa, Fu & Chen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wermuth & Mertens 1977 [as flavomarginata] Zhao & Adler 1993 [as flavomarginata] Genus Geoemyda (Black-breasted Leaf Turtles)   * Geoemyda japonica Ryukyu Black-breasted Leaf Turtle Fritz & Havas 2006 Goris & Maeda 2005 Iverson 1992 Johnson 1969 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wermuth & Mertens 1977 Yasukawa & Ota 2008 Genus Mauremys (Stripe-necked Turtles)   Mauremys mutica Yellow Pond Turtle Goris & Maeda 2005 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021