Reptiles of Zhejiang [China] Related checklists:  China Bibliography of the reptiles of Zhejiang Taxonomic history of the reptiles of Zhejiang Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Alligatoridae (Alligators and Caimans) Genus Alligator (Alligators)   Alligator sinensis Chinese Alligator IUCN Crocodile Specialist Group 1996 Zhao & Adler 1993

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Anguidae (Slowworms, etc.) Genus Dopasia (Oriental Glass Lizards)   Dopasia harti South Chinese Glass Lizard Brygoo 1987 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993  Family Gekkonidae (Typical Geckos) Genus Gekko (Tokays)   Gekko hokouensis Hokou Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Zhao & Adler 1993   Gekko japonicus Japanese Giant Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Zhao & Adler 1993   Gekko subpalmatus Eastern Chinese Palmated Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Lyu, Lin, Ren, Jiang, Zhang, Qi & Wang 2021 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Zhao & Adler 1993   Gekko swinhonis Beijing Gecko Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Zhao & Adler 1993  Family Lacertidae (Typical Lizards) Genus Takydromus (Oriental Grass Lizards)   Takydromus septentrionalis Northern Grass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Cai, Yan, Xu, Lin & Ji 2012 Wang, Lyu, Yang, Li & Wang 2019 Zhao & Adler 1993   Takydromus sexlineatus Six-lined Grass Lizard Zhang 2002   Takydromus sylvaticus Fujian Grass Lizard Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021  Family Scincidae (Skinks) Genus Plestiodon (Blue-tailed Skinks)   Plestiodon chinensis Eastern Chinese Long Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993   Plestiodon elegans Elegant Five-lined Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993   Plestiodon liui Jiangsu Blue-tailed Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Hikida & Zhao 1989 Zhao & Adler 1993 Genus Scincella (South Asian and American Ground Skinks)   Scincella modesta Slender Ground Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus incognitus Koshun Forest Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021   Sphenomorphus indicus Common Indian Forest Skink Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Zhao & Adler 1993

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Amphiesma (South Asian Keelbacks)   Amphiesma stolatum Buff-striped Keelback Lalremsanga, Bal, Vogel & Biakzuala 2022 Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Genus Boiga (Australasian Cat Snakes)   Boiga kraepelini Kelung Cat Snake Zhao & Adler 1993   Boiga multomaculata Marbled Cat Snake Zhao & Adler 1993 Genus Calamaria (Oriental Reed Snakes)   Calamaria pavimentata Brown Reed Snake Zhao & Adler 1993   Calamaria septentrionalis Northern Reed Snake Inger & Marx 1965 Zhao & Adler 1993 Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe bimaculata Twin-spotted Rat Snake Schulz 1996 Zhao & Adler 1993   Elaphe carinata East Asian Keeled Rat Snake Qi, Shi, Ma, Gao, Bu, Grismer, Li & Wang 2021 Schulz 1996 Zhao & Adler 1993   Elaphe schrenckii Amur Rat Snake Schulz 1996 Zhao & Adler 1993   Elaphe taeniura Stripe-tailed Rat Snake Schulz 1996 Zhao & Adler 1993 Genus Euprepiophis (East Asian Rat Snakes)   Euprepiophis mandarinus Mandarin Rat Snake Schulz 1996 Zhao & Adler 1993 Genus Fowlea (Fishing Keelbacks)   Fowlea flavipunctata Yellow-spotted Keelback Vogel & David 2012 Zhao & Adler 1993 [as piscator] Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma frenatum Khasi Hills Rat Snake Schulz 1996 Zhao & Adler 1993 Genus Hebius (Oriental Equatorial Keelbacks)   Hebius craspedogaster Kuatun Keelback Malnate 1962 Wallach, Williams & Boundy 2014 Zhao & Adler 1993 Genus Lycodon (Oriental Wolf Snakes)   Lycodon flavozonatus Yellow Big-toothed Snake Wang, Qi, Lyu, Zeng & Wang 2020 Zhao & Adler 1993   Lycodon rufozonatus Red-banded Big-toothed Snake Wang, Qi, Lyu, Zeng & Wang 2020 Zhao & Adler 1993   Lycodon ruhstrati Taiwanese Wolf Snake Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler 2009 Genus Oligodon (Kukri Snakes)   Oligodon chinensis Chinese Kukri Snake Zhao & Adler 1993   Oligodon formosanus Taiwanese Kukri Snake David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012 Zhao & Adler 1993   Oligodon ornatus Chinese Ornate Kukri Snake Zhao & Adler 1993 Genus Oocatochus (Red-backed Rat Snake)   Oocatochus rufodorsatus Red-backed Rat Snake Schulz 1996 Zhao & Adler 1993 Genus Opisthotropis (Mountain Stream Keelbacks)   Opisthotropis kuatunensis Striped Mountain Stream Keelback Zhao & Adler 1993   Opisthotropis latouchii South Chinese Mountain Stream Keelback Ren, Wang, Guo, Wang, Nguyen & Li 2019 Zhao & Adler 1993   Opisthotropis maxwelli Fujian Mountain Stream Keelback Zhang 2002 Genus Oreocryptophis (Red Mountain Rat Snake)   Oreocryptophis porphyraceus Red Mountain Rat Snake Schulz & Helfenberger 1998 Schulz 1996 Zhao & Adler 1993 Genus Plagiopholis (Blotch-necked Snakes)   Plagiopholis styani Fujian Blotch-necked Snake Zhao & Adler 1993 Genus Pseudagkistrodon (False Habu)   Pseudagkistrodon rudis False Habu Zhao & Adler 1993 Genus Pseudoxenodon (Mock Cobras)   Pseudoxenodon bambusicola Bamboo Mock Cobra Zhao & Adler 1993   Pseudoxenodon stejnegeri Chinese Large-eyed Mock Cobra Zhao & Adler 1993 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas dhumnades Black-striped Rat Snake Zhao & Adler 1993   Ptyas korros Indochinese Rat Snake Zhao & Adler 1993   Ptyas major Greater Green Snake Zhao & Adler 1993   Ptyas mucosa Common Oriental Rat Snake Zhao & Adler 1993 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis lateralis Chinese Tiger Keelback Zhao & Adler 1993 Zhu, Wang, Takeuchi & Zhao 2014 [as R. tigrinus] Genus Sibynophis (Oriental Black-headed Snakes)   Sibynophis chinensis Chinese Black-headed Snake Zhao & Adler 1993 Genus Trimerodytes (Asian Water Snakes)   Trimerodytes aequifasciatus Equal-banded Water Snake Zhao & Adler 1993   Trimerodytes annularis Asian Banded Water Snake Ren, Wang, Guo, Wang, Nguyen & Li 2019 Zhao & Adler 1993   Trimerodytes percarinatus Chinese Keelback Guo, Zhu, Liu, Wang, Che & Nguyen 2020 Zhao & Adler 1993  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Bungarus (Kraits)   Bungarus multicinctus Many-banded Krait Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Zhao & Adler 1993 Genus Naja (Typical Cobras)   Naja atra Chinese Spitting Cobra Shi, Vogel, Ding, Rao, Liu, Zhang, Wu & Chen 2022 Zhao & Adler 1993 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Zhao & Adler 1993 Genus Sinomicrurus (East Asian Coral Snakes)   Sinomicrurus annularis Hong Kong Coral Snake Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021   Sinomicrurus kelloggi Indochinese Coral Snake Zhao & Adler 1993  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis cyanocinctus Blue-banded Sea Snake Zhao & Adler 1993   Hydrophis melanocephalus Black-headed Slender-necked Sea Snake Zhao & Adler 1993   Hydrophis platurus Yellow-bellied Sea Snake Zhao & Adler 1993  Family Homalopsidae (Australasian Mud Snakes) Genus Hypsiscopus (Orange-bellied Mud Snake and Matanna Mud Snake)   Hypsiscopus murphyi Northern Orange-bellied Mud Snake Bernstein, Voris, Stuart, Phimmachak, Seateun, Sivongxay, Neang, Karns, Andrews, Osterhage, Phipps & Ruane 2022 Zhao & Adler 1993 [as plumbea] Genus Myrrophis (Indochinese Mud Snakes)   Myrrophis chinensis Chinese Mud Snake Zhao & Adler 1993  Family Pareidae (Oriental Slug Eaters) Genus Pareas (Typical Oriental Slug Eaters)   Pareas boulengeri Guizhou Slug Eater Ding, Chen, Suwannapoom, Nguyen, Pogarkov & Vogel 2020 Vogel, Nguyen, Zaw & Poyarkov 2020 Wogan & Vogel 2012   Pareas chinensis Chinese Slug Eater Zhao & Adler 1993   Pareas formosensis Taiwanese Red-eyed Slug Eater Le, Tran, Hoang & Stuart 2021   Pareas stanleyi Fujian Slug Eater Zhao & Adler 1993  Family Typhlopidae (Typical Blind Snakes) Genus Virgotyphlops (Brahminy Blind Snake)   Virgotyphlops braminus Brahminy Blind Snake Zhao & Adler 1993  Family Viperidae (Vipers) Genus Azemiops (Asian Mole Vipers)   Azemiops kharini White-headed Mole Viper Orlov, Ryabov & Nguyen 2013 Vogel 2006 [as feae] Zhao & Adler 1993 [as feae] Genus Deinagkistrodon (Sharp-nosed Pitviper)   Deinagkistrodon acutus Sharp-nosed Pitviper Gloyd & Conant 1990 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Zhao & Adler 1993 Genus Gloydius (Mamushis)   Gloydius brevicaudus Short-tailed Mamushi Gloyd & Conant 1990 Vogel 2006 Zhao & Adler 1993 Genus Ovophis (Asian Mountain Pitvipers)   Ovophis makazayazaya Eastern Mountain Pitviper Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Hoge & Romano Hoge 1981 [as monticola orientalis] Zhao & Adler 1993 [as monticola] Genus Protobothrops (Habus and allies)   Protobothrops mucrosquamatus Brown-spotted Pitviper Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Vogel 2006 Zhao & Adler 1993 Genus Viridovipera (Green Tree Pitvipers)   Viridovipera stejnegeri Chinese Green Tree Pitviper David, Vidal & Pauwels 2001 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Vogel 2006 Zhao & Adler 1993  Family Xenodermidae (Asian Litter Snakes) Genus Achalinus (East Asian Burrowing Snakes)   Achalinus rufescens Rufous Burrowing Snake Zhao & Adler 1993   Achalinus spinalis Grey Burrowing Snake Wallach, Williams & Boundy 2014 Zhao & Adler 1993  Family Xenopeltidae (Asian Sunbeam Snakes) Genus Xenopeltis (Asian Sunbeam Snakes)   Xenopeltis hainanensis Hainan Sunbeam Snake McDiarmid, Campbell & Toure 1999 Orlov 2000 Zhao & Adler 1993 Zhao 1997

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Huang in Bjorndal (ed.) 1982 Zhao & Adler 1993 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Huang in Bjorndal (ed.) 1982 Zhao & Adler 1993 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Huang in Bjorndal (ed.) 1982 Zhao & Adler 1993 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Zhao & Adler 1993  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Huang in Bjorndal (ed.) 1982 Zhao & Adler 1993  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora aurocapitata Yellow-headed Box Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Cuora flavomarginata Chinese Yellow-margined Box Turtle Lau & Shi 2000 Ota, Yasukawa, Fu & Chen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 Genus Mauremys (Stripe-necked Turtles)   Mauremys mutica Yellow Pond Turtle Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993   Mauremys reevesii Chinese Three-keeled Pond Turtle Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993   Mauremys sinensis Chinese Stripe-necked Turtle Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993  Family Platysternidae (Big-headed Turtle) Genus Platysternon (Big-headed Turtle)   Platysternon megacephalum Big-headed Turtle Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993  Family Trionychidae (Softshell Turtles) Genus Pelochelys (Indomalayan Giant Softshell Turtles)   Pelochelys cantorii Asian Giant Softshell Turtle Das 2008 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 [as bibroni] Genus Pelodiscus (East Asian Softshell Turtles)   Pelodiscus sinensis Chinese Softshell Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993 Genus Rafetus (Bicallosite Softshell Turtles)   EX Rafetus swinhoei Indochinese Giant Softshell Turtle Fritz & Havas 2006 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993