Reptiles of Mongolia    Bibliography of the reptiles of Mongolia Taxonomic history of the reptiles of Mongolia Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Laudakia (Asian Rock Agamas)   Laudakia stoliczkana Mongolian Rock Agama Ananjeva & Tuniev 1994 Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Baig, Wagner, Ananjeva & Boehme 2012 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Obst 1963 Sindaco & Jeremchenko 2008 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Zhao & Adler 1993 Genus Phrynocephalus (Toad-headed Agamas)   Phrynocephalus helioscopus Central Asian Sunwatcher Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Barabanov & Ananjeva 2007 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Sindaco & Jeremchenko 2008 Szczerbak 2003 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Zhao & Adler 1993   Phrynocephalus hispidus Dzungarian Toad-headed Agama Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Dunayev, Solovyeva & Poyarkov 2021   Phrynocephalus kulagini Tuva Toad-headed Agama Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Dunayev, Solovyeva & Poyarkov 2021 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006   Phrynocephalus versicolor Piebald Toad-headed Agama Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Barabanov & Ananjeva 2007 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Dunayev, Solovyeva & Poyarkov 2021 Obst 1963 Sindaco & Jeremchenko 2008 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Terentjev & Chernov 1965 [1949] Zhao & Adler 1993  Family Gekkonidae (Typical Geckos) Genus Alsophylax (Asian Lowland Pygmy Geckos)   Alsophylax pipiens Caspian Pygmy Gecko Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Obst 1963 Sindaco & Jeremchenko 2008 Szczerbak & Golubev 1996 Szczerbak in Boehme (ed.) 1981 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Wermuth 1965 Zhao & Adler 1993 Genus Tenuidactylus (Central Asian Thin-toed Geckos)   Tenuidactylus elongatus Gobi Thin-toed Gecko Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Szczerbak & Golubev 1996 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Zhao & Adler 1993  Family Lacertidae (Typical Lizards) Genus Eremias (Desert and Steppe Runners)   Eremias argus Mongolian Desert Runner Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Obst 1963 Szczerbak 1974 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Terentjev & Chernov 1965 [1949] Zhao & Adler 1993   Eremias arguta Kazakhstan Steppe Runner Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Sindaco & Jeremchenko 2008 Szczerbak 1974 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Zhao & Adler 1993   Eremias dzungarica Dzungarian Desert Runner Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Orlova, Polyarkov, Chirikova, Nazarov, Munkhbaatar, Munkhbayar & Terbish 2017   Eremias multiocellata Multiocellated Desert Runner Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Munkhbayar & Borkin 2010 Szczerbak 1974 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Zhao & Adler 1993   Eremias przewalskii Gobi Desert Runner Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Sindaco & Jeremchenko 2008 Szczerbak 1974 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Zhao & Adler 1993   Eremias velox Rapid Steppe Runner Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Terentjev & Chernov 1965 [1949]   Eremias vermiculata Central Asian Desert Runner Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Sindaco & Jeremchenko 2008 Szczerbak 1974 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Zhao & Adler 1993 Genus Lacerta (Eurasian Green Lizards)   Lacerta agilis Northern Eurasian Sand Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Sindaco & Jeremchenko 2008 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Terentjev & Chernov 1965 [1949] Zhao & Adler 1993 Genus Zootoca (Eurasian Forest Lizards)   Zootoca vivipara Viviparous Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldan-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 Dely & Boehme in Boehme (ed.) 1984 Obst 1963 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Zhao & Adler 1993  Family Sphaerodactylidae (Least Geckos, etc.) Genus Teratoscincus (Plate-tailed Geckos)   Teratoscincus przewalskii Mongolian Plate-tailed Gecko Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Macey, Ananjeva, Wang & Papenfuss 1997 Nazarov, Radjabizadeh, Poyarkov, Ananjeva, Melnikov & Rastegar-Pouyani 2017 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx tataricus Tatary Sand Boa Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Obst 1963 Sindaco, Venchi & Grieco 2013 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Terentjev & Chernov 1965 [1949] Zhao & Adler 1993  Family Colubridae (Typical Snakes or Colubrids) Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe dione Dione Rat Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Obst & Szczerbak in Boehme (ed.) 1993 Obst 1963 Schulz 1996 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006   Elaphe schrenckii Amur Rat Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Schulz 1996 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Genus Natrix (Eurasian Keelbacks)   Natrix natrix Common European Grass Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Kabisch in Boehme (ed.) 1999 Obst 1963 Sindaco, Venchi & Grieco 2013 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Terentjev & Chernov 1965 [1949] Zhao & Adler 1993 Genus Orientocoluber (North Asian Whip Snake)   Orientocoluber spinalis North Asian Whip Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Obst 1963 Sindaco, Venchi & Grieco 2013 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Zhao & Adler 1993  Family Lamprophiidae (House Snakes, etc.) Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis lineolatus Steppe Sand Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Obst 1963 Sindaco, Venchi & Grieco 2013 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Terentjev & Chernov 1965 [1949] Zhao & Adler 1993  Family Viperidae (Vipers) Genus Gloydius (Mamushis)   Gloydius caraganus Karaganda Mamushi Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006   Gloydius halys Central Asian Mamushi Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 Dirksen & Boehme 2005 Gloyd & Conant 1990 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Kropachev & Orlov 2017 Obst 1963 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Terentjev & Chernov 1965 [1949]   Gloydius ussuriensis Ussuri Mamushi Kropachev, Orlov & Orlova 2016 Genus Vipera (Typical Vipers or Adders)   Vipera berus Northern Eurasian Viper Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Borkin, Munkhbayar, Orlov, Semenov & Terbish 1990 David & Ineich 1999 Obst 1963 Sindaco, Venchi & Grieco 2013 Terbish, Munkhbayar, Clark, Munkhbat & Monks 2006 Zhao & Adler 1993