Reptiles of Mindanao [Philippines] Related checklists:  Philippines Bibliography of the reptiles of Mindanao Taxonomic history of the reptiles of Mindanao Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: April 14th, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus mindorensis Philippine Freshwater Crocodile Alcala 1986 Trutnau & Sommerlad 2006 Wermuth & Mertens 1977

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama David, Pauwels, Lays & Lenglet 2006 Gaulke 1996 Hallermann 2005 Smith 1993 Taylor 1922 Genus Draco (Gliding Dragons)   Draco bimaculatus Two-spotted Gliding Dragon David, Pauwels, Lays & Lenglet 2006 McGuire & Alcala 2000 McGuire & Kiew 2001 Musters 1983 Ross & Lazell 1990 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017 Wermuth 1967   Draco cyanopterus Masked Gliding Dragon Beukema 2011 Manthey 2008 McGuire & Alcala 2000 McGuire & Kiew 2001 Ross & Lazell 1990 [as everetti] Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017   Draco guentheri Southern Philippine Gliding Dragon Alcala 1986 Beukema 2011 Manthey 2008 McGuire & Alcala 2000 McGuire & Kiew 2001   Draco mindanensis Mindanao Gliding Dragon David, Pauwels, Lays & Lenglet 2006 Inger 1983 Manthey 2008 McGuire & Alcala 2000 McGuire & Kiew 2001 Musters 1983 Ross & Lazell 1990 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Wermuth 1967   Draco ornatus White-spotted Gliding Dragon Alcala 1986 McGuire & Alcala 2000 McGuire & Kiew 2001 Ross & Lazell 1990 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Genus Gonocephalus (Western Anglehead Agamas)   Gonocephalus interruptus Mindoro Anglehead Agama Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 [as "cf. interruptus] Welton, Siler, Grismer, Diesmos, Sites & Brown 2017 Wermuth 1967   Gonocephalus semperi White-spotted Anglehead Agama Alcala 1986 David, Pauwels, Lays & Lenglet 2006 Relox, Leano & Ates-Camino 2011 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017 Wermuth 1967   Gonocephalus sophiae Dark-spotted Anglehead Agama Wermuth 1967 Genus Hydrosaurus (Sail-tailed Agamas)   Hydrosaurus pustulatus Philippine Sail-tailed Agama Dela Cruz 2022 Gaulke 1989 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Wermuth 1967  Family Dibamidae (Blind Skinks) Genus Dibamus (Oriental Blind Skinks)   Dibamus leucurus White-tailed Blind Skink Alcala 1986 Greer 1985   Dibamus novaeguineae Indo-Australian Blind Skink Greer 1985  Family Gekkonidae (Typical Geckos) Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus agusanensis Agusan Bent-toed Gecko Alcala 1986 Brown & Alcala 1978 Ross & Lazell 1990 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Welton, Siler, Linkem, Diesmos & Brown 2010   Cyrtodactylus annulatus Mindanao Banded Bent-toed Gecko Alcala 1986 Beukema 2011 Brown & Alcala 1978 David, Pauwels, Lays & Lenglet 2006 Gaulke 1996 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017 Welton, Siler, Diesmos & Brown 2009 Welton, Siler, Linkem, Diesmos & Brown 2010   Cyrtodactylus jambangan Jambangan Bent-toed Gecko Welton, Siler, Diesmos & Brown 2010 Welton, Siler, Linkem, Diesmos & Brown 2010   Cyrtodactylus philippinicus Common Philippine Bent-toed Gecko Supsup, Guinto, Redoblado & Gomez 2017 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Brown & Alcala 1978 David, Pauwels, Lays & Lenglet 2006 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Genus Gekko (Tokays)   Gekko gecko Tokay Brown & Alcala 1978 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016   Gekko intermedius Philippine Gliding Gecko Alcala 1986 [as Ptychozoon intermedium] Beukema 2011 [as Ptychozoon intermedium] Brown & Alcala 1978 [as Ptychozoon intermedium] Brown, Ferner & Diesmos 1997 [as Ptychozoon intermedium] Ross & Lazell 1990 [as Ptychozoon intermedium]   Gekko mindorensis Mindoro Narrow-disked Gecko Relox, Leano & Ates-Camino 2011 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017   Gekko monarchus Spotted House Gecko Alcala 1986 Brown & Alcala 1978 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus frenatus Asian House Gecko Brown & Alcala 1978 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016   Hemidactylus platyurus Flat-tailed Gecko Beukema 2011 Brown & Alcala 1978 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Genus Hemiphyllodactylus (Slender Geckos)   Hemiphyllodactylus insularis Philippine Slender Gecko Brown & Alcala 1978 [as typus] Grismer, Wood, Anuar, Muin, Quah, McGuire, Brown, Ngo & Pham 2013 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 [as "cf. typus"] Zug 2010 Genus Lepidodactylus (Scaly-toed Geckos)   Lepidodactylus aureolineatus Golden-lined Scaly-toed Gecko Alcala 1986 Brown & Alcala 1978 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016   Lepidodactylus labialis Dark-spotted Scaly-toed Gecko Alcala 1986 Brown & Alcala 1978 Eliades, Brown, Huang & Siler 2021 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016   Lepidodactylus planicauda Broad-tailed Scaly-toed Gecko Alcala 1986 Brown & Alcala 1978 Eliades, Brown, Huang & Siler 2021 Genus Luperosaurus (Flap-legged Geckos)   Luperosaurus joloensis Jolo Flap-legged Gecko Alcala 1986 Brown & Alcala 1978 Brown, Supriatna & Ota 2000 Genus Pseudogekko (Smooth-scaled Geckos)   * Pseudogekko chavacano Zamboanga Smooth-scaled Gecko Davis, Watters, Koehler, Whitsett, Huron, Brown, Diesmos & Siler 2015 Siler, Welton, Davis, Watters, Davey, Diesmos, Diesmos & Brown 2014   Pseudogekko pungkaypinit Southern Philippine Smooth-scaled Gecko Davis, Watters, Koehler, Whitsett, Huron, Brown, Diesmos & Siler 2015 Huron, Realubit, Cobb, Brown, Bergmann Morinaga, Diesmos, Diesmos, Brown & Siler 2016 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Siler, Welton, Davis, Watters, Davey, Diesmos, Diesmos & Brown 2014  Family Scincidae (Skinks) Genus Brachymeles (Indo-Malayan Slender Skinks)   Brachymeles gracilis Graceful Slender Skink Alcala 1986 Brown & Alcala 1980 David, Pauwels, Lays & Lenglet 2006 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012   Brachymeles hilong Diuata Slender Skink Alcala 1986 Brown & Alcala 1980 Brown & Rabor 1967 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017   Brachymeles orientalis Bario Dusita Slender Skink Alcala 1986 Brown & Alcala 1980 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012 Smith 1993   * Brachymeles pathfinderi Cotabato Slender Skink Alcala 1986 Brown & Alcala 1980 Siler & Brown 2010 Siler, Crombie, Diesmos & Brown 2011 Siler, Jones, Diesmos, Diesmos & Brown 2012   Brachymeles schadenbergi Southern Philippine Slender Skink Alcala 1986 Brown & Alcala 1980 Siler & Brown 2010 Siler, Jones, Diesmos, Diesmos & Brown 2012 Supsup, Guinto, Redoblado & Gomez 2017   * Brachymeles tiboliorum Western Mindanao Slender Skink Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Siler, Jones, Diesmos, Diesmos & Brown 2012 Genus Dasia (Indo-Malayan Tree Skinks)   Dasia semicincta Southern Keel-scaled Tree Skink Alcala 1986 Brown & Alcala 1980 Inger & Brown 1980 Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Brown & Alcala 1980 Brown 1991 Gaulke 1996   Emoia caeruleocauda Pacific Blue-tailed Skink Brown 1991   Emoia ruficauda Red-tailed Slender Skink Alcala 1986 Brown & Alcala 1980 Brown 1991 Gaulke & Alcala 2009 Sy & Buday 2014 Genus Eutropis (South Asian Sun Skinks)   * Eutropis alcalai Pasonanca Sun Skink Barley, Sanguila & Brown 2021   Eutropis caraga Misamis Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020   * Eutropis cuprea Copper Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020   * Eutropis englei Mindanao Sun Skink Alcala 1986 Barley, Diesmos, Siler, Martinez & Brown 2020 Brown & Alcala 1980 Pitogo, Senarillos, Tubera & Barley 2020 Relox, Leano & Ates-Camino 2011 Supsup, Guinto, Redoblado & Gomez 2017   Eutropis indeprensa Mindoro Sun Skink Brown & Alcala 1980 David, Pauwels, Lays & Lenglet 2006 Gaulke 1996 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 [as "cf. indeprensa"] Supsup, Guinto, Redoblado & Gomez 2017 [as "cf. indeprensa"]   Eutropis lapulapu Mactan Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020   Eutropis multicarinata Many-keeled Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020   Eutropis multifasciata Many-lined Sun Skink Beukema 2011 Brown & Alcala 1980 David, Pauwels, Lays & Lenglet 2006 Nuneza, Ates-Camino & Alicante 2010 O'Shea, Kusuma & Kaiser 2018 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016   Eutropis rudis Three-keeled Sun Skink Brown & Alcala 1980 Genus Lamprolepis (Malay Archipelago Tree Skinks)   Lamprolepis smaragdina Emerald Tree Skink Brown & Alcala 1980 David, Pauwels, Lays & Lenglet 2006 Gaulke 1996 Mertens 1929 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017 Genus Lipinia (Indo-Pacific Striped Skinks)   Lipinia auriculata Bronze Striped Tree Skink Brown & Alcala 1980 David, Pauwels, Lays & Lenglet 2006 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Redoblado & Guinto 2016   Lipinia pulchella Yellow-striped Slender Tree Skink Alcala 1986 Brown & Alcala 1956 Brown & Alcala 1980 Relox, Leano & Ates-Camino 2011 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017 Supsup, Redoblado & Guinto 2016   Lipinia quadrivittata Black-striped Slender Tree Skink Alcala 1986 Brown & Alcala 1956 Brown & Alcala 1980 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017 Supsup, Redoblado & Guinto 2016   Lipinia semperi Mindanao Striped Skink Alcala 1986 Brown & Alcala 1956 Brown & Alcala 1980 Gunther 2000 Smith 1993 Supsup, Redoblado & Guinto 2016   Lipinia subvittata Short-striped Tree Skink Alcala 1986 Brown & Alcala 1980 David, Pauwels, Lays & Lenglet 2006 Supsup, Redoblado & Guinto 2016   Lipinia vulcania Dapitan Peak Striped Skink Alcala 1986 Brown & Alcala 1980   * Lipinia zamboangensis Zamboanga Striped Skink Alcala 1986 Brown & Alcala 1980 Supsup, Redoblado & Guinto 2016 Genus Otosaurus (Philippine Giant Forest Skink)   Otosaurus cumingii Philippine Giant Forest Skink Alcala 1986 Brown & Alcala 1980 Brown & Rico 2009 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Genus Parvoscincus (Philippine Dwarf Forest Skinks)   Parvoscincus decipiens Black-sided Dwarf Forest Skink Alcala 1986 Beukema 2011 Brown & Alcala 1980 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017   * Parvoscincus kitangladensis Mount Kitanglad Dwarf Forest Skink Beukema 2011 Brown 1995 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 [as "cf. kitangladensis"]   Parvoscincus steerei Guimaras Dwarf Forest Skink Brown & Alcala 1980 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Genus Pinoyscincus (Philippine Forest Skinks)   Pinoyscincus abdictus Camiguin Forest Skink Beukema 2011 Brown & Alcala 1980 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016   Pinoyscincus coxi Agusan Forest Skink Alcala 1986 Beukema 2011 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017   Pinoyscincus jagorii Samar Forest Skink Brown & Alcala 1980 Ross & Lazell 1990 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016   Pinoyscincus mindanensis Mindanao Forest Skink Alcala 1986 Beukema 2011 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 David, Pauwels, Lays & Lenglet 2006 Ross & Lazell 1990 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus acutus Sharp-nosed Forest Skink Alcala 1986 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Ross & Lazell 1990 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993   * Sphenomorphus diwata Diwata Mountains Forest Skink Alcala 1986 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017   Sphenomorphus fasciatus Philippine Banded Forest Skink Alcala 1986 Beukema 2011 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 David, Pauwels, Lays & Lenglet 2006 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017   Sphenomorphus variegatus Philippine Black-spotted Forest Skink Alcala 1986 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 David, Pauwels, Lays & Lenglet 2006 Nuneza, Ates-Camino & Alicante 2010 Relox, Leano & Ates-Camino 2011 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017 Genus Tropidophorus (Stream Skinks)   * Tropidophorus davaoensis Davao Stream Skink Brown & Alcala 1980 David, Pauwels, Lays & Lenglet 2006 Supsup, Guinto, Redoblado & Gomez 2017   Tropidophorus misaminius Misamis Stream Skink Alcala 1986 Beukema 2011 Brown & Alcala 1980 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016   Tropidophorus partelloi Mindanao Stream Skink Alcala 1986 Beukema 2011 Brown & Alcala 1980 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Genus Tytthoscincus (Dwarf Leaf-litter Skinks)   Tytthoscincus atrigularis Zamboanga Leaf-litter Skink Alcala 1986 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus cumingi Mindanao Water Monitor Boehme 2003 David, Pauwels, Lays & Lenglet 2006 Koch, Auliya, Schmitz, Kuch & Boehme 2007 Koch, Gaulke & Boehme 2010 Lisle 1996 Mertens 1963 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake David, Pauwels, Lays & Lenglet 2006 Gaulke 1994 Iskandar & Colijn 2001 Leviton 1967 Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017 Wallach, Williams & Boundy 2014 Genus Boiga (Australasian Cat Snakes)   Boiga angulata Leyte Cat Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Orlov & Ryabov 2002   Boiga cynodon Dog-toothed Cat Snake Alcala 1986 Gaulke 1994 Gaulke 1996 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Orlov & Ryabov 2002 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017   Boiga dendrophila Eastern Mangrove Cat Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 Orlov & Ryabov 2002 Relox, Leano & Ates-Camino 2011 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017   Boiga drapiezii White-spotted Cat Snake Leviton, Siler, Weinell & Brown 2018 Genus Calamaria (Oriental Reed Snakes)   Calamaria gervaisi Philippine Reed Snake Beukema 2011 David, Pauwels, Lays & Lenglet 2006 Inger & Marx 1965 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 Relox, Leano & Ates-Camino 2011 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017   Calamaria lumbricoidea Variable Reed Snake Beukema 2011 David, Pauwels, Lays & Lenglet 2006 Inger & Marx 1965 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016   Calamaria virgulata Blunt-tailed Reed Snake Alcala 1986 Inger & Marx 1965 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea paradisi Paradise Flying Tree Snake Gaulke 1994 Iskandar & Colijn 2001 Leviton 1964 Leviton, Siler, Weinell & Brown 2018 Mertens 1968 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus erythrura Philippine Rat Snake David, Pauwels, Lays & Lenglet 2006 Gaulke 1994 Iskandar & Colijn 2001 Leviton 1979 Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Schulz 1996 Smith 1993 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis marenae Philippine Painted Bronzeback Gaulke 1994 [as pictus] Iskandar & Colijn 2001 [as pictus] Leviton 1968 [as pictus] Leviton, Siler, Weinell & Brown 2018 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017 Vogel & Rooijen 2008   Dendrelaphis philippinensis Mindanao Stripe-tailed Bronzeback David, Pauwels, Lays & Lenglet 2006 [as terrificus] Gaulke 1994 [as terrificus] Gunther 1879 Iskandar & Colijn 2001 [as terrificus] Leviton 1968 [as luzonensis] Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 [as caudolineatus] Rooijen & Vogel 2012 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 [as terrificus] Genus Dryophiops (Malayan Vine Snakes)   Dryophiops philippina Philippine Vine Snake Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Smith 1993 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma oxycephalum Red-tailed Green Rat Snake Leviton, Siler, Weinell & Brown 2018 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017 Genus Lycodon (Oriental Wolf Snakes)   Lycodon capucinus Island Wolf Snake Iskandar & Colijn 2001 Leviton 1965 Leviton, Siler, Weinell & Brown 2018 O'Shea, Kusuma & Kaiser 2018 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016   Lycodon dumerilii Mindanao Wolf Snake Alcala 1986 Iskandar & Colijn 2001 Lanza 1999 Leviton 1965 Leviton, Siler, Weinell & Brown 2018 Relox, Leano & Ates-Camino 2011 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017 Genus Oligodon (Kukri Snakes)   * Oligodon maculatus Philippine Spotted Kukri Snake Alcala 1986 David, Pauwels, Lays & Lenglet 2006 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Qian, Qi, Shi, Lu, Jenkins, Mo & Li 2021 Relox, Leano & Ates-Camino 2011 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017   Oligodon modestus Spotted-bellied Kukri Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Genus Opisthotropis (Mountain Stream Keelbacks)   * Opisthotropis alcalai Mindanao Mountain Stream Keelback Alcala 1986 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 Genus Pseudorabdion (Dwarf Reed Snakes)   * Pseudorabdion ater Zamboanga Dwarf Reed Snake Alcala 1986 Iskandar & Colijn 2001 Leviton & Brown 1959 Leviton 1963 Leviton, Siler, Weinell & Brown 2018   * Pseudorabdion taylori Mindanao Dwarf Reed Snake Alcala 1986 Iskandar & Colijn 2001 Leviton & Brown 1959 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis auriculatus White-lined Keelback Alcala 1986 Beukema 2011 David, Pauwels, Lays & Lenglet 2006 Doria, Petri, Bellati, Tiso & Pistarino 2013 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton 1970 Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017   Rhabdophis lineatus Philippine Striped Keelback Alcala 1986 David, Pauwels, Lays & Lenglet 2006 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017 Genus Stegonotus (Indo-Australian Ground Snakes)   Stegonotus muelleri Philippine Ground Snake Alcala 1986 Barnes & Knierim 2018 Iskandar & Colijn 2001 Kaiser, Kaiser & O'Shea 2018 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Genus Tropidonophis (Indo-Australian Keelbacks)   Tropidonophis dendrophiops Mindanao Keelback David, Pauwels, Lays & Lenglet 2006 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Malnate & Underwood 1988 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017  Family Cyclocoridae (Philippine Burrowing Snakes, etc.) Genus Cyclocorus (Triangle-spotted Snakes)   Cyclocorus nuchalis Southern Triangle-spotted Snake Alcala 1986 Beukema 2011 David, Pauwels, Lays & Lenglet 2006 Iskandar & Colijn 2001 Leviton 1965 Leviton, Siler, Weinell & Brown 2018 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017 Genus Oxyrhabdium (Philippine Burrowing Snakes)   Oxyrhabdium leporinum Philippine Banded Burrowing Snake Relox, Leano & Ates-Camino 2011   Oxyrhabdium modestum Philippine Bandless Burrowing Snake Alcala 1986 Beukema 2011 David, Pauwels, Lays & Lenglet 2006 Gaulke & Operiano 2006 Iskandar & Colijn 2001 Leviton 1957 Leviton 1964 Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Calliophis (South Asian Coral Snakes)   Calliophis philippinus Philippine Striped Coral Snake Alcala 1986 Brown, Smart, Leviton & Smith 2018 Gaulke 1994 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017   Calliophis salitan Dinagat Coral Snake Brown, Calvo-Revuelta, Goyes Vallejos, Sanchez-Vialas & Riva 2021 Genus Naja (Typical Cobras)   Naja samarensis Southeastern Philippine Spitting Cobra Alcala 1986 David, Pauwels, Lays & Lenglet 2006 Iskandar & Colijn 2001 Leviton 1964 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Vera, Delima-Baron & Sy 2022 Wuster & Thorpe 1991 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Alcala 1986 David, Pauwels, Lays & Lenglet 2006 Iskandar & Colijn 2001 Leviton 1964 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Sy 2016  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis atriceps Black-headed Sea Snake Alcala 1986 [as H. fasciatus] Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018   Hydrophis curtus Short Sea Snake Leviton, Siler, Weinell & Brown 2018   Hydrophis cyanocinctus Blue-banded Sea Snake Leviton, Siler, Weinell & Brown 2018   Hydrophis platurus Yellow-bellied Sea Snake Leviton, Siler, Weinell & Brown 2018 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 [as "platyurus"] Genus Laticauda (Sea Kraits)   Laticauda colubrina Yellow-lipped Sea Krait Heatwole, Busack & Cogger 2005 Leviton, Siler, Weinell & Brown 2018   Laticauda laticaudata Blue-lipped Sea Krait Iskandar & Colijn 2001 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018  Family Gerrhopilidae (Indo-Malayan Blind Snakes) Genus Gerrhopilus (Indo-Malayan Blind Snakes)   Gerrhopilus hedraeus Negros Blind Snake Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999  Family Homalopsidae (Australasian Mud Snakes) Genus Cerberus (Dog-faced Water Snakes)   Cerberus schneiderii South East Asian Dog-faced Water Snake Gyi 1970 [as C. rynchops] Iskandar & Colijn 2001 [as C. rynchops] Leviton, Siler, Weinell & Brown 2018 Genus Fordonia (White-bellied Mangrove Snake)   Fordonia leucobalia White-bellied Mangrove Snake Gyi 1970 Iskandar & Colijn 2001  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Beukema 2011 David, Pauwels, Lays & Lenglet 2006 Iskandar & Colijn 2001 Leviton 1963 Leviton 1983 Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 Relox, Leano & Ates-Camino 2011 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Smith 1993 Supsup, Guinto, Redoblado & Gomez 2017  Family Pareidae (Oriental Slug Eaters) Genus Aplopeltura (Blunt-headed Slug Eater)   Aplopeltura boa Blunt-headed Slug Eater Alcala 1986 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 Poyarkov, Nguyen, Pawangkhanant, Yushchenko, Brakels, Nguyen, Nguyen, Suwannapoom, Orlov & Vogel 2022 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017 Taylor 1965 Wallach, Williams & Boundy 2014  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python David, Pauwels, Lays & Lenglet 2006 Gaulke 1994 Gaulke 1996 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 Nuneza, Ates-Camino & Alicante 2010 Relox, Leano & Ates-Camino 2011 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017  Family Typhlopidae (Typical Blind Snakes) Genus Malayotyphlops (Malay Archipelago Blind Snakes)   Malayotyphlops sp. (Blind Snake) Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Genus Ramphotyphlops (Western Pacific Blind Snakes)   Ramphotyphlops cumingii Cumming's Blind Snake Alcala 1986 [as longicauda] Hahn 1980 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 [as "cf. cumingi"] Supsup, Guinto, Redoblado & Gomez 2017  Family Viperidae (Vipers) Genus Parias (Malay Archipelago Pitvipers)   Parias flavomaculatus Yellow-spotted Pitviper Alcala 1986 David, Pauwels, Lays & Lenglet 2006 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Iskandar & Colijn 2001 Leviton 1964 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Nuneza, Ates-Camino & Alicante 2010 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 [as "cf. flavomaculatus"] Supsup, Guinto, Redoblado & Gomez 2017 Genus Tropidolaemus (Temple Pitvipers)   Tropidolaemus philippensis South Philippine Temple Pitviper Iskandar & Colijn 2001 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Vogel 2006 Vogel, David, Lutz, Rooijen & Vidal 2007   Tropidolaemus subannulatus North Philippine Temple Pitviper Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Relox, Leano & Ates-Camino 2011 Sanguila, Cobb, Siler, Diesmos, Alcala & Brown 2016 Supsup, Guinto, Redoblado & Gomez 2017 Vogel 2006 [as "cf. subannulatus"] Vogel, David, Lutz, Rooijen & Vidal 2007

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Byrne & Hines 2005 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Byrne & Hines 2005 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Byrne & Hines 2005  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Byrne & Hines 2005  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle Beukema 2011 David, Pauwels, Lays & Lenglet 2006 Diesmos, Brown, Alcala & Sison 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Trionychidae (Softshell Turtles) Genus Dogania (Malayan Softshell Turtle)   Dogania subplana Malayan Softshell Turtle Diesmos, Brown, Alcala & Sison 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelochelys (Indomalayan Giant Softshell Turtles)   Pelochelys cantorii Asian Giant Softshell Turtle Diesmos, Brown, Alcala & Sison 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021