Reptiles of Leyte [Philippines] Related checklists:  Philippines Bibliography of the reptiles of Leyte Taxonomic history of the reptiles of Leyte Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: April 14th, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama Gaulke 1996 Hallermann 2005 Genus Draco (Gliding Dragons)   Draco bimaculatus Two-spotted Gliding Dragon McGuire & Alcala 2000 McGuire & Kiew 2001 Ross & Lazell 1990 Wermuth 1967   Draco mindanensis Mindanao Gliding Dragon McGuire & Alcala 2000 McGuire & Kiew 2001   Draco ornatus White-spotted Gliding Dragon McGuire & Alcala 2000 McGuire & Kiew 2001   Draco reticulatus Reticulated Gliding Dragon McGuire & Alcala 2000 McGuire & Kiew 2001 Genus Gonocephalus (Western Anglehead Agamas)   Gonocephalus semperi White-spotted Anglehead Agama Alcala 1986 Welton, Siler, Grismer, Diesmos, Sites & Brown 2017 Genus Hydrosaurus (Sail-tailed Agamas)   Hydrosaurus pustulatus Philippine Sail-tailed Agama Gaulke 1989  Family Gekkonidae (Typical Geckos) Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus annulatus Mindanao Banded Bent-toed Gecko Alcala 1986 Brown & Alcala 1978 Gaulke 1996 Welton, Siler, Diesmos & Brown 2009   * Cyrtodactylus gubaot Leyte Bent-toed Gecko Alcala 1986 [as agusanensis] Brown & Alcala 1978 [as agusanensis] Ross & Lazell 1990 [as agusanensis] Welton, Siler, Linkem, Diesmos & Brown 2010 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Brown & Alcala 1978 Genus Gekko (Tokays)   Gekko gecko Tokay Brown & Alcala 1978   Gekko intermedius Philippine Gliding Gecko Alcala 1986 [as Ptychozoon intermedium] Brown & Alcala 1978 [as Ptychozoon intermedium] Brown, Ferner & Diesmos 1997 [as Ptychozoon intermedium] Ross & Lazell 1990 [as Ptychozoon intermedium]   Gekko monarchus Spotted House Gecko Alcala 1986 Brown & Alcala 1978 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus frenatus Asian House Gecko Brown & Alcala 1978   Hemidactylus platyurus Flat-tailed Gecko Brown & Alcala 1978 Genus Lepidodactylus (Scaly-toed Geckos)   Lepidodactylus herrei White-lined Scaly-toed Gecko Alcala 1986 Brown & Alcala 1978 Genus Pseudogekko (Smooth-scaled Geckos)   Pseudogekko brevipes Orange-spotted Smooth-scaled Gecko Davis, Watters, Koehler, Whitsett, Huron, Brown, Diesmos & Siler 2015 Siler, Dececchi, Merkord, Davis, Christiani & Brown 2014   Pseudogekko ditoy Leyte Smooth-scaled Gecko Davis, Watters, Koehler, Whitsett, Huron, Brown, Diesmos & Siler 2015 Huron, Realubit, Cobb, Brown, Bergmann Morinaga, Diesmos, Diesmos, Brown & Siler 2016 Siler, Welton, Davis, Watters, Davey, Diesmos, Diesmos & Brown 2014   Pseudogekko pungkaypinit Southern Philippine Smooth-scaled Gecko Davis, Watters, Koehler, Whitsett, Huron, Brown, Diesmos & Siler 2015 Huron, Realubit, Cobb, Brown, Bergmann Morinaga, Diesmos, Diesmos, Brown & Siler 2016 Siler, Welton, Davis, Watters, Davey, Diesmos, Diesmos & Brown 2014  Family Scincidae (Skinks) Genus Brachymeles (Indo-Malayan Slender Skinks)   Brachymeles orientalis Bario Dusita Slender Skink Alcala 1986 Brown & Alcala 1980 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012   * Brachymeles paeforum San Vicente Slender Skink Alcala 1986 [as samarensis] Brown & Alcala 1980 [as samarensis] Siler & Brown 2010 [as samarensis] Siler, Fuiten, Jones, Alcala & Brown 2011 Siler, Jones, Diesmos, Diesmos & Brown 2012   Brachymeles samad Eastern Visayas Slender Skink Brown & Alcala 1980 [as hilong] Siler, Jones, Diesmos, Diesmos & Brown 2012 Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Brown & Alcala 1980 Genus Eutropis (South Asian Sun Skinks)   Eutropis indeprensa Mindoro Sun Skink Brown & Alcala 1980 Gaulke 1996   Eutropis multicarinata Many-keeled Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020 Brown & Alcala 1980   Eutropis multifasciata Many-lined Sun Skink Brown & Alcala 1980 David, Pauwels, Lays & Lenglet 2006 Genus Lamprolepis (Malay Archipelago Tree Skinks)   Lamprolepis smaragdina Emerald Tree Skink Brown & Alcala 1980 Gaulke 1996 Genus Lipinia (Indo-Pacific Striped Skinks)   Lipinia pulchella Yellow-striped Slender Tree Skink Brown & Alcala 1980 Supsup, Redoblado & Guinto 2016   Lipinia quadrivittata Black-striped Slender Tree Skink Brown & Alcala 1980 Genus Otosaurus (Philippine Giant Forest Skink)   Otosaurus cumingii Philippine Giant Forest Skink Alcala 1986 Brown & Alcala 1980 Brown & Rico 2009 Genus Parvoscincus (Philippine Dwarf Forest Skinks)   Parvoscincus steerei Guimaras Dwarf Forest Skink Brown & Alcala 1980 Genus Pinoyscincus (Philippine Forest Skinks)   Pinoyscincus coxi Agusan Forest Skink Alcala 1986 Brown & Alcala 1980   Pinoyscincus jagorii Samar Forest Skink Brown & Alcala 1980 Ross & Lazell 1990   Pinoyscincus llanosi Leyte Forest Skink Alcala 1986 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010   Pinoyscincus mindanensis Mindanao Forest Skink Alcala 1986 Brown & Alcala 1980 Ross & Lazell 1990 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus acutus Sharp-nosed Forest Skink Alcala 1986 Brown & Alcala 1980 Ross & Lazell 1990   Sphenomorphus fasciatus Philippine Banded Forest Skink Alcala 1986 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010   Sphenomorphus variegatus Philippine Black-spotted Forest Skink Brown & Alcala 1980 Genus Tropidophorus (Stream Skinks)   Tropidophorus grayi Philippine Stream Skink Alcala 1986 Brown & Alcala 1980  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus samarensis Samar Water Monitor Boehme 2003 [as V. cumingi] David, Pauwels, Lays & Lenglet 2006 [as V. cumingi] Koch, Auliya, Schmitz, Kuch & Boehme 2007 [as V. cumingi] Koch, Gaulke & Boehme 2010 Lisle 1996 [as V. cumingi] Mertens 1963 [as V. cumingi]

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake Iskandar & Colijn 2001 Leviton 1967 Leviton, Siler, Weinell & Brown 2018 Genus Boiga (Australasian Cat Snakes)   Boiga angulata Leyte Cat Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Orlov & Ryabov 2002   Boiga cynodon Dog-toothed Cat Snake Alcala 1986 Gaulke 1996 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Orlov & Ryabov 2002   Boiga dendrophila Eastern Mangrove Cat Snake Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Genus Calamaria (Oriental Reed Snakes)   Calamaria lumbricoidea Variable Reed Snake Alcala 1986 Grandison 1972 Inger & Marx 1965 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea paradisi Paradise Flying Tree Snake Iskandar & Colijn 2001 Leviton 1964 Leviton, Siler, Weinell & Brown 2018 Mertens 1968 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus erythrura Philippine Rat Snake Iskandar & Colijn 2001 Leviton 1979 Leviton, Siler, Weinell & Brown 2018 Schulz 1996 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis marenae Philippine Painted Bronzeback Leviton, Siler, Weinell & Brown 2018 Vogel & Rooijen 2008   Dendrelaphis philippinensis Mindanao Stripe-tailed Bronzeback Iskandar & Colijn 2001 [as terrificus] Leviton 1968 [as terrificus] Leviton, Siler, Weinell & Brown 2018 Rooijen & Vogel 2012 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma oxycephalum Red-tailed Green Rat Snake Leviton, Siler, Weinell & Brown 2018 Genus Lycodon (Oriental Wolf Snakes)   Lycodon capucinus Island Wolf Snake Iskandar & Colijn 2001 Leviton 1965 Leviton, Siler, Weinell & Brown 2018 O'Shea, Kusuma & Kaiser 2018   Lycodon dumerilii Mindanao Wolf Snake Leviton, Siler, Weinell & Brown 2018 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas luzonensis Smooth-scaled Mountain Rat Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1963 Leviton 1983 Leviton, Siler, Weinell & Brown 2018 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis auriculatus White-lined Keelback Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton 1970 Leviton, Siler, Weinell & Brown 2018   Rhabdophis lineatus Philippine Striped Keelback Leviton, Siler, Weinell & Brown 2018 Genus Stegonotus (Indo-Australian Ground Snakes)   Stegonotus muelleri Philippine Ground Snake Alcala 1986 Barnes & Knierim 2018 Iskandar & Colijn 2001 Kaiser, Kaiser & O'Shea 2018 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Genus Tropidonophis (Indo-Australian Keelbacks)   Tropidonophis dendrophiops Mindanao Keelback Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Malnate & Underwood 1988  Family Cyclocoridae (Philippine Burrowing Snakes, etc.) Genus Cyclocorus (Triangle-spotted Snakes)   Cyclocorus nuchalis Southern Triangle-spotted Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1965 Leviton, Siler, Weinell & Brown 2018 Genus Levitonius (Waray Dwarf Burrowing Snake)   Levitonius mirus Waray Dwarf Burrowing Snake Weinell, Paluh, Siler & Brown 2020 Genus Oxyrhabdium (Philippine Burrowing Snakes)   Oxyrhabdium modestum Philippine Bandless Burrowing Snake Alcala 1986 Gaulke & Operiano 2006 Iskandar & Colijn 2001 Leviton 1957 Leviton 1964 Leviton, Siler, Weinell & Brown 2018  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Naja (Typical Cobras)   Naja samarensis Southeastern Philippine Spitting Cobra Alcala 1986 Iskandar & Colijn 2001 Leviton 1964 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Wuster & Thorpe 1991 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Leviton, Siler, Weinell & Brown 2018 Sy & Boos 2015  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Iskandar & Colijn 2001 Leviton 1963 Leviton 1983 Leviton, Siler, Weinell & Brown 2018  Family Pareidae (Oriental Slug Eaters) Genus Aplopeltura (Blunt-headed Slug Eater)   Aplopeltura boa Blunt-headed Slug Eater Leviton, Siler, Weinell & Brown 2018  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Gaulke 1996 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999  Family Viperidae (Vipers) Genus Parias (Malay Archipelago Pitvipers)   Parias flavomaculatus Yellow-spotted Pitviper Alcala 1986 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Iskandar & Colijn 2001 Leviton 1964 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Genus Tropidolaemus (Temple Pitvipers)   Tropidolaemus philippensis South Philippine Temple Pitviper Iskandar & Colijn 2001 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018   Tropidolaemus subannulatus North Philippine Temple Pitviper Gaulke 1996 [as wagleri] Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 [as wagleri] Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 [as wagleri] Leviton 1964 [as wagleri] Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Vogel, David, Lutz, Rooijen & Vidal 2007

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Sagun, Ramoso & Mejino 2005  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle David, Pauwels, Lays & Lenglet 2006 Diesmos, Brown, Alcala & Sison 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021