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© Rune Midtgaard

Taxonomy of the family Agamidae
Bibliography of the genus Gonocephalus
Biodiversity of the family Agamidae


Western Anglehead Agamas, Angle-headed Agamas, Angle-headed Lizards, Hump-headed Agamas, Narrow-headed Tree Lizards

Westliche Winkelkopfagamen

Vestlige Vinkelhovedagamer

1805 Lophyrus Dumeril [not Lophyrus Poli 1791 (Mollusca); not Lophyrus Latreille 1802 (Insecta: Hymenoptera)] (type species: not designated, but presumably Iguana chamaeleontina Laurenti 1768; syn. Wermuth 1967)
1825 Gonocephalus Kaup (type species: Lophyrus tigrinus Dumeril & Bibron 1837)
1827 Goniocephalus Kaup [emendation]
1830 Gonyocephalus Wagler [emendation]
1845 Dilophyrus Gray (type species: Dilophyrus grandis Gray 1845; syn. Cogger, Cameron & Cogger 1983)
1846 Gonycephalus Agassiz [emendation]
Contents: 17 species, of which 10 (58.8%) are endemic.
Endemism: 0% 100%
Remarks: Previously included Malayodracon robinsonii and the genera Hypsilurus and Lophosaurus.
Distribution: SE. Asia, Malay Archipelago.
Reported from: Brunei, Indonesia (Bangka, Banyak Islands, Batu Islands, Java, Kalimantan, Natuna Islands [Bunguran], Nias, Siberut, Sipura, Sumatra), Laos, Malaysia (Sabah, Sarawak, West Malaysia [incl. Penang Island, Seribuat Archipelago (incl. Tioman)]), Philippines (Bohol, Calamian Islands, Camiguin Sur, Catanduanes, Cebu, Dinagat, Leyte, Luzon, Masbate, Mindanao, Mindoro, Negros, Panay, Polillo, Samar, Sibuyan, Talikud), Thailand, Vietnam.

Gonocephalus abbotti

Southern Thai Anglehead Agama, (Abbott's Anglehead Agama)

Abbotts Winkelkopfagame

Sydthailandsk Vinkelhovedagam

1922 Gonocephalus abbotti Cochran
Gonocephalus doriae abbotti Manthey & Denzer 1993

Remarks: Regarded as a subspecies of doriae by some authors (e.g., Manthey & Denzer 1993; Manthey 2010), but most recent works treat it as a separate species (e.g., Grismer 2008; Grismer & Aun 2008).
Distribution: Malaysia (West Malaysia), Thailand.

Gonocephalus bellii

Blue-necked Anglehead Agama, (Bell's Anglehead Agama)


Blåhalset Vinkelhovedagam

1837 Lophyrus bellii Dumeril & Bibron
Gonyocephalus bellii Boulenger 1885

Remarks: Records from Borneo (e.g., Wermuth 1967) refer to borneensis (Manthey & Denzer 1992; Das 2007c). Sarker & Sarker's (1985) record from Bangladesh is regarded as erroneous.
Distribution: Malaysia (West Malaysia [incl. Penang Island]), Thailand.

Gonocephalus bellii
© Rune Midtgaard

Gonocephalus beyschlagi

Northern Sumatra Anglehead Agama, (Beyschlag's Anglehead Agama)

Beyschlags Winkelkopfagame

Nordsumatransk Vinkelhovedagam

1892 Gonyocephalus beyschlagi Boettger

Distribution: Indonesia (Sumatra).

Gonocephalus borneensis

Bornean Anglehead Agama



1848 Lophyrus borneensis Schlegel
Gonyocephalus borneensis Boulenger 1885
1872 Tiaris miotympanum Günther (Manthey & Denzer 1992)
Gonyocephalus miotympanum Boulenger 1885
1991 Gonocephalus denzeri Manthey (Manthey & Grossmann 1997)

Remarks: Previously regarded as a synonym of bellii (e.g., Wermuth 1967). Revalidated by Manthey & Denzer (1992). Records from the Malay Peninsula probably refer to bellii or liogaster (Denzer & Manthey 1991).
Distribution: Brunei, Indonesia (Kalimantan), Malaysia (Sabah, Sarawak).

Gonocephalus borneensis
© Henrik Bringsøe

Gonocephalus chamaeleontinus

Chameleon Anglehead Agama, (Tioman Anglehead Agama)



1768 Iguana chamaeleontina Laurenti
Gonyocephalus chamaeleontinus Wagler 1830
1802 Lacerta scutata Shaw (Manthey & Denzer 1993)
1820 Agama gigantea Kuhl (Manthey & Denzer 1993)
1820 Agama tigrina Merrem (Manthey & Denzer 1993)
1826 Lophyrus gigas Boie (Manthey & Denzer 1993)
1827 Agama superciliosa Hardwicke & Gray (Manthey & Denzer 1993)
1844 Galeotes lophyrus Schlegel [part] (Manthey & Denzer 1993)
1848 Lophyrus sumatranus Schlegel (Manthey & Denzer 1993)
Gonyocephalus sumatranus Boulenger 1885

Remarks: Das (2010) erroneously reported this species from the West Malaysian mainland (Grismer 2011). Reported from Kalimantan by Iskandar (2004), but no vouchers exist (Das 2007).
Distribution: Indonesia (Bangka, Banyak Islands, Batu Islands, Java, Natuna Islands, Nias, Siberut, Sipura, Sumatra), Malaysia (West Malaysia [incl. Seribuat Archipelago (incl. Tioman)]).

Gonocephalus doriae

Sarawak Anglehead Agama, (Marquis Doria's Anglehead Agama)

Dorias Winkelkopfagame


1871 Gonyocephalus doriae Peters

Remarks: Some authors regard abbotti as a subspecies of doriae (e.g., Manthey & Denzer 1993).
Distribution: Indonesia (Kalimantan), Malaysia (Sabah, Sarawak), Thailand.

Gonocephalus grandis

Greater Anglehead Agama, Great Anglehead Agama, Giant Anglehead Lizard, Creek Anglehead

Große Winkelkopfagame

Stor Vinkelhovedagam

1845 Dilophyrus grandis Gray
Gonyocephalus grandis Schlegel 1858

Distribution: Brunei, Indonesia (Kalimantan, Nias, Siberut, Sipura, Sumatra), Laos, Malaysia (Sabah, Sarawak, West Malaysia [incl. Penang Island, Seribuat Archipelago (incl. Tioman)]), Thailand, Vietnam.

Gonocephalus grandis
© Rune Midtgaard

Gonocephalus interruptus

Mindoro Anglehead Agama, Mindoro Angle-headed Dragon, Mindanao Anglehead Agama



1885 Gonyocephalus interruptus Boulenger

Distribution: Philippines (Bohol, Mindanao, Mindoro, Talikud),

Gonocephalus klossi

Western Sumatra Anglehead Agama, (Kloss's Anglehead Agama)

Kloss' Winkelkopfagame

Vestsumatransk Vinkelhovedagam

1920 Gonyocephalus klossi Boulenger

Distribution: Indonesia (Sumatra).

Gonocephalus kuhlii

Java Anglehead Agama, (Kuhl's Anglehead Agama)

Kuhls Winkelkopfagame

Javansk Vinkelhovedagam

1848 Lophyrus kuhlii Schlegel
Gonyocephalus kuhlii Boulenger 1885
1844 Galeotes lophyrus Schlegel [part] (Manthey & Denzer 1993)

Remarks: Previously regarded as a synonym of chamaeleontinus (e.g., Wermuth 1967).
Distribution: Indonesia (Java, Sumatra).

Gonocephalus lacunosus

Sikulikap Anglehead Agama



1991 Gonocephalus lacunosus Manthey & Denzer

Distribution: Indonesia (Sumatra).

Gonocephalus liogaster

Blue-eyed Anglehead Agama, Orange-ringed Anglehead Agama, Bornean Anglehead Agama, Comb-crested Anglehead


Blåøjet Vinkelhovedagam

1872 Tiaris liogaster Günther
Gonyocephalus liogaster Boulenger 1885
1887 Gonyocephalus herveyi Boulenger (Dring 1979; Manthey & Denzer 1992)

Remarks: Das (2010) regarded herveyi as a valid species.
Distribution: Brunei, Indonesia (Kalimantan, Natuna Islands [Bunguran], Sumatra), Malaysia (Sabah, Sarawak, West Malaysia).

Gonocephalus megalepis

Sumatran Large-scaled Anglehead Agama, Green Anglehead Agama

Grüne Winkelkopfagame

Storskællet Vinkelhovedagam

1860 Lophyrus megalepis Bleeker
Gonyocephalus megalepis Boulenger 1887
1872 Tiaris tuberculatus Günther (Boulenger 1887; Manthey & Denzer 1991)

Remarks: Presence in Borneo needs confirmation (Denzer & Manthey 1991).
Distribution: Indonesia (Sumatra).

Gonocephalus mjoebergi

Mount Murud Anglehead Agama, (Mjöberg's Anglehead Agama)

Mjöbergs Winkelkopfagame


1925 Gonocephalus mjöbergi Smith
Gonocephalus mjoebergi [emendation] Wermuth 1967

Distribution: Malaysia (Sarawak).

Gonocephalus pyrius

Barisan Range Anglehead Agama, Fiery Anglehead Dragon



2021 Gonocephalus pyrius Harvey, Rech, Riyanto, Kurniawan & Smith

Distribution: Indonesia (Sumatra).

Gonocephalus semperi

White-spotted Anglehead Agama, White-spotted Angle-headed Dragon, (Semper's Anglehead Agama)

Sempers Winkelkopfagame

Hvidplettet Vinkelhovedagam

1867 Lophyrus semperi Peters
Gonyocephalus semperi Boulenger 1885

Distribution: Philippines (Bohol, Calamian Islands, Camiguin Sur, Cebu, Leyte, Mindanao, Mindoro, Negros, Panay).

Gonocephalus sophiae

Dark-spotted Anglehead Agama, Dark-spotted Anglehead Agama, Negros Forest Dragon, (Sophia's Anglehead Agama)

Sophias Winkelkopfagame

Mørkplettet Vinkelhovedagam

1845 Tiaris sophiae Gray
Gonyocephalus sophiae Boulenger 1885
1845 Tiaris bellii Gray [not Lophyrus bellii Duméril & Bibron 1837] (Taylor 1922)
1868 Tiaris petersii Günther (Wermuth 1967)

Distribution: Philippines (Cebu, Luzon, Masbate, Mindanao, Negros, Panay, Polillo, Sibuyan).