Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. (www.repfocus.dk).
Latest update: May 12th, 2024.
 
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Genus
Phrynocephalus
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Toad-headed Agamas, Toad Agamas
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Krötenkopfagamen
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Tudsehovedagamer
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1825 |  |
Phrynocephalus Kaup (type species: Lacerta caudivolvula Pallas 1814) |
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1831 | |
Megalochilus Eichwald (type species: Lacerta aurita Gmelin 1789; syn. Wermuth 1967; Szczerbak 1994; Barabanov & Ananjeva 2007) |
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1843 | |
Helioscopus Fitzinger (type species: Phrynocephalus helioscopus Wagler; syn. Wermuth 1967) |
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1843 | |
Phrynosaurus Fitzinger (type species: Phrynocephalus olivieri Dumeril & Bibron 1837; syn. Wermuth 1967) |
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1843 | |
Saccostoma Fitzinger (type species: Phrynocephalus auritus Fitzinger [=Lacerta aurita Gmelin 1789]; syn. Wermuth 1967) |
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2007 | |
Oreosaura Barabanov & Ananjeva (type species: Phrynocephalus vlangalii Strauch 1876) |
Contents:
62 species, of which 29 (46.8%) are endemic.
Endemism: 0%  100%
Remarks:
Previously included
Bufoniceps laungwalaensis.
Taxonomy of Phrynocephalus is still far from clear.
Status of some forms is a matter of controversy, and various authors have disagreed regarding validity of forms and synonymies.
The species list below follows Barabanov & Ananjeva (2007), with some adjustments based on subsequent works.
Distribution:
Middle East, C. and E. Asia
Reported from:
Afghanistan,
Armenia,
Azerbaijan,
China
(Gansu,
Hebei,
Inner Mongolia,
Ningxia,
Qinghai,
Shaanxi,
Shanxi,
Sichuan,
Tibet,
Xinjiang),
India
(Jammu & Kashmir),
Iran,
Iraq,
Jordan,
Kazakhstan,
Kuwait,
Kyrgyzstan,
Mongolia,
Nepal,
Oman,
Pakistan,
Qatar,
Russia,
Saudi Arabia,
Tajikistan,
Turkey,
Turkmenistan,
United Arab Emirates,
Uzbekistan,
Yemen.
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Phrynocephalus sp.
© Rune Midtgaard
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Phrynocephalus ahvazicus
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Ahvaz Toad-headed Agama
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Ahvaz-Krötenkopfagame
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Ahvaz-tudsehovedagam
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2014 | |
Phrynocephalus ahvazicus Melnikov, Melnikova, Nazarov, Rajabizadeh, Al-Johany, Amr & Ananjeva |
Remarks:
Previously confused with arabicus (see remarks under that species).
Records of arabicus from Kuwait (e.g., Al-Sirhan & Brown 2010), southern Iraq (Mohammed, Rhadi, Rastegar-Pouyani, Rastegar-Pouyani & Yousefkhani 2017), and southwestern Iran
(e.g., Kamali 2020) are doubtful and provisionally assigned to ahvazicus, the species of the arabicus group occurring most closely to this region.
Distribution:
Iran,
Iraq,
Kuwait.
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Phrynocephalus alpherakii
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Korgas Toad-headed Agama
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Korgas-Krötenkopfagame
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Korgas-tudsehovedagam
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1905 | |
Phrynocephalus alpherakii Bedriaga in Nikolsky |
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Phrynocephalus caudivolvulus alpherakii Tsarevsky 1926 |
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Phrynocephalus guttatus alpherakii Golubev 1992 |
Remarks:
Previously regarded as a subspecies of guttatus (e.g., Barabanov & Ananjeva 2007).
Melville, Hale, Mantziou, Ananjeva, Milto & Clemann (2009) suggested that alpherakii deserved recognition as separate species,
but this was regarded as premature by Sindaco, Venchi & Grieco (2013).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
China
(Xinjiang),
Kazakhstan.
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Phrynocephalus ananjevae
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Zagros Toad-headed Agama
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Zagros-Krötenkopfagame
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Zagros-tudsehovedagam
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2013 | |
Phrynocephalus ananjevae Melnikov, Melnikova, Nazarov & Rajabizadeh |
Distribution:
Iran.
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Phrynocephalus arabicus
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Arabian Toad-headed Agama
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Arabische Krötenkopfagame
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Arabisk Tudsehovedagam
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1894 | |
Phrynocephalus arabicus Anderson |
Remarks:
Now restricted to southern Arabia.
Previously included macropeltis, nejdensis, and the population now assigned to ahvazicus.
Records of arabicus from Kuwait (e.g., Al-Sirhan & Brown 2010), southern Iraq (Mohammed, Rhadi, Rastegar-Pouyani, Rastegar-Pouyani & Yousefkhani 2017), and southwestern Iran
(e.g., Kamali 2020) are doubtful and provisionally assigned to ahvazicus, the species of the arabicus group occurring most closely to this region.
Nevertheless, Kamali (2020) listed both ahvazicus and arabicus from Iran.
A record from Qatar (Gardner 2013) is provisionally assigned to macropeltis, the species of the arabicus group occurring most closely to Qatar.
Distribution:
Oman,
Saudi Arabia,
Yemen.
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Phrynocephalus axillaris
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East Turkestan Toad-headed Agama
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Ostturkestanische Krötenkopfagame
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Østturkestansk Tudsehovedagam
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1875 | |
Phrynocephalus axillaris Blanford |
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1906 | |
Phrynocephalus axillaris var. ericae Bedriaga "1905" (Barabanov & Ananjeva 2007) |
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1906 | |
Phrynocephalus axillaris var. idae Bedriaga "1905" (Barabanov & Ananjeva 2007) |
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1906 | |
Phrynocephalus axillaris var. murielis Bedriaga "1905" (Barabanov & Ananjeva 2007) |
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1906 | |
Phrynocephalus koslowi Bedriaga "1905" (Barabanov & Ananjeva 2007) |
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1907 | |
Phrynocephalus axillaris var. klemenzi Bedriaga (Barabanov & Ananjeva 2007) |
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1907 | |
Phrynocephalus axillaris var. maculata Bedriaga [not Phrynocephalus maculata Anderson 1872] (Barabanov & Ananjeva 2007) |
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1910 | |
Phrynocephalus Ludovici Mocquard (Barabanov & Ananjeva 2007) |
Remarks:
Previously included nasatus.
Distribution:
China
(Gansu,
Xinjiang).
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Phrynocephalus bannikovi
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Krasnovodsky Toad-headed Agama
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Krasnovodsky-Krötenkopfagame
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Krasnovodsky-tudsehovedagam
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1976 | |
Phrynocephalus reticulatus bannikovi Darevsky, Rustamov & Shammakov |
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Phrynocephalus bannikovi Lichnova 1992 |
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Phrynocephalus ocellatus bannikovi Szczerbak 1994 |
Remarks:
Previously treated as a subspecies of reticulatus.
Lichnova (1992) raised bannikovi to status as a separate species.
Validity confirmed by Macey, Schulte, Ananjeva, Dyke, Wang, Orlov, Shafiei, Robinson, Dujsebayeva, Freund, Fischer, Liu & Papenfuss (2018).
Distribution:
Turkmenistan.
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Phrynocephalus boettgeri
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Sherabad Toad-headed Agama
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Sherabad-Krötenkopfagame
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Sherabad-tudsehovedagam
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1905 | |
Phrynocephalus raddei var. boettgeri Bedriaga in Nikolsky |
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Phrynocephalus reticulatus boettgeri Tsarevsky 1926 |
Remarks:
Previously regarded as a synonym of raddei (e.g., Barabanov & Ananjeva 2007).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
Afghanistan,
Tajikistan,
Turkmenistan,
Uzbekistan.
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Phrynocephalus brevipes
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Short-toed Toad-headed Agama
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Kurzzehen-Krötenkopfagame
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Korttået Tudsehovedagam
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1907 | |
Phrynocephalus olivieri var. brevipes Nikolski |
Remarks:
Previously regarded as a synonym of scutellatus (e.g., Barabanov & Ananjeva 2007).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
Iran.
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Phrynocephalus cameranoi
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Yining Shi Toad-headed Agama
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Yining-Shi-Krötenkopfagame
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Yining Shi-tudsehovedagam
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1907 | |
Phrynocephalus helioscopus var. Cameranoi Bedriaga |
Remarks:
Previously regarded as a synonym of helioscopus (e.g., Barabanov & Ananjeva 2007).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
Kazakhstan.
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Phrynocephalus carinipes
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Pudesch Kupa Toad-headed Agama
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Pudesch-Kupa-Krötenkopfagame
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Pudesch Kupa-tudsehovedagam
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1907 | |
Phrynocephalus olivieri var. carinipes Nikolski |
Remarks:
Previously regarded as a synonym of scutellatus (e.g., Barabanov & Ananjeva 2007).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
Iran.
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Phrynocephalus clarkorum
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Afghan Toad-headed Agama
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Afghanische Krötenkopfagame
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Afghansk Tudsehovedagam
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1967 | |
Phrynocephalus clarkorum Anderson & Leviton |
Distribution:
Afghanistan,
Pakistan.
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Phrynocephalus erythrurus
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Northwest Tibetan Toad-headed Agama
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Nordwesttibetanische Krötenkopfagame
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Nordvesttibetansk Tudsehovedagam
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1909 | |
Phrynocephalus erythrurus Zugmayer |
Remarks:
Regarded as a synonym of theobaldi by Barabanov & Ananjeva (2007), but as a valid species by Jin & Liu (2010). The taxon parva was previously regarded as a synonym of vlangalii (Barabanov & Ananjeva 2007). This species occupies the highest regions of any reptile on earth (Jin & Liu 2010).
Distribution:
China
(Qinghai,
Tibet,
Xinjiang).
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Phrynocephalus euptilopus
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Darband Toad-headed Agama, Sand Toad Agama, (Spotted Toad-headed Agama)
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Darband-Krötenkopfagame
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Darband-tudsehovedagam
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1897 | |
Phrynocephalus euptilopus Alcock & Finn "1896" |
Remarks:
Although reported from India by several authors (e.g., Daniel 1983; Tikader & Sharma 1992; Das 1997; Daniel 2002), there seem to be no verified records from the country (Das 1996; Barabanov & Ananjeva 2007; Venugopal 2010).
Distribution:
Afghanistan,
Pakistan.
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Phrynocephalus forsythii
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Yarkand Toad-headed Agama
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Yarkand-Krötenkopfagame
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Yarkand-tudsehovedagam
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1872 | |
Phrynocephalus forsythii Anderson |
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Phrynocephalus theobaldi forsythii Boulenger 1885 |
Distribution:
China
(Xinjiang).
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Phrynocephalus frontalis
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Shanxi Toad-headed Agama
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Shanxi-Krötenkopfagame
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Shanxi-tudsehovedagam
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1876 | |
Phrynocephalus frontalis Strauch |
Remarks:
Previously regarded as a synonym of przewalskii (e.g., Barabanov & Ananjeva 2007).
Retained herein as a valid species, following Melville, Hale, Mantziou, Ananjeva, Milto & Clemann (2009) and Dunayev, Solovyeva & Poyarkov (2021).
Distribution:
China
(Beijing,
Gansu,
Hebei,
Inner Mongolia,
Ningxia,
Shaanxi,
Shanxi).
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Phrynocephalus golubewii
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Bami Toad-headed Agama, (Golubev's Toad-headed Agama)
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Bami-Krötenkopfagame
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Bami-tudsehovedagam
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1990 | |
Phrynocephalus golubewii Shenbrot & Semyonov |
Distribution:
Turkmenistan.
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Phrynocephalus guinanensis
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Guinan Toad-headed Agama
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Guinan-Krötenkopfagame
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Guinan-tudsehovedagam
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2009 | |
Phrynocephalus guinanensis Ji, Wang & Wang |
Remarks:
Jin & Brown (2019) suggested that guinanensis might be a synonym of of putjatai.
Not listed for China by Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che (2020).
Distribution:
China
(Qinghai).
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Phrynocephalus guttatus
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Spotted Toad-headed Agama, Tail Roller
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Gefleckter Krötenkopf, Schwanzroller
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Plettet Tudsehovedagam
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1789 | |
Lacerta guttata Gmelin |
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Agama guttata Daudin 1802 |
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Phrynocephalus guttatus Kaup 1827 |
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1814 | |
Lacerta caudivolvula Pallas (Barabanov & Ananjeva 2007) |
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Agama caudivolvula Lichtenstein 1823 |
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Phrynocephalus caudivolvulus Fitzinger 1826 |
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1831 | |
Phrynocephalus Pallasii Gray (Barabanov & Ananjeva 2007) |
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1905 | |
Phrynocephalus suschkini Bedriaga in Nikolsky (Barabanov & Ananjeva 2007) |
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1907 | |
Phrynocephalus acutirostris Bedriaga (Barabanov & Ananjeva 2007) |
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1983 | |
Phrynocephalus guttatus kalmykus Badmaeva & Shcherbak (Barabanov & Ananjeva 2007) |
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1987 | |
Phrynocephalus versicolor paraskiwi Semenov, Brushko, Shenbrot & Kubykin (Barabanov & Ananjeva 2007) |
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1995 | |
Phrynocephalus guttatus salsatus Gorelov, Dunayev & Kotenko in Golubev, Gorelov, Dunayev & Kotenko (Barabanov & Ananjeva 2007) |
Remarks:
Previously included alpherakii, incertus, melanurus, and moltschanovi.
See under reticulatus.
Distribution:
China
(Xinjiang),
Kazakhstan,
Russia,
Turkmenistan,
Uzbekistan.
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Phrynocephalus helioscopus
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Central Asian Sunwatcher
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Zentralasiatische Sonnengucker
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Centralasiatisk Solkigger
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1771 | |
Lacerta helioscopa Pallas |
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Agama helioscopus Lichtenstein 1823 |
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Phrynocephalus helioscopus Kaup 1827 |
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1789 | |
Lacerta uralensis Gmelin (Barabanov & Ananjeva 2007) |
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1831 | |
Phrynocephalus nigricans Eichwald (Barabanov & Ananjeva 2007) |
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1831 | |
Phrynocephalus varius Eichwald (Barabanov & Ananjeva 2007) |
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Phrynocephalus helioscopus varius Barabanov & Ananjeva 2007 |
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1841 | |
Lacerta gibba Eichwald [unavailable name] (Barabanov & Ananjeva 2007) |
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1909 | |
Phrynocephalus helioscopus saposhnikovi Kascenko (Barabanov & Ananjeva 2007) |
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1912 | |
Phrynocephalus helioscopus orientalis Bedriaga (Barabanov & Ananjeva 2007) |
Remarks:
Previously included cameranoi, horvathi, meridionalis, persicus, saidalievi, sergeevi, and turcomanus.
Distribution:
China
(Inner Mongolia,
Xinjiang),
Iran,
Kazakhstan,
Kyrgyzstan,
Mongolia,
Russia,
Tajikistan,
Turkmenistan,
Uzbekistan.
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Phrynocephalus hispidus
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Dzungarian Toad-headed Agama, (Spotted Toad-headed Agama)
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Dsungarischer Krötenkopfagame
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Dzungarisk Tudsehovedagam
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1909 | |
Phrynocephalus versicolor var. hispida Bedriaga |
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Phrynocephalus hispidus Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldán-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 |
Remarks:
Previously regarded as a synonym of versicolor (e.g., Barabanov & Ananjeva 2007).
Regarded as a separate species herein, following
Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldán-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer (2021) and
Dunayev, Solovyeva & Poyarkov (2021).
Distribution:
China
(Gansu,
Xinjiang),
Mongolia.
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Phrynocephalus hongyuanensis
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Sichuan Toad-headed Agama
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Sichuan-Krötenkopfagame
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Sichuan-tudsehovedagam
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1980 | |
Phrynocephalus vlangalii hongyuanensis Zhao, Jiang & Huang in Jiang, Huang & Zhao |
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Phrynocephalus hongyuanensis Wang & Jiang 1992 |
Remarks:
Previously treated as a subspecies of vlangalii.
Wang & Jiang (1992) raised hongyuanensis to status as a separate species.
Validity confirmed by Macey, Schulte, Ananjeva, Dyke, Wang, Orlov, Shafiei, Robinson, Dujsebayeva, Freund, Fischer, Liu & Papenfuss (2018).
Not listed for China by Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che (2020).
Distribution:
China
(Sichuan).
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Phrynocephalus horvathi
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Turkish Toad-headed Agama, (Horvath's Toad-headed Agama)
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Türkische Krötenkopfagame
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Tyrkisk Tudsehovedagam
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1894 | |
Phrynocephalus helioscopus var. horvathi Mehely |
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Phrynocephalus horvathi Melnikov, Melnikova, Nazarov & Rajabizadeh 2013 |
Remarks:
Previously regarded as a subspecies or a synonym of helioscopus (e.g., Mehely 1894) or of persicus (e.g., Anderson 1999; Barabanov & Ananjeva 2007). Arakelyan, Danielyan, Corti, Sindaco & Leviton (2011) suggested that horvathi deserved recognition as a separate species. It was formally revalidated by Melnikov, Melnikova, Nazarov & Rajabizadeh (2013).
Distribution:
Armenia,
Azerbaijan,
Iran,
Turkey.
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Phrynocephalus incertus
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Almaty Toad-headed Agama
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Almaty-Krötenkopfagame
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Almaty-tudsehovedagam
|
|
1905 | |
Phrynocephalus caudivolvolus var. incerta Bedriaga in Nikolsky |
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Phrynocephalus guttatus incerta Melville, Hale, Mantziou, Ananjeva, Milto & Clemann 2009 |
Remarks:
Previously regarded as a synonym of guttatus (e.g., Barabanov & Ananjeva 2007).
Melville, Hale, Mantziou, Ananjeva, Milto & Clemann (2009) suggested that incerta deserved recognition as a separate species,
but this was regarded as premature by Sindaco, Venchi & Grieco (2013).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
Kazakhstan.
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Phrynocephalus interscapularis
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Rose-shouldered Toad-headed Agama, Spiny Toad-headed Agama
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Rosaschulter-Krötenkopfagame, Stachelkopf
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Rosenskuldret Tudsehovedagam
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|
1856 | |
Phrynocephalus interscapularis Lichtenstein |
Remarks:
Previously included sogdianus.
Listed for Iran by Kamali & Anderson (2015) and (Nasrabadi, Rastegar-Pouyani, Rastegar-Pouyani & Gharzi 2017), but omitted(?) by Kamali (2020).
Distribution:
Afghanistan,
Iran,
Kazakhstan,
Turkmenistan,
Uzbekistan.
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Phrynocephalus kangsuensis
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Kangsu Toad-headed Agama
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Kangsu-Krötenkopfagame
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Kangsu-tudsehovedagam
|
|
2024 | |
Phrynocephalus kangsuensis Liang & Shi |
Distribution:
China
(Xinjiang).
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Phrynocephalus khorasanus
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Khorasan Bearded Toad-headed Agama
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Bärtiger Khorasan-Krötenkopfagame
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Skægget Khorasan-tudsehovedagam
|
|
2018 | |
Phrynocephalus mystaceus khorasanus Solovyeva, Dunayev, Nazarov, Radjabizadeh & Poyarkov |
Remarks:
Previously regarded as a subspecies of mystaceus.
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
Iran.
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Phrynocephalus kulagini
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Tuva Toad-headed Agama
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|
Tuva-Krötenkopfagame
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Tuva-tudsehovedagam
|
|
1909 | |
Phrynocephalus versicolor var. Kulagini Bedriaga |
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| |
Phrynocephalus kulagini Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldán-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer 2021 |
Remarks:
Previously regarded as a synonym or a subspecies of versicolor (e.g., Barabanov & Ananjeva 2007; Dunayev & Orlova 2023).
Regarded as a separate species herein, following
Buehler, Zoljargal, Purvee, Munkhbayar, Munkhbaatar, Batsaikhan, Ananjeva, Orlov, Papenfuss, Roldán-Pina, Douchindorj, Grismer, Oaks, Brown & Grismer (2021) and
Dunayev, Solovyeva & Poyarkov (2021).
Distribution:
Mongolia,
Russia.
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Phrynocephalus kuschakewitschi
|
Ajaguz Toad-headed Agama
|
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Ajaguz-Krötenkopfagame
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Ajaguz-tudsehovedagam
|
|
1905 | |
Phrynocephalus kuschakewitschi Bedriaga in Nikolsky |
|
| |
Phrynocephalus caudivolvulus kuschakewitschi Tsarevsky 1926 |
|
| |
Phrynocephalus guttatus kuschakewitschi Terentjev & Chernov 1949 |
Remarks:
Previously regarded as a synonym of melanurus (e.g., Barabanov & Ananjeva 2007).
Melville, Hale, Mantziou, Ananjeva, Milto & Clemann (2009) suggested that kuschakewitschi deserved recognition as a separate species,
but this was regarded as premature by Sindaco, Venchi & Grieco (2013).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
Kazakhstan.
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Phrynocephalus lhasaensis
|
Lhasa Toad-headed Agama
|
|
Lhasa-Krötenkopfagame
|
|
Lhasa-tudsehovedagam
|
|
1999 | |
Phrynocephalus theobaldi orientalis Wang, Papenfuss & Zeng in Wang, Zheng, Fang, Wu, Papenfuss & Macey [not Phrynocephalus helioscopus var. orientalis Bedriaga 1912] (Barabanov & Ananjeva 2007) |
|
2002 | |
Phrynocephalus theobaldi lhasaensis Barabanov, Ananjeva, Papenfuss & Wang [substitute name for Phrynocephalus theobaldi orientalis Wang, Papenfuss & Zeng in Wang, Zheng, Fang, Wu, Papenfuss & Macey 1999] |
Remarks:
Previously regarded as a synonym of theobaldi (e.g., Barabanov & Ananjeva 2007).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
China
(Tibet).
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|
|
Phrynocephalus longicaudatus
|
Long-tailed Toad-headed Agama
|
|
Langschwanz-Krötenkopfagame
|
|
Langhalet Tudsehovedagam
|
|
1957 | |
Phrynocephalus maculatus longicaudatus Haas |
|
| |
Phrynocephalus longicaudatus Macey, Schulte, Ananjeva, Dyke, Wang, Orlov, Shafiei, Robinson, Dujsebayeva, Freund, Fischer, Liu & Papenfuss 2018 |
Remarks:
Not mentioned for Saudi Arabia by Aloufi, Amr, Baker & Hamidan (2019).
Distribution:
Iraq,
Jordan,
Kuwait,
Oman,
Saudi Arabia,
United Arab Emirates,
Yemen.
|
|
|
Phrynocephalus lutensis
|
Lut Toad-headed Agama
|
|
Lut-Krötenkopfagame
|
|
Lut-tudsehovedagam
|
|
2015 | |
Phrynocephalus lutensis Kamali & Anderson |
Distribution:
Iran.
|
|
|
Phrynocephalus luteoguttatus
|
Yellow-speckled Toad-headed Agama, Spiny-headed Toad-headed Agama
|
|
Gelbfleckiger Krötenkopfagame
|
|
Gulprikket Tudsehovedagam
|
|
1887 | |
Phrynocephalus luteoguttatus Boulenger |
Remarks:
Although reported from India by Das (1997), there are no verified records from the country (Venugopal 2010).
Distribution:
Afghanistan,
Pakistan.
|
|
|
Phrynocephalus macropeltis
|
Dhahran Toad-headed Agama
|
|
Dhahran-Krötenkopfagame
|
|
Dhahran-tudsehovedagam
|
|
1957 | |
Phrynocephalus nejdensis macropeltis Haas |
|
| |
Phrynocephalus macropeltis Melnikov, Melnikova, Nazarov, Rajabizadeh, Al-Johany, Amr & Ananjeva 2014 |
Remarks:
Previously regarded as a synonym of arabicus (e.g., Barabanov & Ananjeva 2007). Revalidated as a separate species by Melnikov, Melnikova, Nazarov, Rajabizadeh, Al-Johany, Amr & Ananjeva (2014). A record from Qatar of arabicus (Gardner 2013) is provisionally assigned to macropeltis, the species of the arabicus group occurring most closely to Qatar.
Distribution:
Qatar,
Saudi Arabia,
United Arab Emirates.
|
|
|
Phrynocephalus maculatus
|
Black-tailed Toad-headed Agama, Whip-tailed Toad-headed Agama, (Spotted Toad-headed Agama)
|
|
Schwarzschwanz-Krötenkopfagame
|
|
Sorthalet Tudsehovedagam
|
|
1872 | |
Phrynocephalus maculatus Anderson |
|
1896 | |
Phrynocephalus spiniventris Nikolsky (Barabanov & Ananjeva 2007) |
|
| |
Phrynocephalus maculatus spiniventris Nikolsky 1900 |
Remarks:
Previously included longicaudatus.
Records from Turkmenistan (e.g., Szczerbak 1994) refer to golubewii (Shenbrot & Semenov 1990; Barabanov & Ananjeva 2007).
Distribution:
Afghanistan,
Iran,
Pakistan.
|
|
|
Phrynocephalus melanurus
|
Kurchum Toad-headed Agama
|
|
Kurchum-Krötenkopfagame
|
|
Kurchum-tudsehovedagam
|
|
1831 | |
Phrynocephalus melanurus Eichwald |
|
| |
Phrynocephalus guttatus melanurus Melville, Hale, Mantziou, Ananjeva, Milto & Clemann 2009 |
|
1906 | |
Phrynocephalus bergi Elpatjevsky "1907" (Barabanov & Ananjeva 2007) |
|
1907 | |
Phrynocephalus Arcellazzii Bedriaga (Barabanov & Ananjeva 2007) |
|
1907 | |
Phrynocephalus Haeckeli Bedriaga (Barabanov & Ananjeva 2007) |
|
1907 | |
Phrynocephalus Isseli Bedriaga (Barabanov & Ananjeva 2007) |
|
1907 | |
Phrynocephalus Salenskyi Bedriaga (Barabanov & Ananjeva 2007) |
|
| |
Phrynocephalus guttatus salenskyi Szczerbak 2003 |
|
1909 | |
Phrynocephalus Grum-Grzimailoi [grumgrzimaloi] Bedriaga (Barabanov & Ananjeva 2007) |
|
1909 | |
Phrynocephalus helioscopus levis Kascenko (Barabanov & Ananjeva 2007) |
|
| |
Phrynocephalus helioscopus laevis Nikolski 1915 [substitute name for Phrynocephalus helioscopus var. levis Kascenko 1909] |
|
1926 | |
Phrynocephalus bedrjagai Zarevsky (Barabanov & Ananjeva 2007) |
|
1979 | |
Phrynocephalus albo-lineatus Zhao (Barabanov & Ananjeva 2007) |
Remarks:
Formerly included kuschakewitschi.
Previously regarded as a subspecies of guttatus (e.g., Barabanov & Ananjeva 2007).
Melville, Hale, Mantziou, Ananjeva, Milto & Clemann (2009) suggested that melanurus deserved recognition as a separate species,
but this was regarded as premature by Sindaco, Venchi & Grieco (2013).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Ananjeva, Guo & Wang (2011) listed albolineatus as a separate species.
Liu, Song, Li & Shi (2012) treated grumgrzimaloi as a separate species.
Distribution:
China
(Xinjiang),
Kazakhstan.
|
|
|
Phrynocephalus meridionalis
|
Surkhandarya Toad-headed Agama
|
|
Surkhandarya-Krötenkopfagame
|
|
Surkhandarya-tudsehovedagam
|
|
2012 | |
Phrynocephalus helioscopus meridionalis Solovyeva, Dunayev & Poyarkov |
Remarks:
Previously regarded as a subspecies of helioscopus.
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
Turkmenistan,
Uzbekistan.
|
|
|
Phrynocephalus moltschanovi
|
Bel-Tau Toad-headed Agama, (Molchanov's Toad-headed Agama)
|
|
Bel-Tau-Krötenkopfagame
|
|
Bel-Tau-tudsehovedagam
|
|
1913 | |
Phrynocephalus caudivolvolus var. moltschanovi Nikolski |
|
| |
Phrynocephalus moltschanovi Semenov & Shenbrot 1982 |
Remarks:
Previously regarded as a synonym of guttatus (e.g., Barabanov & Ananjeva 2007).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023), although treated as such already by Semenov & Shenbrot (1982).
Distribution:
Kazakhstan.
|
|
|
Phrynocephalus mystaceus
|
Bearded Toad-headed Agama
|
|
Bärtiger Krötenkopfagame
|
|
Skægget Tudsehovedagam
|
|
1776 | |
Lacerta mystacea Pallas |
|
| |
Agama mystacea Merrem 1820 |
|
| |
Phrynocephalus mystacia [sic] Kaup 1827 |
|
| |
Megalochilus mystaceus Ananjeva 1986 |
|
1776 | |
Lacerta aurita Pallas (Barabanov & Ananjeva 2007) |
|
| |
Gekko auritus Latreille 1801 |
|
| |
Agama aurita Daudin 1802 |
|
| |
Phrynocephalus auritus Fitzinger 1826 |
|
| |
Megalochilus auritus Eichwald 1831 |
|
| |
Saccostoma auritum Fitzinger 1843 |
|
1802 | |
Lacerta lobata Shaw [substitute name for Lacerta aurita Gmelin 1789] (Barabanov & Ananjeva 2007) |
|
1932 | |
Phrynocephalus mystaceus galli Krassowsky (Barabanov & Ananjeva 2007) |
|
1932 | |
Phrynocephalus mystaceus mystaceus natio Dagestanica Krassowsky (unavailable name; Barabanov & Ananjeva 2007) |
|
| |
Megalochilus mystaceus dagestanica Ananjeva 1987 "1986" (Barabanov & Ananjeva 2007) |
|
1990 | |
Phrynocephalus mystaceus aurantiacocaudatus Semyonov & Shenbrot |
Remarks:
Previously included khorasanus.
Introduced to:
Azerbaijan.
Distribution:
Afghanistan,
China
(Xinjiang),
Iran,
Kazakhstan,
Russia,
Tajikistan,
Turkmenistan,
Uzbekistan.
|
|
|
Phrynocephalus nanschanicus
|
Nanshan Toad-headed Agama
|
|
Nanshan-Krötenkopfagame
|
|
Nanshan-tudsehovedagam
|
|
1906 | |
Phrynocephalus vlangalii var. nanschanica Bedriaga ["1905"] |
|
| |
Phrynocephalus nanschanicus Tsarevsky 1927 |
Remarks:
Previously regarded as a synonym of vlangalii (e.g., Barabanov & Ananjeva 2007).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
China
(Qinghai).
|
|
|
Phrynocephalus nasatus
|
Aksu Toad-headed Agama
|
|
Aksu-Krötenkopfagame
|
|
Aksu-tudsehovedagam
|
|
1995 | |
Phrynocephalus nasatus Golubev & Dunayev |
Remarks:
Previously regarded as a synonym of axillaris (e.g., Barabanov & Ananjeva 2007).
Revalidated as a separate species by Dunayev (2020).
Distribution:
China
(Xinjiang).
|
|
|
Phrynocephalus nejdensis
|
Nejd Toad-headed Agama
|
|
Nejd-Krötenkopfagame
|
|
Nejd-tudsehovedagam
|
|
1957 | |
Phrynocephalus nejdensis Haas |
Remarks:
Previously regarded as a synonym of arabicus (e.g., Barabanov & Ananjeva 2007). Revalidated as a separate species by Melnikov, Melnikova, Nazarov, Rajabizadeh, Al-Johany, Amr & Ananjeva (2014).
Distribution:
Jordan,
Saudi Arabia.
|
|
|
Phrynocephalus ornatus
|
Southern Striped Toad-headed Agama, Black-and-White Toad-headed Agama
|
|
Südliche Gestreifte Krötenkopfagame
|
|
Sydlig Stribet Tudsehovedagam
|
|
1887 | |
Phrynocephalus ornatus Boulenger |
Remarks:
Previously included vindumi.
Distribution:
Afghanistan,
Pakistan.
|
|
|
Phrynocephalus parvus
|
Dytschu Toad-headed Agama
|
|
Dytschu-Krötenkopfagame
|
|
Dytschu-tudsehovedagam
|
|
1909 | |
Phrynocephalus Vlangalii var. parva Bedriaga |
|
| |
Phrynocephalus parvus Tsarevsky 1927 |
|
| |
Phrynocephalus erythrurus parva Jin & Liu 2010 |
Remarks:
Previously regarded as a subspecies of erythrurus (e.g., Jin & Liu 2010).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023), although treated as such already by Tsarevsky (1927).
Distribution:
China
(Qinghai).
|
|
|
Phrynocephalus persicus
|
Persian Toad-headed Agama
|
|
Persische Krötenkopfagame
|
|
Persisk Tudsehovedagam
|
|
1863 | |
Phrynocephalus persicus De Filippi |
|
| |
Phrynocephalus helioscopus persicus Terentjev & Chernov 1949 |
Remarks:
Previously regarded as a subspecies of helioscopus. Revalidated as a separate species by Anderson (1999). Previously included horvathi.
Distribution:
Armenia,
Azerbaijan,
Iran.
|
|
|
Phrynocephalus przewalskii
|
Alashan Toad-headed Agama
|
|
Alasha-Krötenkopfagame
|
|
Alasha-tudsehovedagam
|
|
1876 | |
Phrynocephalus Przewalskii Strauch |
|
1876 | |
Phrynocephalus affinis Strauch (Barabanov & Ananjeva 2007) |
|
1906 | |
Phrynocephalus pewzowi Bedriaga "1905" [nomen nudum] (Barabanov & Ananjeva 2007) |
|
1907 | |
Phrynocephalus Pewzowi Bedriaga (Barabanov & Ananjeva 2007) |
|
1907 | |
Phrynocephalus steindachneri Bedriaga (Barabanov & Ananjeva 2007) |
|
1909 | |
Phrynocephalus carinilabris Bedriaga (Barabanov & Ananjeva 2007) |
|
1909 | |
Phrynocephalus ciliaris Bedriaga (Barabanov & Ananjeva 2007) |
|
1909 | |
Phrynocephalus potanini Bedriaga (Barabanov & Ananjeva 2007) |
|
1909 | |
Phrynocephalus Reldæ Bedriaga (Barabanov & Ananjeva 2007) |
|
1927 | |
Phrynocephalus birulai Zarevsky (Barabanov & Ananjeva 2007) |
|
1927 | |
Phrynocephalus carinatus Zarevsky (Barabanov & Ananjeva 2007) |
|
1927 | |
Phrynocephalus elegans Zarevsky (Barabanov & Ananjeva 2007) |
|
1927 | |
Phrynocephalus parvulus Zarevsky (Barabanov & Ananjeva 2007) |
|
1927 | |
Phrynocephalus suschkinianus Zarevsky (Barabanov & Ananjeva 2007) |
|
1984 | |
Phrynocephalus werneri Peters [unavailable name] (Barabanov & Ananjeva 2007) |
Remarks:
Previously included frontalis.
Distribution:
China
(Beijing,
Gansu,
Inner Mongolia,
Ningxia).
|
|
|
Phrynocephalus putjatai
|
Guide Toad-headed Agama
|
|
Guide-Krötenkopfagame
|
|
Guide-tudsehovedagam
|
|
1909 | |
Phrynocephalus Putjatai Bedriaga |
Remarks:
Zhao & Adler (1993) regarded the status of this species as uncertain.
Barabanov & Ananjeva (2007) treated it as a valid species.
Validity supported by Guo & Wang (2007) and Noble, Qi & Fu (2010).
Specific epithet occasionally misspelled putjatia (e.g., Noble, Qi & Fu 2010).
Distribution:
China
(Gansu,
Qinghai).
|
|
|
Phrynocephalus raddei
|
Trans-Caspian Toad-headed Agama
|
|
Trans-Kaspische Krötenkopfagame
|
|
Transkaspisk Tudsehovedagam
|
|
1888 | |
Phrynocephalus raddei Boettger |
|
1915 | |
Phrynocephalus raddei var. bilkewitschi Nikolski (Barabanov & Ananjeva 2007) |
|
1915 | |
Phrynocephalus raddei var. zarudnyi Nikolski (Barabanov & Ananjeva 2007) |
Remarks:
Previously included boettgeri.
Records from Afghanistan refer to boettgeri (Wagner, Bauer, Leviton, Wilms & Böhme 2016).
Previously regarded as a synonym of reticulatus (e.g., Terentjev & Chernov 1949; Wermuth 1967), but as a valid species by Szczerbak (1994) and Dunayev (1996).
Although presence in Iran has previously been questioned (e.g., Anderson 1999; Safaei-Mahroo & al. 2015), raddei was recently confirmed for the country
(Khosravani, Khani, Oraie & Rasyegar-Pouyani 2014), although not listed for Iran by Kamali (2020).
Distribution:
Iran,
Tajikistan,
Turkmenistan,
Uzbekistan.
|
|
|
Phrynocephalus reticulatus
|
Reticulated Toad-headed Agama, Netted Toad-headed Agama, (Ladakh Toad-headed Agama)
|
|
Netz-Krötenkopfagame
|
|
Retikuleret Tudsehovedagam
|
|
1823 | |
Agama ocellata Lichtenstein in Eversmann (Barabanov & Ananjeva 2007; see remarks) |
|
| |
Phrynocephalus ocellatus Golubev 1991 |
|
1831 | |
Phrynocephalus reticulatus Eichwald |
|
| |
Phrynocephalus caudivolvolus reticulatus Nikolsky 1915 |
|
| |
Phrynocephalus theobaldi reticulatus Das 2003 |
|
1905 | |
Phrynocephalus boulengeri Bedriaga in Nikolsky (Barabanov & Ananjeva 2007) |
|
| |
Phrynocephalus nikolskii boulengeri Nikolsky 1915 |
|
1905 | |
Phrynocephalus nikolskii Bedriaga in Nikolsky (Barabanov & Ananjeva 2007) |
Remarks:
Previously included bannikovi.
Agama ocellata Lichtenstein in Eversmann 1823 was formerly regarded as a synonym of caudivolvolus (e.g., Boulenger 1885) or of guttatus (e.g., Lantz 1924; Wermuth 1967). Golubev (1991) demonstrated that ocellata is a senior synonym of reticulatus and referred to the species as ocellatus. In the interest of taxonomic stability, the species is here referred to as reticulatus, following Barabanov & Ananjeva (2007), since this name has been used many times for more than a century. P. raddei was previously confused with reticulatus (e.g., Wermuth 1967), and older records may refer to either species. Records from Afghanistan (e.g., Leviton & Anderson 1970 [as reticulatus boettgeri]) refer to raddei (Clark 1990).
Distribution:
Turkmenistan,
Uzbekistan.
|
|
|
Phrynocephalus roborowskii
|
Zaidam Toad-headed Agama
|
|
Zaidam-Krötenkopfagame
|
|
Zaidam-tudsehovedagam
|
|
1906 | |
Phrynocephalus vlangalii var. roborowskii Bedriaga "1905" |
|
| |
Phrynocephalus roborowskii Peters 1984 |
Remarks:
Regarded as a synonym of vlangalii by Zhao & Adler (1993), but as a valid species by Barabanov & Ananjeva (2007).
Not listed for China by Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che (2020).
Distribution:
China
(Qinghai).
|
|
|
Phrynocephalus rossikowi
|
Khentau Toad-headed Agama
|
|
Khentau-Krötenkopfagame
|
|
Khentau-tudsehovedagam
|
|
1899 | |
Phrynocephalus rossikowi Nikolsky |
|
1915 | |
Phrynocephalus rossikowi relictus Nikolski (Barabanov & Ananjeva 2007) |
|
1979 | |
Phrynocephalus rossikowi shammakowi Shcherbak & Golubev (Barabanov & Ananjeva 2007) |
Distribution:
Turkmenistan,
Uzbekistan.
|
|
|
Phrynocephalus saidalievi
|
Konibodom Toad-headed Agama
|
|
Konibodom-Krötenkopfagame
|
|
Konibodom-tudsehovedagam
|
|
1981 | |
Phrynocephalus helioscopus saidalievi Sattorov |
|
| |
Phrynocephalus saidalievi Eremchenko & Panfilov 1999 |
Remarks:
Previously regarded as a subspecies of helioscopus (e.g., Barabanov & Ananjeva 2007).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023), although treated as such already by Eremchenko & Panfilov (1999).
Distribution:
Kyrgyzstan,
Tajikistan,
Uzbekistan.
|
|
|
Phrynocephalus sakoi
|
Sharqiyah Toad-headed Agama
|
|
Sharqiyah-Krötenkopfagame
|
|
Sharqiyah-tudsehovedagam
|
|
2015 | |
Phrynocephalus sakoi Melnikov, Melnikova, Nazarov, Al-Johany & Ananjeva |
Distribution:
Oman.
|
|
|
Phrynocephalus scutellatus
|
Grey Toad-headed Agama, Banded Toad-headed Agama
|
|
Graue Krötenkopfagame
|
|
Grå Tudsehovedagam
|
|
1807 | |
Agame scutellata [incorrect original spelling] Olivier |
|
| |
Phrynocephalus scutellatus Moquard 1910 |
|
1837 | |
Phrynocephalus olivieri Duméril & Bibron (Barabanov & Ananjeva 2007) |
Remarks:
Previously included brevipes, cameranoi, and tickelii.
Distribution:
Afghanistan,
Iran,
Pakistan.
|
|
|
Phrynocephalus sergeevi
|
Navoi Toad-headed Agama
|
|
Navoi-Krötenkopfagame
|
|
Navoi-tudsehovedagam
|
|
2012 | |
Phrynocephalus helioscopus sergeevi Solovyeva, Dunayev & Poyarkov |
Remarks:
Previously regarded as a subspecies of helioscopus.
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
Uzbekistan.
|
|
|
Phrynocephalus sogdianus
|
Tajik Toad-headed Agama
|
|
Tadschikische Krötenkopfagame
|
|
Tadsjikisk Tudsehovedagam
|
|
1948 | |
Phrynocephalus interscapularis sogdianus Chernov |
|
| |
Phrynocephalus sogdianus Sokolovsky 1975 |
Remarks:
Previously treated as a subspecies of interscapularis.
Sokolovsky (1975) raised sogdianus to status as a separate species.
Validity confirmed by Macey, Schulte, Ananjeva, Dyke, Wang, Orlov, Shafiei, Robinson, Dujsebayeva, Freund, Fischer, Liu & Papenfuss (2018).
Distribution:
Afghanistan,
Tajikistan,
Uzbekistan.
|
|
|
Phrynocephalus strauchi
|
Fergana Valley Toad-headed Agama, (Strauch's Toad-headed Agama)
|
|
Fergana-Krötenkopfagame
|
|
Fergana-tudsehovedagam
|
|
1899 | |
Phrynocephalus strauchi Nikolsky |
|
| |
Phrynocephalus reticulatus strauchi Terentjev & Chernov 1949 |
Remarks:
Previously treated as a subspecies of reticulatus.
Mesherin & Golubev (1989) revalidated strauchi to status as a separate species.
Validity confirmed by Macey, Schulte, Ananjeva, Dyke, Wang, Orlov, Shafiei, Robinson, Dujsebayeva, Freund, Fischer, Liu & Papenfuss (2018).
Distribution:
Kyrgyzstan,
Tajikistan,
Uzbekistan.
|
|
|
Phrynocephalus theobaldi
|
Kashmir Toad Agama, (Theobald's Toad Agama)
|
|
Himalaya-Krötenkopfagame
|
|
Kashmir-tudsehovedagam
|
|
1863 | |
Phrynocephalus theobaldi Blyth |
|
1867 | |
Phrynocephalus stoliczkai Steindachner (Barabanov & Ananjeva 2007) |
|
1909 | |
Phrynocephalus vlangalii var. lidskii Bedriaga (Barabanov & Ananjeva 2007) |
|
| |
Phrynocephalus lidskii Tsarevsky 1927 |
|
1984 | |
Phrynocephalus alticola Peters (Barabanov & Ananjeva 2007) |
|
1996 | |
Phrynocephalus zetangensis Wang, Zeng, Fang, Liu, Wu, Papenfuss & Macey (the authors also used the spelling zetanensis in the original decription) (Barabanov & Ananjeva 2007; Guo & Wang 2007) |
Remarks:
Previously included erythrurus and lhasaensis.
Ananjeva, Guo & Wang (2011) questioned the presence of theobaldi in Xinjiang, referring to Zhao (1979).
Distribution:
China
(Tibet,
Xinjiang),
India
(Jammu & Kashmir),
Nepal.
|
|
|
Phrynocephalus tickelii
|
1845 | |
Phrynocephalus Tickelii Gray |
Remarks:
Previously regarded as a synonym of scutellatus (e.g., Barabanov & Ananjeva 2007).
Raised to status as a separate species by Solovyeva, Dunayev, Nazarov, Bondarenko & Poyarkov (2023).
Distribution:
Afghanistan.
|
|
|
Phrynocephalus turcomanus
|
Turkmen Sunwatcher
|
|
Turkmenische Sonnengucker
|
|
Turkmensk Solkigger
|
|
2012 | |
Phrynocephalus helioscopus turcomanus Solovyeva, Dunayev & Poyarkov |
|
| |
Phrynocephalus turcomanus Macey, Schulte, Ananjeva, Dyke, Wang, Orlov, Shafiei, Robinson, Dujsebayeva, Freund, Fischer, Liu & Papenfuss 2018 |
Distribution:
Iran,
Turkmenistan.
|
|
|
Phrynocephalus versicolor
|
Piebald Toad-headed Agama
|
|
Mongolischer Krötenkopfagame
|
|
Mongolsk Tudsehovedagam
|
|
1876 | |
Phrynocephalus versicolor Strauch |
|
1906 | |
Phrynocephalus guentheri Bedriaga "1905" (Barabanov & Ananjeva 2007) |
|
1906 | |
Phrynocephalus versicolor var. doriai Bedriaga "1905" [nomen nudum] (Barabanov & Ananjeva 2007) |
|
1909 | |
Phrynocephalus Blanfordi Bedriaga (Barabanov & Ananjeva 2007) |
|
1909 | |
Phrynocephalus versicolor var. Bogdanowi Bedriaga (Barabanov & Ananjeva 2007) |
|
1909 | |
Phrynocephalus versicolor var. Doriai Bedriaga (Barabanov & Ananjeva 2007) |
|
1909 | |
Phrynocephalus versicolor var. Siebenrocki Bedriaga (Barabanov & Ananjeva 2007) |
|
1930 | |
Phrynocephalus rostralis Zarevsky (Barabanov & Ananjeva 2007) |
|
1958 | |
Phrynocephalus stejnegeri Bannikov [nomen nudum] (Barabanov & Ananjeva 2007) |
|
1984 | |
Phrynocephalus steynegeri Peters [unavailable name] (Barabanov & Ananjeva 2007) |
|
1995 | |
Phrynocephalus immaculatus Zhao (Barabanov & Ananjeva 2007) |
Remarks:
Previously included hispidus and kulagini.
Records from Kazakhstan refer to the former subspecies paraskiwi, now considered a synonym of guttatus (Golubev 1989; Szczerbak 2003; Barabanov & Ananjeva 2007).
Introduced to:
Philippines
(Luzon).
Distribution:
China
(Gansu,
Inner Mongolia,
Ningxia,
Xinjiang),
Mongolia.
|
|
|
Phrynocephalus vindumi
|
Northern Striped Toad-headed Agama
|
|
Nördliche Gestreifte Krötenkopfagame
|
|
Nordlig Stribet Tudsehovedagam
|
|
1998 | |
Phrynocephalus ornatus vindumi Golubev |
|
| |
Phrynocephalus vindumi Macey, Schulte, Ananjeva, Dyke, Wang, Orlov, Shafiei, Robinson, Dujsebayeva, Freund, Fischer, Liu & Papenfuss 2018 |
Distribution:
Iran.
|
|
|
Phrynocephalus vlangalii
|
Quinghai Toad-headed Agama
|
|
Quinghai-Krötenkopfagame
|
|
Quinghai-tudsehovedagam
|
|
1876 | |
Phrynocephalus Vlangalii Strauch |
|
1909 | |
Phrynocephalus Vlangalii var. geckoides Bedriaga (Barabanov & Ananjeva 2007) |
|
| |
Phrynocephalus geckoides Tsarevsky 1927 |
|
1909 | |
Phrynocephalus Vlangalii var. Grombtschewskii Bedriaga (Barabanov & Ananjeva 2007) |
|
| |
Phrynocephalus grombtschewskii Tsarevsky 1927 |
|
1909 | |
Phrynocephalus Vlangalii var. Pylzowi Bedriaga (Barabanov & Ananjeva 2007) |
|
| |
Phrynocephalus pylzowi Tsarevsky 1927 |
|
| |
Phrynocephalus parvus pylzowi Tsarevsky 1927 |
|
1984 | |
Phrynocephalus occidentalis Peters [unavailable name] (Barabanov & Ananjeva 2007) |
Remarks:
Previously included nanschanicus, hongyuanensis, and parva.
Distribution:
China
(Gansu,
Qinghai,
Sichuan,
Xinjiang).
|
|
|