Reptiles of Luzon [Philippines] Related checklists:  Philippines Bibliography of the reptiles of Luzon Taxonomic history of the reptiles of Luzon Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: April 14th, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus mindorensis Philippine Freshwater Crocodile Alcala 1986 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Trutnau & Sommerlad 2006 Wermuth & Mertens 1977   Crocodylus porosus Saltwater Crocodile Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama Brown, Ferner, Sison, Gonzales & Kennedy 1996 David, Pauwels, Lays & Lenglet 2006 Lagat 2009   Bronchocela marmorata Marbled Long-tailed Agama Alcala 1986 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Hallermann 2005 Manthey 2008 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Wermuth 1967 Genus Draco (Gliding Dragons)   Draco spilopterus Common Philippine Gliding Dragon Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Lagat 2009 Manthey 2008 McGuire & Alcala 2000 McGuire & Kiew 2001 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Genus Gonocephalus (Western Anglehead Agamas)   Gonocephalus sophiae Dark-spotted Anglehead Agama Lagat 2009 [as G. sp.] Manthey 2010 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Welton, Siler, Grismer, Diesmos, Sites & Brown 2017 Genus Hydrosaurus (Sail-tailed Agamas)   Hydrosaurus pustulatus Philippine Sail-tailed Agama Gaulke 1989 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Wermuth 1967  Family Gekkonidae (Typical Geckos) Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus philippinicus Common Philippine Bent-toed Gecko Alcala 1986 Brown & Alcala 1978 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Lagat 2009 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Welton, Siler, Linkem, Diesmos & Brown 2010 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Brown & Alcala 1978 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Lagat 2009 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Genus Gekko (Tokays)   * Gekko carusadensis Luzon Karst Gecko Linkem, Siler, Diesmos, Sy & Brown 2010   Gekko gecko Tokay Brown & Alcala 1978 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Lagat 2009 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011   Gekko kikuchii Lanyu Gecko Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013   Gekko mindorensis Mindoro Narrow-disked Gecko Brown, Ferner, Sison, Gonzales & Kennedy 1996 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Gekko monarchus Spotted House Gecko Alcala 1986 Brown & Alcala 1978 Lagat 2009 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus frenatus Asian House Gecko Brown & Alcala 1978 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Lagat 2009 Lajmi, Bansal, Giri & Karanth 2019 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Hemidactylus platyurus Flat-tailed Gecko Brown & Alcala 1978 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Lagat 2009 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Hemidactylus stejnegeri Taiwanese Half-toed Gecko Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Lagat 2009 Zhao & Adler 1993 Genus Lepidodactylus (Scaly-toed Geckos)   * Lepidodactylus bakingibut Ilocano Scaly-toed Gecko Eliades, Brown, Huang & Siler 2021   * Lepidodactylus bisakol Bicol Scaly-toed Gecko Eliades, Brown, Huang & Siler 2021   Lepidodactylus lugubris Common Mourning Gecko Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 [as "cf. lugubris"] Eliades, Brown, Huang & Siler 2021 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 [as "L. cf. lugubris"] McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011   Lepidodactylus planicauda Broad-tailed Scaly-toed Gecko Devan-Song & Brown 2012 [as "cf. planicaudus"] Genus Luperosaurus (Flap-legged Geckos)   * Luperosaurus angliit Aurora Flap-legged Gecko Brown, Diesmos & Oliveiros 2011   * Luperosaurus cumingii Southern Luzon Flap-legged Gecko Alcala 1986 Brown & Alcala 1978 Brown, Supriatna & Ota 2000   * Luperosaurus kubli Mount Lataan Flap-legged Gecko Brown, Diesmos & Duya 2007 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 [as "cf. kubli"] Genus Pseudogekko (Smooth-scaled Geckos)   Pseudogekko compressicorpus Flat-bodied Smooth-scaled Gecko Alcala 1986 Brown & Alcala 1978 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Davis, Watters, Koehler, Whitsett, Huron, Brown, Diesmos & Siler 2015 Ross & Lazell 1990 Siler, Welton, Davis, Watters, Davey, Diesmos, Diesmos & Brown 2014   * Pseudogekko hungkag Mount Bulusan Smooth-scaled Gecko Brown, Meneses, Wood, Fernandez, Cuesta, Clores, Tracy, Buehler & Siler 2020   Pseudogekko smaragdinus Emerald Smooth-scaled Gecko Davis, Watters, Koehler, Whitsett, Huron, Brown, Diesmos & Siler 2015 Siler, Dececchi, Merkord, Davis, Christiani & Brown 2014   * Pseudogekko sumiklab Bicol Smooth-scaled Gecko Siler, Davis, Watters, Freitas, Griffith, Binaday, Lobos, Amarga & Brown 2017  Family Scincidae (Skinks) Genus Brachymeles (Indo-Malayan Slender Skinks)   * Brachymeles bicolandia Central Bicol Slender Skink Siler, Fuiten, Jones, Alcala & Brown 2011 Siler, Jones, Diesmos, Diesmos & Brown 2012   * Brachymeles bicolor Elongate Slender Skink Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Siler & Brown 2010 Siler, Crombie, Diesmos & Brown 2011 Siler, Diesmos & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Brachymeles bonitae Stub-limbed Slender Skink Alcala 1986 Brown & Alcala 1980 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Davis, Feller, Brown & Siler 2014 Lagat 2009 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Schaper, Bergmann, Morinaga, Fernandez, Freitas & Siler 2018 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Brachymeles boulengeri Polillo Slender Skink Alcala 1986 Brown & Alcala 1980 Lagat 2009 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   * Brachymeles brevidactylus Southern Bicol Slender Skink Siler, Fuiten, Jones, Alcala & Brown 2011 Siler, Jones, Diesmos, Diesmos & Brown 2012   * Brachymeles elerae Four-toed Slender Skink Brown & Alcala 1980 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Siler & Brown 2010 Siler 2010 Siler, Diesmos & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012 Siler, Jones, Welton & Brown 2011   Brachymeles ilocandia Ilokano Slender Skink Siler, Davis, Freitas, Huron, Geheber, Watters, Penrod, Papes, Amrein, Anwar, Cooper, Hein, Manning, Patel, Pinaroc, Diesmos, Diesmos, Oliveros & Brown 2016   * Brachymeles isangdaliri Aurora Slender Skink Davis, Feller, Brown & Siler 2014   Brachymeles kadwa Zabali Slender Skink Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   * Brachymeles lukbani Luzon Limbless Slender Skink Siler & Brown 2010 Siler, Balete, Diesmos & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012   Brachymeles makusog Tungaw Slender Skink Siler & Brown 2010 Siler, Diesmos & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012   * Brachymeles muntingkamay Caraballo Mountains Slender Skink Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012 Siler, Rico, Duya & Brown 2009 Siler, Welton, Rock & Brown 2013   * Brachymeles wrighti Benguet Slender Skink Alcala 1986 Brown & Alcala 1980 Siler, Jones, Diesmos, Diesmos & Brown 2012 Siler, Jones, Welton & Brown 2011 Genus Dasia (Indo-Malayan Tree Skinks)   Dasia grisea Grey Tree Skink Brown & Alcala 1980 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Lagat 2009 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Brown & Alcala 1980 Brown 1991 Devan-Song & Brown 2012 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Genus Eutropis (South Asian Sun Skinks)   Eutropis bontocensis Luzon Mountain Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020 Brown & Alcala 1980   Eutropis borealis Subic Bay Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020 Brown & Alcala 1980 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Lagat 2009 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Eutropis cumingi Zambales Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020 Brown & Alcala 1980 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Eutropis gubataas Upland Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020   Eutropis indeprensa Mindoro Sun Skink David, Pauwels, Lays & Lenglet 2006   Eutropis lapulapu Mactan Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020   Eutropis multifasciata Many-lined Sun Skink Brown & Alcala 1980 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Lagat 2009 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 O'Shea, Kusuma & Kaiser 2018 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Genus Lamprolepis (Malay Archipelago Tree Skinks)   Lamprolepis smaragdina Emerald Tree Skink Brown & Alcala 1980 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gaulke 1996 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Lagat 2009 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Mertens 1929 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Genus Lipinia (Indo-Pacific Striped Skinks)   Lipinia auriculata Bronze Striped Tree Skink Supsup, Redoblado & Guinto 2016   Lipinia pulchella Yellow-striped Slender Tree Skink Alcala 1986 Brown & Alcala 1956 Brown & Alcala 1980 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Lagat 2009 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Supsup, Redoblado & Guinto 2016   Lipinia vulcania Dapitan Peak Striped Skink Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 [as "cf. vulcania"] Genus Otosaurus (Philippine Giant Forest Skink)   Otosaurus cumingii Philippine Giant Forest Skink Alcala 1986 Brown & Alcala 1980 Brown & Rico 2009 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Lagat 2009 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Genus Parvoscincus (Philippine Dwarf Forest Skinks)   Parvoscincus abstrusus Mount Makiling Dwarf Forest Skink Linkem & Brown 2013   * Parvoscincus agtorum Agta Dwarf Forest Skink Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Linkem & Brown 2013   * Parvoscincus arvindiesmosi Southern Luzon Dwarf Forest Skink Linkem & Brown 2013   * Parvoscincus aurorus Aurora Dwarf Forest Skink Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Linkem & Brown 2013   * Parvoscincus banahaoensis Mount Banahao Dwarf Forest Skink Linkem & Brown 2013   * Parvoscincus beyeri Lalo Dwarf Forest Skink Alcala 1986 Brown & Alcala 1980 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018   * Parvoscincus boyingi Zambales Dwarf Forest Skink Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010   Parvoscincus decipiens Black-sided Dwarf Forest Skink Alcala 1986 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Lagat 2009 Linkem & Brown 2013 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   * Parvoscincus duwendorum Mount Pao Dwarf Forest Skink Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Siler, Linkem, Cobb, Watters, Cummings, Diesmos & Brown 2014   * Parvoscincus hadros Mount Mingan Dwarf Forest Skink Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010   * Parvoscincus igorotorum Igorot Mountains Dwarf Forest Skink Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010   * Parvoscincus jimmymcguirei Mount Cagua Dwarf Forest Skink Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Linkem & Brown 2013   Parvoscincus laterimaculatus Bicol Peninsula Dwarf Forest Skink Brown & Alcala 1980 Brown, Ferner & Ruedas 1995 [as knollmanae] Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010   Parvoscincus lawtoni Balbalan Dwarf Forest Skink Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010   * Parvoscincus leucospilos White-spotted Dwarf Forest Skink Alcala 1986 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Linkem, Cobb, Watters, Cummings, Diesmos & Brown 2014 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   * Parvoscincus luzonensis Highland Dwarf Forest Skink Alcala 1986 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010   * Parvoscincus manananggalae Lipimental Dwarf Forest Skink Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Siler, Linkem, Cobb, Watters, Cummings, Diesmos & Brown 2014   * Parvoscincus palaliensis Mount Palali Dwarf Forest Skink Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Linkem & Brown 2013   Parvoscincus steerei Guimaras Dwarf Forest Skink Brown & Alcala 1980 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Lagat 2009 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   * Parvoscincus tagapayo Kabatangan River Dwarf Forest Skink Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Brown, McGuire, Ferner & Alcala 1999 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   * Parvoscincus tikbalangi Sierra Madres Dwarf Forest Skink Siler, Linkem, Cobb, Watters, Cummings, Diesmos & Brown 2014 Genus Pinoyscincus (Philippine Forest Skinks)   Pinoyscincus abdictus Camiguin Forest Skink Brown & Alcala 1980 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Pinoyscincus coxi Agusan Forest Skink Alcala 1986 Brown & Alcala 1980 Lagat 2009   Pinoyscincus jagorii Samar Forest Skink Brown & Alcala 1980 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Lagat 2009 Ross & Lazell 1990 Genus Tropidophorus (Stream Skinks)   Tropidophorus grayi Philippine Stream Skink Alcala 1986 Brown & Alcala 1980  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   * Varanus bitatawa Northern Luzon Forest Monitor Abano-Sarigumba, Brown, Welton & Ibanez 2018 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 [as "V. cf. bitatawa"] Meneses, Alviola, Parcon, Cosico, Lumibao, Gonzalez & Brown 2020 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Welton, Siler, Bennett, Diesmos, Duya, Dugay, Rico, Van Weerd & Brown 2010   Varanus dalubhasa Bicol Water Monitor Welton, Travers, Siler & Brown 2014   Varanus marmoratus Marbled Water Monitor Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gaulke 1996 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Koch, Auliya, Schmitz, Kuch & Boehme 2007 Koch, Gaulke & Boehme 2010 Lagat 2009 Lisle 1996 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Mertens 1963 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Welton, Travers, Siler & Brown 2014   Varanus olivaceus Olive Monitor Auffenberg 1976 [as V. grayi] Auffenberg 1988 Boehme 2003 Lisle 1996 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Pianka in Pianka, King & King (eds.) 2004

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Acrochordidae (Australasian File Snakes) Genus Acrochordus (Australasian File Snakes)   Acrochordus granulatus Lesser File Snake Bernstein & Bautista 2020 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Iskandar & Colijn 2001 McDiarmid, Campbell & Toure 1999 Wallach, Williams & Boundy 2014  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake Alcala 1986 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Iskandar & Colijn 2001 Leviton 1967 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Wallach, Williams & Boundy 2014 Genus Boiga (Australasian Cat Snakes)   Boiga angulata Leyte Cat Snake Alcala 1986 Devan-Song & Brown 2012 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Orlov & Ryabov 2002   Boiga cynodon Dog-toothed Cat Snake Alcala 1986 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gaulke 1996 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Orlov & Ryabov 2002 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Boiga dendrophila Eastern Mangrove Cat Snake Alcala 1986 Brongersma 1934 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Orlov & Ryabov 2002 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Boiga drapiezii White-spotted Cat Snake Binaday & Lobos 2016 Leviton, Siler, Weinell & Brown 2018   Boiga philippina Luzon Cat Snake Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Iskandar & Colijn 2001 Kroon 1973 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Orlov & Ryabov 2002 Genus Calamaria (Oriental Reed Snakes)   Calamaria bitorques Banded Reed Snake Alcala 1986 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Inger & Marx 1965 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Calamaria gervaisi Philippine Reed Snake Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Inger & Marx 1965 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea paradisi Paradise Flying Tree Snake Brown, Ferner, Sison, Gonzales & Kennedy 1996 Devan-Song & Brown 2012 Iskandar & Colijn 2001 Leviton 1964 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Mertens 1968 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus erythrura Philippine Rat Snake Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Iskandar & Colijn 2001 Leviton 1979 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Schulz 1996 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis luzonensis Luzon Stripe-tailed Bronzeback Alcala 1986 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Iskandar & Colijn 2001 Leviton 1961 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Rooijen & Vogel 2012 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Dendrelaphis marenae Philippine Painted Bronzeback Brown, Ferner, Sison, Gonzales & Kennedy 1996 [as pictus] Brown, McGuire, Ferner, Icarangal & Kennedy 2000 [as pictus] Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Iskandar & Colijn 2001 [as pictus] Leviton 1968 [as pictus] Leviton, Siler, Weinell & Brown 2018 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 [as pictus] Vogel & Rooijen 2008   Dendrelaphis philippinensis Mindanao Stripe-tailed Bronzeback Leviton, Siler, Weinell & Brown 2018 Rooijen & Vogel 2012 Genus Dryophiops (Malayan Vine Snakes)   Dryophiops philippina Philippine Vine Snake Alcala 1986 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Iskandar & Colijn 2001 Leviton 1964 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma oxycephalum Red-tailed Green Rat Snake Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Schulz 1996 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Genus Lycodon (Oriental Wolf Snakes)   Lycodon capucinus Island Wolf Snake Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Iskandar & Colijn 2001 Leviton 1965 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 O'Shea, Kusuma & Kaiser 2018 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Lycodon muelleri Luzon Wolf Snake Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Iskandar & Colijn 2001 Lanza 1999 Leviton 1965 Leviton, Siler, Weinell & Brown 2018 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   * Lycodon solivagus Pale-banded Wolf Snake Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Iskandar & Colijn 2001 Lanza 1999 Leviton, Siler, Weinell & Brown 2018 Ota & Ross 1994   * Lycodon tessellatus Manila Wolf Snake Alcala 1986 Iskandar & Colijn 2001 Lanza 1999 Genus Oligodon (Kukri Snakes)   Oligodon ancorus Luzon Kukri Snake Alcala 1986 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Supsup 2016   Oligodon modestus Spotted-bellied Kukri Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Genus Pseudorabdion (Dwarf Reed Snakes)   Pseudorabdion mcnamarae Negros Dwarf Reed Snake Alcala 1986 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 [as "cf. mcnamarae"] Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 [as "P. cf. mcnamarae"] Iskandar & Colijn 2001 Leviton & Brown 1959 Leviton 1963 Leviton, Siler, Weinell & Brown 2018   Pseudorabdion talonuran Panay Dwarf Reed Snake Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 [as "cf. talonuran"] Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas luzonensis Smooth-scaled Mountain Rat Snake Alcala 1986 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Iskandar & Colijn 2001 Leviton 1963 Leviton 1983 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Genus Rhabdophis (Asian Keelbacks)   * Rhabdophis barbouri Luzon Keelback Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Genus Tropidonophis (Indo-Australian Keelbacks)   Tropidonophis dendrophiops Mindanao Keelback Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018   Tropidonophis spilogaster Northern Philippine Keelback Alcala 1986 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Doria, Petri, Bellati, Tiso & Pistarino 2013 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Zhu, Wang, Takeuchi & Zhao 2014  Family Cyclocoridae (Philippine Burrowing Snakes, etc.) Genus Cyclocorus (Triangle-spotted Snakes)   Cyclocorus lineatus Northern Triangle-spotted Snake Alcala 1986 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Iskandar & Colijn 2001 Leviton 1965 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Genus Hologerrhum (Philippine Cylindrical Snakes)   Hologerrhum philippinum Luzon Cylindrical Snake Alcala 1986 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Leviton, Ferner & Sison 2001 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Genus Myersophis (Luzon Alpine Snake)   * Myersophis alpestris Luzon Alpine Snake Alcala 1986 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Taylor 1963 Genus Oxyrhabdium (Philippine Burrowing Snakes)   Oxyrhabdium leporinum Philippine Banded Burrowing Snake Alcala 1986 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gaulke & Operiano 2006 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Iskandar & Colijn 2001 Leviton 1957 Leviton 1964 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Calliophis (South Asian Coral Snakes)   Calliophis philippinus Philippine Striped Coral Snake Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Genus Hemibungarus (Barred Coral Snakes)   Hemibungarus calligaster Luzon Barred Coral Snake Alcala 1986 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Brown, Smart, Leviton & Smith 2018 Devan-Song & Brown 2012 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011   Hemibungarus mcclungi Polillo Barred Coral Snake Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Genus Naja (Typical Cobras)   Naja philippinensis Northern Philippine Cobra Alcala 1986 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Iskandar & Colijn 2001 Leviton 1964 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Wuster & Thorpe 1991 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Alcala 1986 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Iskandar & Colijn 2001 Leviton 1964 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Sy, Baniqued & Diesmos 2016 Sy, Layola, Yu & Diesmos 2015 Urriza, Elec & Palacpac 2017  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis atriceps Black-headed Sea Snake Herre 1942 Leviton, Siler, Weinell & Brown 2018 Smith 1926   Hydrophis curtus Short Sea Snake Brown, Ferner, Sison, Gonzales & Kennedy 1996 [as Lapemis hardwickii] Devan-Song & Brown 2012 [as Lapemis hardwickii] Herre 1942 [as Lapemis hardwickii] Leviton, Siler, Weinell & Brown 2018 Smith 1926 [as Lapemis hardwickii]   Hydrophis cyanocinctus Blue-banded Sea Snake Alcala 1986 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Smith 1935   Hydrophis lamberti Siamese Sea Snake Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018   Hydrophis ornatus Reef Sea Snake Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018   Hydrophis platurus Yellow-bellied Sea Snake Leviton, Siler, Weinell & Brown 2018   * Hydrophis semperi Lake Taal Sea Snake Alcala 1986 Bucherl, Buckley & Deulofeu (eds.) 1968 Herre 1942 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Smith 1926 Genus Laticauda (Sea Kraits)   Laticauda colubrina Yellow-lipped Sea Krait Brown, Ferner, Sison, Gonzales & Kennedy 1996 Devan-Song & Brown 2012 Heatwole, Busack & Cogger 2005 Iskandar & Colijn 2001 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018   Laticauda laticaudata Blue-lipped Sea Krait Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Smith 1926   Laticauda semifasciata Erabu Sea Krait Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Genus Microcephalophis (Small-headed Sea Snakes)   Microcephalophis gracilis Graceful Small-headed Sea Snake Leviton, Siler, Weinell & Brown 2018  Family Gerrhopilidae (Indo-Malayan Blind Snakes) Genus Gerrhopilus (Indo-Malayan Blind Snakes)   Gerrhopilus hedraeus Negros Blind Snake Iskandar & Colijn 2001 McDiarmid, Campbell & Toure 1999 Wallach 1996   Gerrhopilus manilae Manila Blind Snake Leviton, Siler, Weinell & Brown 2018  Family Homalopsidae (Australasian Mud Snakes) Genus Cerberus (Dog-faced Water Snakes)   * Cerberus microlepis Lake Buhi Dog-faced Water Snake Gyi 1970 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Murphy & Voris 2014 Murphy 2007 Murphy, Voris & Karns 2012   Cerberus schneiderii South East Asian Dog-faced Water Snake Brown, Ferner, Sison, Gonzales & Kennedy 1996 [as C. rynchops] Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 [as C. rynchops] Dunson & Minton 1978 [as C. rynchops] Gyi 1970 [as C. rynchops] Iskandar & Colijn 2001 [as C. rynchops] Leviton, Siler, Weinell & Brown 2018  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Iskandar & Colijn 2001 Leviton 1963 Leviton 1983 Leviton, Siler, Weinell & Brown 2018 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011  Family Pareidae (Oriental Slug Eaters) Genus Aplopeltura (Blunt-headed Slug Eater)   Aplopeltura boa Blunt-headed Slug Eater Poyarkov, Nguyen, Pawangkhanant, Yushchenko, Brakels, Nguyen, Nguyen, Suwannapoom, Orlov & Vogel 2022 Sy & Binaday 2016  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gaulke 1996 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011  Family Typhlopidae (Typical Blind Snakes) Genus Acutotyphlops (Sharp-nosed Blind Snakes)   * Acutotyphlops banaorum Balbalan Blind Snake Leviton, Siler, Weinell & Brown 2018 Wallach, Brown, Diesmos & Gee 2007 Wallach, Williams & Boundy 2014 Genus Malayotyphlops (Malay Archipelago Blind Snakes)   * Malayotyphlops andyi Cagayan Blind Snake Leviton, Siler, Weinell & Brown 2018 Wynn, Diesmos & Brown 2016   * Malayotyphlops collaris Collared Blind Snake Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 Wynn & Leviton 1993 Wynn, Diesmos & Brown 2016   * Malayotyphlops denrorum Isabela Blind Snake Leviton, Siler, Weinell & Brown 2018 Wynn, Diesmos & Brown 2016   Malayotyphlops luzonensis Luzon Blind Snake Alcala 1986 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Hahn 1980 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Wynn, Diesmos & Brown 2016   Malayotyphlops ruber Samar Red Blind Snake Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999   Malayotyphlops ruficauda Red-tailed Blind Snake Alcala 1986 [as jagorii] Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Hahn 1980 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Wynn, Diesmos & Brown 2016  Family Viperidae (Vipers) Genus Parias (Malay Archipelago Pitvipers)   Parias flavomaculatus Yellow-spotted Pitviper Alcala 1986 Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Gojo Cruz, Afuang, Gonzalez & Gruezo 2018 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Iskandar & Colijn 2001 Leviton 1964 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011 Vogel 2006 Genus Tropidolaemus (Temple Pitvipers)   Tropidolaemus subannulatus North Philippine Temple Pitviper Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Gaulke 1996 [as wagleri] Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 [as wagleri] Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 [as wagleri] Leviton 1964 [as wagleri] Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Vogel 2006 Vogel, David, Lutz, Rooijen & Vidal 2007

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Sagun, Ramoso & Mejino 2005 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle Brown, Ferner, Sison, Gonzales & Kennedy 1996 Brown, McGuire, Ferner, Icarangal & Kennedy 2000 Brown, Oliveros, Siler, Fernandez, Welton, Buenavente, Diesmos & Diesmos 2012 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Devan-Song & Brown 2012 Diesmos, Brown, Alcala & Sison 2008 McLeod, Siler, Diesmos, Diesmos, Garcia, Arkonceo, Balaquit, Uy, Villaseran, Yarra & Brown 2011 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Siler, Welton, Siler, Brown, Bucol, Diesmos & Brown 2011  Family Trionychidae (Softshell Turtles) Genus Dogania (Malayan Softshell Turtle)   Dogania subplana Malayan Softshell Turtle Diesmos, Brown, Alcala & Sison 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelochelys (Indomalayan Giant Softshell Turtles)   Pelochelys cantorii Asian Giant Softshell Turtle Bonin, Devaux & Dupre 2006 Brown, Siler, Oliveiros, Welton, Rock, Swab, Weerd, Beijnen, Jose, Rodriguez, Jose & Diesmos 2013 Diesmos, Brown, Alcala & Sison 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021