Reptiles of Montenegro    Bibliography of the reptiles of Montenegro Taxonomic history of the reptiles of Montenegro Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Anguidae (Slowworms, etc.) Genus Anguis (Slowworms)   Anguis graeca Greek Slowworm Gvozdik, Benkovsky, Crottini, Bellati, Moravec, Romano, Sacchi & Jandzik 2013 Gvozdik, Jandzik, Lymberakis, Jablonski & Moravec 2010 Jovanovic 2009 [as A. fragilis] Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 [as "fragilis complex"] Mikulicek, Jablonski, Palenik, Gvozdik & Jandzik 2018 Polovic & Cadenovic 2014 [as A. fragilis] Pozzi 1966 [as A. fragilis] Genus Pseudopus (Eurasian Glass Lizard)   Pseudopus apodus Eurasian Glass Lizard Brelih & Dzukic 1974 Jovanovic 2009 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Pozzi 1966  Family Gekkonidae (Typical Geckos) Genus Hemidactylus (Half-toed Geckos)   Hemidactylus turcicus Turkish Half-toed Gecko Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Speybroeck, Beukema, Bok & Voort 2016  Family Lacertidae (Typical Lizards) Genus Algyroides (Keeled Lizards)   Algyroides nigropunctatus Blue-throated Keeled Lizard Ajtic, Tomovic, Aleksic & Crnobrnja-Isailovic 2005 Bischoff in Boehme (ed.) 1981 Brelih & Dzukic 1974 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Speybroeck, Beukema, Bok & Voort 2016 Genus Dalmatolacerta (Sharp-snouted Rock Lizard)   Dalmatolacerta oxycephala Sharp-snouted Rock Lizard Arnold & Ovenden 2002 Bischoff in Boehme (ed.) 1984 Brelih & Dzukic 1974 Crnobrnja-Isailovic, Corovic & Cosic 2020 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Mayer & Podnar 2003 Sindaco & Jeremchenko 2008 Speybroeck, Beukema, Bok & Voort 2016 Genus Dinarolacerta (Montenegro Rock Lizards)   Dinarolacerta montenegrina Prokletije Rock Lizard Ljubisavljevic, Arribas, Dzukic & Carranza 2007 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Speybroeck, Beukema, Bok & Voort 2016 Trapp 2011   Dinarolacerta mosorensis Mosor Rock Lizard Arnold & Ovenden 2002 Bischoff in Boehme (ed.) 1984 Brelih & Dzukic 1974 Ljubisavljevic, Arribas, Dzukic & Carranza 2007 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Sindaco & Jeremchenko 2008 Speybroeck, Beukema, Bok & Voort 2016 Trapp 2011 Genus Lacerta (Eurasian Green Lizards)   Lacerta agilis Northern Eurasian Sand Lizard Bischoff in Boehme (ed.) 1984 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Polovic & Cadenovic 2014 Schweigger 2008 Trapp 2011   Lacerta bilineata Western Green Lizard Boehme, Fritz, Kotenko, Dzukic, Ljubisavljevic, Tzankov & Berendonk 2007 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 [as "viridis complex" [part]] Marzahn, Mayer, Joger, Ilgaz, Jablonski, Kindler, Kumlutas, Nistri, Schneewiess, Vamberger, Zagar & Fritz 2015 [as "Adriatic lineage"]   Lacerta trilineata Western Balkan Green Lizard Brelih & Dzukic 1974 Jovanovic 2009 Kornilios, Thanou, Lymberakis, Ilgaz, Kumlutas & Leache 2020 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Mayer & Podnar 2003 Nettmann & Rykena in Boehme (ed.) 1984 Trapp 2011   Lacerta viridis Eastern Green Lizard Boehme, Fritz, Kotenko, Dzukic, Ljubisavljevic, Tzankov & Berendonk 2007 Jovanovic 2009 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 [as "viridis complex" [part]] Marzahn, Mayer, Joger, Ilgaz, Jablonski, Kindler, Kumlutas, Nistri, Schneewiess, Vamberger, Zagar & Fritz 2015 Mayer & Podnar 2003 Polovic & Cadenovic 2014 Genus Podarcis (Wall Lizards)   Podarcis melisellensis Dalmatian Wall Lizard Arnold & Ovenden 2002 Brelih & Dzukic 1974 Jovanovic 2009 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Mayer & Podnar 2003 Speybroeck, Beukema, Bok & Voort 2016 Tiedemann & Henle in Boehme (ed.) 1986 Trapp 2011   Podarcis muralis Common European Wall Lizard Brelih & Dzukic 1974 Gruschwitz & Boehme in Boehme (ed.) 1986 Jovanovic 2009 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Mayer & Podnar 2003 Polovic & Cadenovic 2014 Trapp 2011   Podarcis siculus Italian Wall Lizard Brelih & Dzukic 1974 Henle & Klaver in Boehme (ed.) 1986 Jovanovic 2009 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Mayer & Podnar 2003 Podnar, Mayer & Tvrtkovic 2005 Genus Zootoca (Eurasian Forest Lizards)   Zootoca vivipara Viviparous Lizard Brelih & Dzukic 1974 Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Pozzi 1966  Family Scincidae (Skinks) Genus Ablepharus (Eurasian Snake-eyed Skinks)   Ablepharus kitaibelii European Snake-eyed Skink Ljubisavljevic, Tomovic, Urosevic, Gvozdenovic, Ikovic, Zagora & Labus 2018 Speybroeck, Beukema, Bok & Voort 2016 Vergilov, Hristov, Lukanov, Lambevska & Tzankov 2016

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Coronella (Palearctic Smooth Snakes)   Coronella austriaca Northern Smooth Snake Brelih & Dzukic 1974 Engelmann in Boehme (ed.) 1993 Geniez 2018 Jablonski, Nagy, Avci, Olgun, Kukushkin, Safaei-Mahroo & Jandzik 2019 Polovic & Cadenovic 2014 Genus Dolichophis (Greater Whip Snakes)   Dolichophis caspius Caspian Whip Snake Brelih & Dzukic 1974 Jovanovic 2009 Krcmar, Mikuska & Kletecki 2007 Polovic & Cadenovic 2014 Speybroeck, Beukema, Bok & Voort 2016 Szczerbak & Boehme in Boehme (ed.) 1993 Trapp 2011 Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe quatuorlineata Western Four-lined Rat Snake Boehme & Szczerbak in Boehme (ed.) 1993 Brelih & Dzukic 1974 Geniez 2018 Schulz 1996 Genus Hierophis (Eurasian Whip Snakes)   Hierophis gemonensis Balkan Whip Snake Brelih & Dzukic 1974 Geniez 2018 Henle in Boehme (ed.) 1993 Genus Natrix (Eurasian Keelbacks)   Natrix natrix Common European Grass Snake Brelih & Dzukic 1974 Geniez 2018 Jovanovic 2009 Kabisch in Boehme (ed.) 1999 Kindler, Boehme, Corti, Gvozdik, Jablonski, Jandzik, Metallinou, Siroky & Fritz 2013 Polovic & Cadenovic 2014 Trapp 2011   Natrix tessellata Dice Snake Brelih & Dzukic 1974 Geniez 2018 Gruschwitz, Lenz, Mebert & Lanka in Boehme (ed.) 1999 Jovanovic 2009 Polovic & Cadenovic 2014 Trapp 2011 Genus Platyceps (Flat-headed Whip Snakes)   Platyceps najadum European Slender Whip Snake Brelih & Dzukic 1974 Geniez 2018 Speybroeck, Beukema, Bok & Voort 2016 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus fallax European Cat Snake Brelih & Dzukic 1974 Geniez 2018 Grillitsch & Grillitsch in Boehme (ed.) 1999 Speybroeck, Beukema, Bok & Voort 2016 Genus Zamenis (Western Palearctic Rat Snakes)   Zamenis longissimus Common Aesculapian Snake Brelih & Dzukic 1974 Geniez 2018 Jovanovic 2009 Polovic & Cadenovic 2014   Zamenis situla Leopard Snake Brelih & Dzukic 1974 Geniez 2018 Obst, Szczerbak & Boehme in Boehme (ed.) 1993  Family Lamprophiidae (House Snakes, etc.) Genus Malpolon (Montpellier Snakes)   Malpolon insignitus Eastern Montpellier Snake Brelih & Dzukic 1974 Geniez 2018 Haan in Boehme 1999 Jovanovic 2009  Family Xenotyphlopidae (Round-nosed Blind Snakes) Genus Xerotyphlops (Desert Blind Snakes)   Xerotyphlops vermicularis European Worm Snake Arnold & Ovenden 2002 Brelih & Dzukic 1974 Geniez 2018 Grillitsch & Grillitsch in Boehme (ed.) 1993 Speybroeck, Beukema, Bok & Voort 2016  Family Viperidae (Vipers) Genus Vipera (Typical Vipers or Adders)   Vipera ammodytes European Nose-horned Viper Brelih & Dzukic 1974 David & Vogel 2010 Geniez 2018 Heckes, Gruber & Stumpel in Joger & Stumpel 2005 Jelic, Ajtic, Sterijovski, Crnobrnja-Isailovic, Lelo & Tomovic 2013 Jovanovic 2009 McDiarmid, Campbell & Toure 1999 Polovic & Cadenovic 2014 Tomovic & Dzukic 2003 Trapp 2011 Ursenbacher, Schweiger, Tomovic, Crnobrnja-Isailovic, Fumagalli & Mayer 2008   Vipera berus Northern Eurasian Viper Brelih & Dzukic 1974 Crnobrnja-Isailovic, Tomovic & Ajtic 2003 David & Vogel 2010 Geniez 2018 Nilson, Andren & Voelkl in Joger & Stumpel 2005 Polovic & Cadenovic 2014 Trapp 2011   Vipera ursinii Meadow and Karst Vipers Arnold & Ovenden 2002 Brelih & Dzukic 1974 Crnobrnja-Isailovic, Tomovic & Ajtic 2003 David & Vogel 2010 Dely & Joger in Joger & Stumpel 2005 Geniez 2018 Jelic, Ajtic, Sterijovski, Crnobrnja-Isailovic, Lelo & Tomovic 2013 Nilson & Andren 2001 Tomovic, Crnobrnja-Isailovic & Ajtic 2004 Trapp 2011 Venchi & Sindaco 2007

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Gramentz in Fritz (ed.) 2005 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Frazier, Gramentz & Fritz in Fritz (ed.) 2005 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Emydidae (Western Pond Turtles) Genus Emys (European Pond Turtles)   Emys orbicularis Common European Pond Turtle Brelih & Dzukic 1974 Fritz in Fritz (ed.) 2001 Jovanovic 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Geoemydidae (Eastern Pond Turtles) Genus Mauremys (Stripe-necked Turtles)   Mauremys rivulata Balkan Pond Turtle Arnold & Ovenden 2002 Brelih & Dzukic 1974 Mantziou & Rifai 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Speybroeck, Beukema, Bok & Voort 2016 Wischuf & Busack in Fritz (ed.) 2001  Family Testudinidae (Tortoises) Genus Testudo (Palearctic Tortoises)   Testudo graeca Spur-thighed Tortoise Tomovic, Ajtic, Dokovic & Antic 2004   Testudo hermanni Greek Tortoise Bertolero, Cheylan, Hailey, Livoreil & Willemsen 2011 Bonin, Devaux & Dupre 2006 Brelih & Dzukic 1974 Cheylan in Fritz (ed.) 2001 Jovanovic 2009 Ljubisavljevic, Dzukic, Vukov & Kalezic 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Speybroeck, Beukema, Bok & Voort 2016 Trapp 2011