Reptiles of Yemen  Related checklists:  Yemeni mainland - Hanish Islands - Socotra Archipelago Bibliography of the reptiles of Yemen Taxonomic history of the reptiles of Yemen Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Trogonophidae (Short-headed Worm Lizards) Genus Agamodon (Chisel-headed Worm Lizards)   * Agamodon arabicus Arabian Chisel-headed Worm Lizard Arnold 1986 Gans 2005 Schatti & Desvoignes 1999 Sindaco & Jeremchenko 2008 Genus Pachycalamus (Socotra Worm Lizard)   * Pachycalamus brevis Socotra Worm Lizard Gans 1967 Gans 2005 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wranik 1998

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Acanthocercus (Ridgeback Agamas)   Acanthocercus adramitanus Hadhramaut Ridgeback Agama Fritz & Schutte 1988 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Showler 1995 Sindaco & Jeremchenko 2008   Acanthocercus yemenensis Yemen Ridgeback Agama Arnold 1986 Fritz & Schutte 1988 Schatti & Gasperetti 1994 Schatti 1989 Showler 1995 Sindaco & Jeremchenko 2008 Genus Phrynocephalus (Toad-headed Agamas)   Phrynocephalus arabicus Arabian Toad-headed Agama Melnikov, Melnikova, Nazarov, Rajabizadeh, Al-Johany, Amr & Ananjeva 2014 Schatti & Desvoignes 1999 Showler 1995   Phrynocephalus longicaudatus Long-tailed Toad-headed Agama Schatti & Desvoignes 1999 [as maculatus] Genus Pseudotrapelus (Afro-Arabian Desert Agamas)   * Pseudotrapelus neumanni Yemen Desert Agama Arnold 1986 [as sinaitus] Fritz & Schutte 1988 [as sinaitus] Melnikov, Ananjeva & Papenfuss 2013 Schatti & Desvoignes 1999 [as sinaitus] Schatti & Gasperetti 1994 [as sinaitus] Schatti 1989 [as sinaitus] Showler 1995 [as sinaitus] Tamar, Scholz, Crochet, Geniez, Meiri, Schmitz, Wilms & Carranza 2016 Genus Trapelus (Afro-Asian Ground Agamas)   Trapelus flavimaculatus Yellow-spotted Agama Fritz & Schutte 1988 Rastegar-Pouyani 1998 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Genus Uromastyx (Western Spiny-tailed Agamas)   Uromastyx aegyptia Egyptian Spiny-tailed Agama Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Showler 1995 Wilms & Boehme 2000 Wilms & Boehme 2007 Wilms 2001 Wilms, Boehme, Wagner, Lutzmann & Schmitz 2009   Uromastyx benti South Arabian Spiny-tailed Agama Arnold 1986 Schatti 1989 Wilms & Boehme 2000 Wilms & Boehme 2007 Wilms 2001 Wilms, Boehme, Wagner, Lutzmann & Schmitz 2009   Uromastyx ornata Ornate Spiny-tailed Agama Arnold 1986 Schatti & Desvoignes 1999 [as ocellata] Schatti & Gasperetti 1994 [as ocellata] Wilms & Boehme 2000 Wilms & Boehme 2007   * Uromastyx shobraki Western Yemen Spiny-tailed Agama Wilms & Boehme 2007 Wilms, Boehme, Wagner, Lutzmann & Schmitz 2009   * Uromastyx yemenensis Southern Yemen Spiny-tailed Agama Wilms & Boehme 2007 Wilms & Schmitz 2007 Wilms, Boehme, Wagner, Lutzmann & Schmitz 2009  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo arabicus Arabian Chameleon Arnold 1986 Glaw 2015 Klaver & Boehme 1997 Meerman & Boomsma 1987 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Showler 1995 Sindaco & Jeremchenko 2008 Tilbury 2010   Chamaeleo calyptratus Veiled Chameleon Arnold 1986 Glaw 2015 Klaver & Boehme 1997 Schatti & Desvoignes 1999 Schatti 1989 Showler 1995 Sindaco & Jeremchenko 2008 Tilbury 2010   Chamaeleo chamaeleon Mediterranean Chameleon Arnold 1986 Glaw 2015 Klaver & Boehme 1997 Schatti & Gasperetti 1994 Schatti 1989 Tilbury 2010   * Chamaeleo monachus Socotran Chameleon Glaw 2015 Klaver & Boehme 1997 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Showler 1996 Sindaco, Venchi & Grieco 2013 Tilbury 2010 Wranik 1998  Family Gekkonidae (Typical Geckos) Genus Bunopus (South West Asian Tuberculated Geckos)   Bunopus tuberculatus Southern Tuberculated Gecko Schatti & Desvoignes 1999 Schatti 1989 Showler 1995 Szczerbak & Golubev 1996 Genus Cyrtopodion (Palearctic Bent-toed Geckos)   Cyrtopodion scabrum Keeled Rock Gecko Arnold 1986 Roesler & Wranik 1998 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Genus Hemidactylus (Half-toed Geckos)   * Hemidactylus adensis Aden Half-toed Gecko Smid, Moravec, Kratochvil, Nasher, Masuch, Gvozdik & Carranza 2015   Hemidactylus alkiyumii Tawi Atair Half-toed Gecko Carranza & Arnold 2012 Sindaco, Venchi & Grieco 2013   * Hemidactylus dracaenacolus Dragon Tree Half-toed Gecko Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 1999 Roesler & Wranik 2004 Sindaco, Venchi & Grieco 2013   Hemidactylus festivus Wadi Ayoun Half-toed Gecko Carranza & Arnold 2012 Sindaco, Venchi & Grieco 2013   Hemidactylus flaviviridis Yellow-bellied House Gecko Fritz & Schutte 1987 Roesler & Wranik 1998 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Schmidt 1953 Showler 1995   * Hemidactylus forbesii Abd al Kuri Half-toed Gecko Loveridge 1947 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wermuth 1965 Wranik 1998   * Hemidactylus granti Socotra Half-toed Gecko Loveridge 1947 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wranik 1998   * Hemidactylus homoeolepis Socotra Even-scaled Gecko Carranza & Arnold 2012 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 1998 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Showler 1996 Sindaco & Jeremchenko 2008 Sindaco, Venchi & Grieco 2013 Wranik 1998   * Hemidactylus inintellectus Socotra Rock Gecko Razzetti, Sindaco, Grieco & al. 2011 Sindaco, Venchi & Grieco 2013 Sindaco, Ziliani, Razzetti, Carugati, Grieco, Pupin, Al-Aseily, Pella & Fasola 2009   * Hemidactylus jumailiae Ibb Half-toed Gecko Busais & Joger 2011 Sindaco, Venchi & Grieco 2013   Hemidactylus lemurinus Dhofar Half-toed Gecko Carranza & Arnold 2012 Schatti & Desvoignes 1999   * Hemidactylus mandebensis Taizz Half-toed Gecko Smid, Moravec, Kratochvil, Nasher, Masuch, Gvozdik & Carranza 2015   Hemidactylus minutus Dwarf Half-toed Gecko Arnold 1986 [as H. homoeolepis] Carranza & Arnold 2012 [as H. homoeolepis] Schatti & Desvoignes 1999 [as H. homoeolepis] Schatti & Gasperetti 1994 [as H. homoeolepis] Sindaco & Jeremchenko 2008 [as H. homoeolepis] Vasconcelos & Carranza 2014   Hemidactylus montanus Mountain Half-toed Gecko Busais & Joger 2011   * Hemidactylus oxyrhinus Sharp-nosed Half-toed Gecko Loveridge 1947 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wermuth 1965 Wranik 1998   * Hemidactylus pumilio Pygmy Half-toed Gecko Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wranik 1998   Hemidactylus robustus Red Sea Half-toed Gecko Arnold 1986 [as turcicus] Busais & Joger 2011 Carranza & Arnold 2012 Fritz & Schutte 1987 [as turcicus parkeri] Roesler & Wranik 1998 [as turcicus] Schatti & Desvoignes 1999 [as turcicus] Schatti & Gasperetti 1994 [as turcicus] Schatti 1989 [as turcicus] Schmidt 1953 [as turcicus] Showler 1995 [as turcicus]   * Hemidactylus saba Sheba Half-toed Gecko Busais & Joger 2011 Sindaco, Venchi & Grieco 2013 Smid, Moravec, Kratochvil, Gvozdik, Nasher, Busais, Wilms, Shobrak & Carranza 2013   * Hemidactylus shihraensis Al Shihr Half-toed Gecko Busais & Joger 2011 Sindaco, Venchi & Grieco 2013   Hemidactylus sinaitus Sudan Half-toed Gecko Arnold 1986 Busais & Joger 2011 Carranza & Arnold 2012 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Sindaco & Jeremchenko 2008 Smid, Mazuch, Novakova, Modry, Malonza, Elmi, Carranza & Moravec 2020   * Hemidactylus ulii Al Hababi Half-toed Gecko Smid, Moravec, Kratochvil, Gvozdik, Nasher, Busais, Wilms, Shobrak & Carranza 2013   * Hemidactylus yerburii Lahej Half-toed Gecko Arnold 1986 Fritz & Schutte 1987 Roesler & Wranik 1998 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Schmidt 1953 Showler 1995 Sindaco & Jeremchenko 2008 Genus Stenodactylus (Afro-Asian Thin-toed Geckos)   Stenodactylus doriae Middle Eastern Thin-toed Gecko Arnold 1980 Schatti & Desvoignes 1999 Showler 1995   Stenodactylus leptocosymbotes Eastern Arabian Thin-toed Gecko Arnold 1980 Arnold 1986 Leviton, Anderson, Adler & Minton 1992 Schatti & Desvoignes 1999 Sindaco & Jeremchenko 2008   Stenodactylus slevini Dharan Thin-toed Gecko Arnold 1980 Schatti & Desvoignes 1999   Stenodactylus yemenensis Yemen Thin-toed Gecko Arnold 1980 Arnold 1986 Roesler & Wranik 1998 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Showler 1995 Sindaco & Jeremchenko 2008 Genus Trachydactylus (Arabian Tuberculated Geckos)   Trachydactylus spatalurus South Arabian Tuberculated Gecko Arnold 1986 Gardner 2013 Pous, Machado, Metallinou, Cervenka, Kratochvil, Paschou, Mazuch, Smid, Simo-Riudalbas, Sanuy & Carranza 2016 Roesler & Wranik 1998 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Showler 1995 Sindaco & Jeremchenko 2008 Genus Trigonodactylus (Triangle-scaled Geckos)   Trigonodactylus arabicus Arabian Triangle-scaled Gecko Arnold 1980 Metallinou & Carranza 2013 Schatti & Desvoignes 1999 Sindaco & Jeremchenko 2008   * Trigonodactylus pulcher Yemen Triangle-scaled Gecko Arnold 1980 Arnold 1986 Metallinou & Carranza 2013 Schatti & Desvoignes 1999 Sindaco & Jeremchenko 2008 Genus Tropiocolotes (Afro-Asian Desert Dwarf Geckos)   Tropiocolotes scorteccii Yemen Dwarf Gecko Arnold 1986 Machado, Smid, Mazuch, Sindaco, Al Shukaili & Carranza 2018 Ribeiro Junior, Tamar, Maza, Flecks, Wagner, Shacham, Calvo, Geniez, Crochet, Koch & Meiri 2022 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Showler 1995 Sindaco & Jeremchenko 2008 Szczerbak & Golubev 1996 Wilms, Shobrak & Wagner 2010  Family Lacertidae (Typical Lizards) Genus Acanthodactylus (Fringe-toed Lizards)   * Acanthodactylus arabicus Arabian Fringe-toed Lizard Arnold 1986 Salvador 1982 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Showler 1995 Sindaco & Jeremchenko 2008   Acanthodactylus boskianus Afro-Arabian Fringe-toed Lizard Salvador 1982 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Schmidt 1953 Showler 1995   Acanthodactylus felicis South Arabian Fringe-toed Lizard Arnold 1983 Arnold 1986 Salvador 1982 Schatti & Desvoignes 1999   Acanthodactylus masirae Masirah Fringe-toed Lizard Salvador 1982   Acanthodactylus opheodurus Snake-tailed Fringe-toed Lizard Salvador 1982 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Showler 1995 Sindaco & Jeremchenko 2008   Acanthodactylus schmidti Middle East Fringe-toed Lizard Salvador 1982 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994   * Acanthodactylus yemenicus Yemen Fringe-toed Lizard Arnold 1986 Salvador 1982 Schatti & Gasperetti 1994 Sindaco & Jeremchenko 2008 Genus Latastia (African Long-tailed Lizards)   Latastia longicaudata Common Long-tailed Lizard Arnold 1986 Bischoff 1991 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schmidt 1953 Genus Mesalina (Afro-Asian Desert Lizards)   Mesalina adramitana Hadramaut Desert Lizard Arnold 1986 Schatti & Desvoignes 1999 Schatti 1989 Sindaco & Jeremchenko 2008 Yousefkhani, Marin & Rastegar-Pouyani 2015   Mesalina arnoldi Yemen Desert Lizard Sindaco, Simo-Riudalbas, Sacchi & Carranza 2018   Mesalina austroarabica South Arabian Desert Lizard Sindaco, Simo-Riudalbas, Sacchi & Carranza 2018   * Mesalina balfouri Socotra Desert Lizard Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2000 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Showler 1994 Showler 1996 Sindaco, Venchi & Grieco 2013 Szczerbak 1975 Wranik 1998 Yousefkhani, Marin & Rastegar-Pouyani 2015   * Mesalina kuri Abd al Kuri Desert Lizard Joger & Mayer 2002 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 [as balfouri] Sindaco, Venchi & Grieco 2013 Wranik 1998 [as balfouri] Yousefkhani, Marin & Rastegar-Pouyani 2015   Mesalina martini Red Sea Desert Lizard Arnold 1986 Schatti & Desvoignes 1999 Sindaco & Jeremchenko 2008 Szczerbak 1975 Yousefkhani, Marin & Rastegar-Pouyani 2015 Genus Ophisops (Snake-eyed Lizards)   Ophisops elbaensis Mount Elba Snake-eyed Lizard Schatti 1989 [as Ophisops sp.] Sindaco & Jeremchenko 2008 Genus Philochortus (Shield-backed Ground Lizards)   Philochortus neumanni Arabian Shield-backed Ground Lizard Arnold 1986 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Schmidt 1953 Sindaco & Jeremchenko 2008  Family Phyllodactylidae (Leaf-toed Geckos) Genus Haemodracon (Socotra Leaf-toed Geckos)   * Haemodracon riebeckii Common Socotra Leaf-toed Gecko Bauer, Good & Branch 1997 Loveridge 1947 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Tamar, Simo-Riudalbas, Garcia-Porta, Santos, Llorente, Vasconcelos & Carranza 2019 Wranik 1998   * Haemodracon trachyrhinus Large-nosed Socotra Leaf-toed Gecko Bauer, Good & Branch 1997 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 1998 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Showler 1996 Sindaco, Venchi & Grieco 2013 Tamar, Simo-Riudalbas, Garcia-Porta, Santos, Llorente, Vasconcelos & Carranza 2019 Wranik 1998 Genus Ptyodactylus (Fan-toed Geckos)   Ptyodactylus hasselquistii Egyptian Fan-toed Gecko Arnold & Gallagher 1977 Fritz & Schutte 1987 Heimes 1987 Metallinou, Cervenka, Crochet, Kratochvil, Wilms, Geniez, Shobrak, Brito & Carranza 2015 Roesler & Wranik 1998 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Showler 1995  Family Scincidae (Skinks) Genus Ablepharus (Eurasian Snake-eyed Skinks)   Ablepharus pannonicus Southwest Asian Snake-eyed Skink Arnold 1986 Grossmann, Kowalski & Schillert 2013 Leviton, Anderson, Adler & Minton 1992 Schatti & Gasperetti 1994 Schatti 1989 Sindaco & Jeremchenko 2008 Genus Chalcides (Cylindrical Skinks)   Chalcides ocellatus Mediterranean Ocellated Skink Arnold 1986 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Schmidt 1953 Showler 1995 Genus Eurylepis (Asian Mole Skinks)   Eurylepis taeniolatus Yellow-bellied Mole Skink Arnold 1986 Leviton, Anderson, Adler & Minton 1992 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Genus Hakaria (Socotra Slender Skink)   * Hakaria simonyi Socotra Slender Skink Loveridge 1935 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wranik 1998 [as socotranus] Genus Scincus (Sand Fish)   Scincus conirostris Persian Sand Fish Arnold & Leviton 1977 Arnold 1986 Leviton, Anderson, Adler & Minton 1992 Schatti & Desvoignes 1999   Scincus hemprichii Western Arabian Sand Fish Arnold & Leviton 1977 Arnold 1986 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schmidt 1953 Sindaco & Jeremchenko 2008 Smid, Uvizl, Shobrak, Salim, AlGethami, Algethami, Alanazi, Alsubaie, Busais & Carranza 2021   Scincus mitranus Eastern Arabian Sand Fish Arnold & Leviton 1977 Arnold 1986 Leviton, Anderson, Adler & Minton 1992 Schatti & Desvoignes 1999 Showler 1995 Sindaco & Jeremchenko 2008 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis brevicollis Short-necked Skink Arnold 1986 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Showler 1995   * Trachylepis cristinae Abd al Kuri Rock Skink Razzetti, Sindaco, Grieco & al. 2011 [as socotrana] Roesler & Wranik 2000 [as socotrana] Roesler & Wranik 2004 [as socotrana] Schatti & Desvoignes 1999 [as socotrana] Sindaco, Metallinou, Pupin, Fasola & Carranza 2012 Sindaco, Venchi & Grieco 2013 Wranik 1998 [as socotrana]   * Trachylepis socotrana Socotra Rock Skink Boulenger 1887 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2000 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Showler 1996 Sindaco, Metallinou, Pupin, Fasola & Carranza 2012 Sindaco, Venchi & Grieco 2013 Wranik 1998   Trachylepis tessellata Tessellated Skink Arnold 1986 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Schmidt 1953 Showler 1995 Sindaco & Jeremchenko 2008  Family Sphaerodactylidae (Least Geckos, etc.) Genus Pristurus (Semaphore Geckos)   * Pristurus abdelkuri Abd al Kuri Semaphore Gecko Arnold 1993 Arnold 2009 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wranik 1998   Pristurus carteri Masirah Semaphore Gecko Arnold 1986 Arnold 1993 Arnold 2009 Schatti & Desvoignes 1999 Sindaco & Jeremchenko 2008   * Pristurus collaris Collared Semaphore Gecko Arnold 1986 Arnold 1993 Arnold 2009 Roesler & Wranik 1998 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Showler 1995 Sindaco & Jeremchenko 2008   Pristurus crucifer Cross-marked Semaphore Gecko Arnold 1986 Arnold 1993 Arnold 2009 Fritz & Schutte 1987 Roesler & Wranik 1998 Schatti & Desvoignes 1999 Schmidt 1953 Showler 1995 Sindaco & Jeremchenko 2008   Pristurus flavipunctatus Yellow-spotted Semaphore Gecko Arnold 1986 Arnold 1993 Arnold 2009 Fritz & Schutte 1987 Roesler & Wranik 1998 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Showler 1995 Sindaco & Jeremchenko 2008   * Pristurus guichardi Orange-spotted Semaphore Gecko Arnold 1993 Arnold 2009 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wranik 1998   * Pristurus insignis Socotra Crestless Semaphore Gecko Arnold 1993 Arnold 2009 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Showler 1996 Sindaco, Venchi & Grieco 2013 Wranik 1998   * Pristurus insignoides Adho Dimellus Semaphore Gecko Arnold 1993 Arnold 2009 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wranik 1998   * Pristurus mazbah Mazbah Semaphore Gecko Sindaco & Jeremchenko 2008   Pristurus minimus Lesser Semaphore gecko Schatti & Desvoignes 1999 Showler 1995 Showler 1996 Sindaco & Jeremchenko 2008 Tamar, Mitsi, Simo-Riudalbas, Tejero-Cicuendez, Al-Sariri & Carranza 2019   * Pristurus obsti Orange-lined Semaphore Gecko Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 1999 Roesler & Wranik 2004 Sindaco, Venchi & Grieco 2013   * Pristurus ornithocephalus Bird-headed Semaphore gecko Arnold 1986 Arnold 1993 Arnold 2009 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Showler 1995 Sindaco & Jeremchenko 2008   Pristurus popovi Asir Semaphore Gecko Arnold 1986 Arnold 1993 Arnold 2009 Fritz & Schutte 1987 Sindaco & Jeremchenko 2008   Pristurus rupestris Rock Semaphore Gecko Arnold 1993 Arnold 2009 Roesler & Wranik 1998 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Schmidt 1953 Showler 1995   * Pristurus samhaensis Samha Semaphore Gecko Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2000 Roesler & Wranik 2004 Sindaco, Venchi & Grieco 2013   * Pristurus schneideri Hanish al Kabir Island Semaphore Gecko Arnold 2009 Roesler, Koehler & Boehme 2008 Sindaco, Venchi & Grieco 2013   * Pristurus sokotranus Socotra Throat-spotted Semaphore Gecko Arnold 1993 Arnold 2009 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 1998 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Showler 1996 Sindaco, Venchi & Grieco 2013 Wranik 1998   * Pristurus saada Yemen Semaphore Gecko Arnold 1986 Arnold 1993 Arnold 2009 Schatti & Gasperetti 1994 Schatti 1989 Sindaco & Jeremchenko 2008  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus griseus Desert Monitor Bayless 2002 Schatti & Desvoignes 1999   Varanus yemenensis Yemen Monitor Boehme 2003 Boehme, Joger & Schatti 1989 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Sindaco & Jeremchenko 2008

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx jayakari Arabian Sand Boa Gasperetti 1988 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994  Family Colubridae (Typical Snakes or Colubrids) Genus Dasypeltis (African Egg Eaters)   Dasypeltis arabica Arabian Egg Eater Bates & Broadley 2018 Gasperetti 1988 [as scabra] Schatti & Desvoignes 1999 [as scabra] Sindaco, Venchi & Grieco 2013 [as scabra] Genus Hemerophis (Socotra Whip Snake)   * Hemerophis socotrae Socotra Whip Snake Corkill & Cochrane 1965 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2000 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Schatti & Utiger 2001 Showler 1996 Sindaco, Venchi & Grieco 2013 Wranik 1998 Genus Lytorhynchus (Awl-headed Snakes)   Lytorhynchus diadema Crowned Awl-headed Snake Gasperetti 1988 Schatti & Desvoignes 1999 Schatti 1989   Lytorhynchus gaddi Iranian Awl-headed Snake Alshammari, Badry, Busais, Ibrahim & El-Abd 2021 Genus Platyceps (Flat-headed Whip Snakes)   Platyceps rhodorachis Wadi Whip Snake Gasperetti 1988 Perry 2012 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Schmidt 1953 Sindaco, Venchi & Grieco 2013   Platyceps variabilis Arabian Whip Snake Gasperetti 1988 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Sindaco, Venchi & Grieco 2013 Genus Rhynchocalamus (Middle East Black-headed Snakes)   Rhynchocalamus arabicus South Arabian Black-headed Snake Gasperetti 1988 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Smid, Martinez, Gebhart, Aznar, Gallego, Goecmen, Pous, Tamar & Carranza 2015 Genus Spalerosophis (Diadem Snakes)   Spalerosophis diadema Common Diadem Snake Gasperetti 1988 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus dhara Arabian Cat Snake Crochet, Rasmussen, Wilms, Geniez, Trape & Boehme 2008 Els, Al-Johany, Amr, Baha-el-Din, Disi, Boehme, Geniez & Werner 2021 Gasperetti 1988 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Naja (Typical Cobras)   Naja arabica Arabian Cobra David & Vogel 2010 Gasperetti 1988 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis platurus Yellow-bellied Sea Snake David & Ineich 1999 Gasperetti 1988 Walczak 1978  Family Lamprophiidae (House Snakes, etc.) Genus Atractaspis (Stiletto Snakes)   Atractaspis andersonii Arabian Stiletto Snake Gasperetti 1988 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Laurent 1950 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wallach, Williams & Boundy 2014 Genus Boaedon (Afro-Arabian House Snakes)   * Boaedon arabicus Arabian House Snake Gasperetti 1988 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Schmidt 1953 Genus Ditypophis (Socotra Night Snake)   * Ditypophis vivax Socotra Night Snake Corkill & Cochrane 1965 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Showler 1996 Sindaco, Venchi & Grieco 2013 Wranik 1998 Genus Malpolon (Montpellier Snakes)   Malpolon moilensis Moila Snake Gasperetti 1988 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis schokari Arabian Sand Snake Gasperetti 1988 Marx 1988 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schatti 1989 Schmidt 1953  Family Leptotyphlopidae (Thread Snakes) Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis burii Arabian Thread Snake Broadley & Wallach 2007 Gasperetti 1988 Hahn 1978 McDiarmid, Campbell & Toure 1999 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013   * Myriopholis filiformis Socotra Slender Thread Snake Hahn 1978 Hahn 1980 McDiarmid, Campbell & Toure 1999 Razzetti, Sindaco, Grieco & al. 2011 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wranik 1998   Myriopholis macrorhyncha Large-snouted Thread Snake Gasperetti 1988 Hahn 1978 Schatti & Desvoignes 1999   * Myriopholis macrura Socotra Long-tailed Thread Snake Hahn 1978 Hahn 1980 McDiarmid, Campbell & Toure 1999 Razzetti, Sindaco, Grieco & al. 2011 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wranik 1998   Myriopholis nursii Aden Thread Snake Broadley & Wallach 2007 Gasperetti 1988 Hahn 1978 McDiarmid, Campbell & Toure 1999 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994   * Myriopholis wilsoni Round-snouted Socotra Thread Snake Hahn 1978 Hahn 1980 McDiarmid, Campbell & Toure 1999 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013   * Myriopholis yemenica Yemen Thread Snake Broadley & Wallach 2007 Sindaco, Venchi & Grieco 2013  Family Typhlopidae (Typical Blind Snakes) Genus Xerotyphlops (Desert Blind Snakes)   * Xerotyphlops socotranus Socotra Blind Snake Hahn 1980 McDiarmid, Campbell & Toure 1999 Razzetti, Sindaco, Grieco & al. 2011 Roesler & Wranik 2004 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013 Wranik 1998  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder David & Ineich 1999 Gasperetti 1988 McDiarmid, Campbell & Toure 1999 Schatti & Desvoignes 1999 Schatti & Gasperetti 1994 Schmidt 1953 Genus Cerastes (Horned Vipers)   Cerastes cerastes Sahara Horned Viper McDiarmid, Campbell & Toure 1999 Schatti & Desvoignes 1999 [as "cf. cerastes"] Schatti & Gasperetti 1994 Werner, Sivan, Kushnir & Motro 1999   Cerastes gasperettii Middle Eastern Horned Viper David & Vogel 2010 Gasperetti 1988 McDiarmid, Campbell & Toure 1999 Schatti & Desvoignes 1999 Schatti 1989 Werner, Verdier, Rosenman & Sivan 1991 Genus Echis (Carpet Vipers)   Echis borkini Yemen Carpet Viper David & Vogel 2010 Pook, Joger, Stumpel & Wuster 2009 Sindaco, Venchi & Grieco 2013   Echis coloratus Painted Carpet Viper Gasperetti 1988 McDiarmid, Campbell & Toure 1999 Schatti & Desvoignes 1999   Echis khosatzkii Dhofar Carpet Viper Pook, Joger, Stumpel & Wuster 2009 Schatti & Desvoignes 1999 Sindaco, Venchi & Grieco 2013

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Roesler & Wranik 2004 Showler 1996 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Gasperetti, Stimson, Miller, Ross & Gasperetti 1993 King & Burke 1989 Pritchard 1979 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Roesler & Wranik 2004 Ross & Barwani in Bjorndal (ed.) 1982 Showler 1994 Showler 1996 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Gasperetti, Stimson, Miller, Ross & Gasperetti 1993 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Roesler & Wranik 2004 Ross & Barwani in Bjorndal (ed.) 1982 Showler 1994 Showler 1996 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Gasperetti, Stimson, Miller, Ross & Gasperetti 1993 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Gasperetti, Stimson, Miller, Ross & Gasperetti 1993 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Roesler & Wranik 2004  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa barbata Arabian Helmeted Turtle Boycott & Bourquin 2008 [as subrufa] Els, Alqahtani, Fritz, Shobrak, Busais & Burriel-Carranza 2022 Gasperetti, Stimson, Miller, Ross & Gasperetti 1993 [as subrufa] Iverson 1992 [as subrufa] McCord, Joseph-Ouni & Bour 2004 [as subrufa] Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Testudinidae (Tortoises) Genus Centrochelys (African Spurred Tortoise)   Centrochelys sulcata African Spurred Tortoise Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021