Reptiles of Panay [Philippines] Related checklists:  Philippines Bibliography of the reptiles of Panay Taxonomic history of the reptiles of Panay Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: April 14th, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama David, Pauwels, Lays & Lenglet 2006 Gaulke 2011 Hallermann 2005 Genus Draco (Gliding Dragons)   Draco spilopterus Common Philippine Gliding Dragon Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Manthey 2008 McGuire & Alcala 2000 McGuire & Kiew 2001 Genus Gonocephalus (Western Anglehead Agamas)   Gonocephalus semperi White-spotted Anglehead Agama Welton, Siler, Grismer, Diesmos, Sites & Brown 2017   Gonocephalus sophiae Dark-spotted Anglehead Agama Gaulke 2011 Manthey 2010 Genus Hydrosaurus (Sail-tailed Agamas)   Hydrosaurus pustulatus Philippine Sail-tailed Agama Ferner, Brown, Sison & Kennedy 2001 Gaulke 1989 Gaulke 2011  Family Gekkonidae (Typical Geckos) Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus annulatus Mindanao Banded Bent-toed Gecko David, Pauwels, Lays & Lenglet 2006 Ferner, Brown, Sison & Kennedy 2001   Cyrtodactylus philippinicus Common Philippine Bent-toed Gecko Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko David, Pauwels, Lays & Lenglet 2006 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Genus Gekko (Tokays)   * Gekko ernstkelleri Panay Limestone Gecko Gaulke 2011 Linkem, Siler, Diesmos, Sy & Brown 2010 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Roesler, Siler, Brown, Demegillo & Gaulke 2006   Gekko gecko Tokay Brown & Alcala 1978 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011   Gekko mindorensis Mindoro Narrow-disked Gecko Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Linkem, Siler, Diesmos, Sy & Brown 2010 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Sison, Gonzales & Ferner 1995 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus frenatus Asian House Gecko Brown & Alcala 1978 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011   Hemidactylus platyurus Flat-tailed Gecko Brown & Alcala 1978 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011   Hemidactylus stejnegeri Taiwanese Half-toed Gecko Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Sison, Gonzales & Ferner 1995 Genus Hemiphyllodactylus (Slender Geckos)   Hemiphyllodactylus insularis Philippine Slender Gecko Brown & Alcala 1978 [as typus] Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Zug 2010 Genus Lepidodactylus (Scaly-toed Geckos)   Lepidodactylus lugubris Common Mourning Gecko Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011   Lepidodactylus planicauda Broad-tailed Scaly-toed Gecko Alcala 1986 Brown & Alcala 1978 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Genus Luperosaurus (Flap-legged Geckos)   Luperosaurus corfieldi Panay Flap-legged Gecko Gaulke 2011 Gaulke, Demegillo, Reiter & Tacud 2003 [as "cf. cumingii"] Gaulke, Roesler & Brown 2007 Genus Pseudogekko (Smooth-scaled Geckos)   Pseudogekko brevipes Orange-spotted Smooth-scaled Gecko Gaulke 2011 Gaulke, Demegillo, Reiter & Tacud 2003 [as "cf. brevipes"] Siler, Dececchi, Merkord, Davis, Christiani & Brown 2014  Family Scincidae (Skinks) Genus Brachymeles (Indo-Malayan Slender Skinks)   Brachymeles talinis Greater Striped Slender Skink Gaulke 2011 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012   Brachymeles taylori Common Negros Slender Skink Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011   Brachymeles tridactylus Negros Three-toed Slender Skink Brown & Alcala 1980 Davis, Feller, Brown & Siler 2014 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012 Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Brown 1991 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Genus Eutropis (South Asian Sun Skinks)   Eutropis borealis Subic Bay Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011   Eutropis indeprensa Mindoro Sun Skink Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Sison, Gonzales & Ferner 1995   Eutropis lapulapu Mactan Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020   Eutropis multifasciata Many-lined Sun Skink David, Pauwels, Lays & Lenglet 2006 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011   * Eutropis sibalom Sibalom Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020 Genus Insulasaurus (Island Forest Skinks)   Insulasaurus arborens Negros Forest Skink Alcala 1986 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Genus Lamprolepis (Malay Archipelago Tree Skinks)   Lamprolepis smaragdina Emerald Tree Skink Brown & Alcala 1980 Ferner, Brown, Sison & Kennedy 2001 Gaulke 1996 Gaulke 2011 Mertens 1929 Genus Lipinia (Indo-Pacific Striped Skinks)   Lipinia pulchella Yellow-striped Slender Tree Skink Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Supsup, Redoblado & Guinto 2016   Lipinia rabori Black Slender Striped Skink Gaulke 2011 Supsup, Redoblado & Guinto 2016 Genus Otosaurus (Philippine Giant Forest Skink)   Otosaurus cumingii Philippine Giant Forest Skink Ferner, Brown, Sison & Kennedy 2001 Genus Parvoscincus (Philippine Dwarf Forest Skinks)   * Parvoscincus sisoni Panay Dwarf Forest Skink Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Ferner, Brown & Greer 1997 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011   Parvoscincus steerei Guimaras Dwarf Forest Skink Brown & Alcala 1980 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Genus Pinoyscincus (Philippine Forest Skinks)   Pinoyscincus coxi Agusan Forest Skink Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011   Pinoyscincus jagorii Samar Forest Skink Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus fasciatus Philippine Banded Forest Skink Sison, Gonzales & Ferner 1995 Genus Tropidophorus (Stream Skinks)   Tropidophorus grayi Philippine Stream Skink Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Sison, Gonzales & Ferner 1995  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   * Varanus mabitang Panay Monitor Boehme 2003 Gaulke & Curio 2001 Gaulke 2011 Gaulke in Pianka, King & King (eds.) 2004   Varanus nuchalis Rough-necked Water Monitor Boehme 2003 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Koch, Auliya, Schmitz, Kuch & Boehme 2007 Koch, Gaulke & Boehme 2010 Lisle 1996

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Acrochordidae (Australasian File Snakes) Genus Acrochordus (Australasian File Snakes)   Acrochordus granulatus Lesser File Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Leviton, Siler, Weinell & Brown 2018 Wallach, Williams & Boundy 2014  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake Ferner, Brown, Sison & Kennedy 2001 Iskandar & Colijn 2001 Leviton 1967 Leviton, Siler, Weinell & Brown 2018 Wallach, Williams & Boundy 2014 Genus Boiga (Australasian Cat Snakes)   Boiga angulata Leyte Cat Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 [as "cf. angulata"] Leviton, Siler, Weinell & Brown 2018   Boiga cynodon Dog-toothed Cat Snake Ferner, Brown, Sison & Kennedy 2001 [as "cf. cynodon"] Gaulke 2011 Leviton, Siler, Weinell & Brown 2018   Boiga dendrophila Eastern Mangrove Cat Snake Ferner, Brown, Sison & Kennedy 2001 [as "cf. dendrophila"] Gaulke 2011 Gaulke, Demegillo & Vogel 2004 Leviton, Siler, Weinell & Brown 2018   Boiga drapiezii White-spotted Cat Snake Leviton, Siler, Weinell & Brown 2018 Genus Calamaria (Oriental Reed Snakes)   Calamaria bitorques Banded Reed Snake Gaulke & Vogel 2005 Gaulke 2011 [as "cf. bitorques"] Leviton, Siler, Weinell & Brown 2018   Calamaria gervaisi Philippine Reed Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Inger & Marx 1965 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea paradisi Paradise Flying Tree Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Sison, Gonzales & Ferner 1995 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus erythrura Philippine Rat Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Iskandar & Colijn 2001 Leviton 1979 Leviton, Siler, Weinell & Brown 2018 Schulz 1996 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis fuliginosus Negros Bronzeback Alcala 1986 [as terrificus] Ferner, Brown, Sison & Kennedy 2001 [as terrificus] Gaulke 2011 [as terrificus] Iskandar & Colijn 2001 [as terrificus] Leviton, Siler, Weinell & Brown 2018 Rooijen & Vogel 2012 Sison, Gonzales & Ferner 1995 [as D. caudolineatus]   Dendrelaphis marenae Philippine Painted Bronzeback Ferner, Brown, Sison & Kennedy 2001 [as pictus] Gaulke 2011 Iskandar & Colijn 2001 [as pictus] Leviton 1968 [as pictus] Leviton, Siler, Weinell & Brown 2018 Vogel & Rooijen 2008 Genus Dryophiops (Malayan Vine Snakes)   Dryophiops philippina Philippine Vine Snake Gaulke 2011 Gaulke, Demegillo, Reiter & Tacud 2003 Leviton, Siler, Weinell & Brown 2018 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma oxycephalum Red-tailed Green Rat Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Leviton, Siler, Weinell & Brown 2018 Genus Lycodon (Oriental Wolf Snakes)   Lycodon capucinus Island Wolf Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Iskandar & Colijn 2001 Leviton 1965 Leviton, Siler, Weinell & Brown 2018 O'Shea, Kusuma & Kaiser 2018   * Lycodon fausti Panay Wolf Snake Gaulke 2002 Gaulke 2011 Gaulke, Demegillo, Reiter & Tacud 2003 Leviton, Siler, Weinell & Brown 2018 Genus Oligodon (Kukri Snakes)   Oligodon modestus Spotted-bellied Kukri Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Sison, Gonzales & Ferner 1995 Genus Pseudorabdion (Dwarf Reed Snakes)   Pseudorabdion mcnamarae Negros Dwarf Reed Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Sison, Gonzales & Ferner 1995   Pseudorabdion oxycephalum Sharp-nosed Dwarf Reed Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Leviton, Siler, Weinell & Brown 2018   Pseudorabdion talonuran Panay Dwarf Reed Snake Brown, Leviton & Sison 1999 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas luzonensis Smooth-scaled Mountain Rat Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Leviton, Siler, Weinell & Brown 2018 Genus Tropidonophis (Indo-Australian Keelbacks)   Tropidonophis negrosensis Negros Spotted Keelback Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Malnate & Underwood 1988  Family Cyclocoridae (Philippine Burrowing Snakes, etc.) Genus Cyclocorus (Triangle-spotted Snakes)   Cyclocorus lineatus Northern Triangle-spotted Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Leviton, Siler, Weinell & Brown 2018 Genus Hologerrhum (Philippine Cylindrical Snakes)   * Hologerrhum dermali Panay Cylindrical Snake Brown, Leviton, Ferner & Sison 2001 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Leviton, Siler, Weinell & Brown 2018 Genus Oxyrhabdium (Philippine Burrowing Snakes)   Oxyrhabdium leporinum Philippine Banded Burrowing Snake Gaulke & Operiano 2006 Gaulke 2011 Leviton, Siler, Weinell & Brown 2018   Oxyrhabdium modestum Philippine Bandless Burrowing Snake Leviton, Siler, Weinell & Brown 2018  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Hemibungarus (Barred Coral Snakes)   Hemibungarus gemianulis Visayas Barred Coral Snake Alcala 1986 Brown, Smart, Leviton & Smith 2018 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Vogel 2006 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra David, Pauwels, Lays & Lenglet 2006 Gaulke 2011 Gaulke, Demegillo, Reiter & Tacud 2003 Leviton, Siler, Weinell & Brown 2018  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis belcheri Western Faint-banded Sea Snake Ferner, Brown, Sison & Kennedy 2001 Smith 1926   Hydrophis curtus Short Sea Snake Ferner, Brown, Sison & Kennedy 2001 [as Lapemis hardwickii]   Hydrophis cyanocinctus Blue-banded Sea Snake Ferner, Brown, Sison & Kennedy 2001 Genus Laticauda (Sea Kraits)   Laticauda colubrina Yellow-lipped Sea Krait Ferner, Brown, Sison & Kennedy 2001 Leviton, Siler, Weinell & Brown 2018  Family Homalopsidae (Australasian Mud Snakes) Genus Cerberus (Dog-faced Water Snakes)   Cerberus schneiderii South East Asian Dog-faced Water Snake Dunson & Minton 1978 [as C. rynchops] Ferner, Brown, Sison & Kennedy 2001 [as C. rynchops] Gaulke 2011 [as C. rynchops] Gyi 1970 [as C. rynchops] Iskandar & Colijn 2001 [as C. rynchops] Leviton, Siler, Weinell & Brown 2018  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Leviton 1963 Leviton, Siler, Weinell & Brown 2018  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Ferner, Brown, Sison & Kennedy 2001 Gaulke 1996 Gaulke 2011 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999  Family Typhlopidae (Typical Blind Snakes) Genus Malayotyphlops (Malay Archipelago Blind Snakes)   Malayotyphlops castanotus Brown-bellied Blind Snake Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 Wynn & Leviton 1993 Wynn, Diesmos & Brown 2016 Genus Ramphotyphlops (Western Pacific Blind Snakes)   Ramphotyphlops cumingii Cumming's Blind Snake Ferner, Brown, Sison & Kennedy 2001  Family Viperidae (Vipers) Genus Parias (Malay Archipelago Pitvipers)   Parias flavomaculatus Yellow-spotted Pitviper Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Leviton, Siler, Weinell & Brown 2018 Vogel 2006 Genus Tropidolaemus (Temple Pitvipers)   Tropidolaemus subannulatus North Philippine Temple Pitviper Ferner, Brown, Sison & Kennedy 2001 [as "cf. wagleri"] Gaulke 2011 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 [as wagleri] Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Vogel, David, Lutz, Rooijen & Vidal 2007

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Sagun, Ramoso & Mejino 2005  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle David, Pauwels, Lays & Lenglet 2006 Diesmos, Brown, Alcala & Sison 2008 Ferner, Brown, Sison & Kennedy 2001 Gaulke 2011 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021