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© Rune Midtgaard


Taxonomy of the family Colubridae
Bibliography of the genus Fowlea
Biodiversity of the family Colubridae








Genus
Fowlea

Fishing Keelbacks, (Oriental Painted Keelbacks, Striped Water Snakes)

Fischnattern

Fiskesnoge

1868 Fowlea Theobald (type species: Fowlea peguensis Theobald 1868)
1893 Diplophallus Cope (type species: Hydrus piscator Schneider 1799)
Contents: 7 species, of which 3 (42.9%) are endemic.
Endemism: 0% 100%
Remarks: Previously included in Natrix (e.g., Smith 1943) and, more recently, Xenochrophis (e.g., Williams & Wallach 1989). Revalidated as a separate genus by Purkayastha, Kalita, Brahma, Doley & Das (2018).
Distribution: S. and E. Asia.
Reported from: Afghanistan, Bangladesh, Bhutan, Cambodia, China (Anhui, Fujian, Guangdong [incl. Nan Ao Island], Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Jiangsu, Jiangxi, Tibet, Yunnan, Zhejiang), India (Andaman & Nicobar Islands [Andaman Islands], Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Chhattisgarh, Delhi, Goa, Gujarat, Himachal Pradesh, Jammu & Kashmir, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Punjab, Rajasthan, Sikkim, Tamil Nadu, Telangana, Tripura, Uttar Pradesh, Uttarakhand, West Bengal), Indonesia (Bali, Java, Sumatra), Laos, Malaysia (West Malaysia), Myanmar, Nepal, Pakistan, Sri Lanka, Taiwan, Thailand, Vietnam (incl. Cat Ba Island).

Fowlea asperrimus

Sri Lankan Painted Keelback

Srilankanische Fischnatter

Srilankansk Fiskesnog

1891 Tropidonotus asperrimus Boulenger
Natrix piscator asperrimus Smith 1943
Xenochrophis asperrimus Welch 1988
Fowlea asperrimus Purkayastha, Kalita, Brahma, Doley & Das 2018

Remarks: Validity as a distinct species confirmed by Vogel & David (2006).
Distribution: Sri Lanka.


Fowlea flavipunctatus

Yellow-spotted Keelback, Common Keelback

Gelbfleckiger Fischnatter

Gulplettet Fiskesnog

1860 Amphiesma flavipunctatum Hallowell
Natrix piscator flavipunctata Smith 1943
Natrix flavipunctata Taylor 1965
Xenochrophis flavipunctatus Welch 1988
Fowlea flavipunctatus Purkayastha, Kalita, Brahma, Doley & Das 2018
1907 Tropidonotus piscator ornata Wall (Vogel & David 2012)

Remarks: Previously regarded as a synonym of piscator. Taylor (1965) separated the two species, based on sympatry in Thailand. Validity confirmed by Vogel & David (2006). It is still not clear, however, whether one or both species are present in parts of the range of piscator (in the strict sense) (see for instance Zhao & Adler 1993). A record from Assam (India) (Smith 1943) refer to schnurrenbergeri (Vogel & David 2006). Listed for India by Aengals, Kumar, Palot & Ganesh (2018), but without discussion or references. Das & Yaakob (2007) included Sarawak in the distribution of flavipunctatus, but the species was not regarded as a Bornean species by Stuebing, Inger & Lardner (2014), which is followed herein. A record of piscator from Shaanxi (Song 2002), presumably referring to flavipunctatus, needs confirmation.
Introduced to: Singapore.
Distribution: Cambodia, China (Anhui, Fujian, Guangdong [incl. Nan Ao Island], Guangxi, Guizhou, Hainan, Hong Kong, Hunan, Jiangsu, Jiangxi, Tibet, Yunnan, Zhejiang), Laos, Malaysia (West Malaysia), Myanmar, Taiwan, Thailand, Vietnam (incl. Cat Ba Island).


Fowlea flavipunctatus
© Rune Midtgaard

Fowlea melanzostus

Southern Keelback, Javan Keelback

Südliche Fischnatter

Sydlig Fiskesnog

1807 Coluber Melanzostus Gravenhorst
Tropidonotus melanozostus Boie 1827 [unjustified emendation]
Tropidonotus quincunciatus melanozostus Müller 1890
Tropidonotus piscator melanozosta Boettger 1892
Natrix piscator melanzostus Smith 1943
Xenochrophis piscator melanzostus Bosch 1985
Xenochrophis flavipunctatus melanozostus Manthey & Grossmann 1997
Xenochrophis melanzostus Das 1999
Fowlea melanzostus Purkayastha, Kalita, Brahma, Doley & Das 2018
1834 Coluber lippus Reuss (Vogel & David 2012)

Remarks: Validity as a distinct species confirmed by Vogel & David (2006). Reports from the Andaman Islands (India) refer to tytleri (Vogel & David 2006). Presence in Sulawesi needs confirmation, since only a few voucher specimens are available from the island (Lang & Vogel 2005). Reports from Borneo are erroneous (Vogel & David 2006).
Distribution: Indonesia (Bali, Java, Sumatra).


Fowlea piscator

Checkered Keelback, Common Pond Snake, Asian Fishing Snake, Fish Snake, (Asian Water Snake)

Schachbrett-Fischnatter

Indisk Fiskesnog

1799 Hydrus piscator Schneider
Tropidonotus piscator Boie 1827
Nerodia piscator Wall 1923
Natrix piscator Smith 1930
Fowlea piscator Malnate 1960
Xenochrophis piscator Malnate & Minton 1965
1799 Hydrus palustris Schneider (Vogel & David 2012)
1803 Coluber anostomosatus Daudin (Vogel & David 2012)
1803 Coluber braminus Daudin (Vogel & David 2012)
1803 Coluber dora Daudin (Vogel & David 2012)
1803 Coluber mortuarius Daudin (Vogel & David 2012)
1803 Coluber umbratus Daudin (Vogel & David 2012)
1834 Coluber bengalensis Gray (Vogel & David 2012)
1834 Coluber rectangulus Gray (Vogel & David 2012)
1837 Tropidonotus quincunciatus Schlegel (Vogel & David 2012)
1890 Tropidonotus sancti-johannis Boulenger (Vogel & David 2012)
Nerodia piscator sancti-johannis Bourret 1935
Natrix piscator sancti-johannis Bourret 1936
Natrix sancti-johannis Khan 1984
Xenochrophis piscator sanctijohannis Kramer 1977
Xenochrophis sanctijohannis Welch 1988
Fowlea sanctijohannis Purkayastha, Kalita, Brahma, Doley & Das 2018
1907 Tropidonotus piscator lateralis Wall (Vogel & David 2012)
1907 Tropidonotus piscator obscurus Wall (Vogel & David 2012)
1907 Tropidonotus piscator punctatus Wall (Vogel & David 2012)
1907 Tropidonotus piscator unicolor Wall (Vogel & David 2012)

Other common names:
sanctijohannis: St. John's Keelback, Olive Water Snake
Remarks: Previously included asperrimus, flavipunctatus, melanzostus, and tytleri. Records from the Andaman Islands refer to tytleri. Records from Cambodia, Singapore, Vietnam, West Malaysia, some records from Laos, and most Chinese records refer to flavipunctatus. Records from Indonesia refer to melanzostus. A record from Shaanxi (Song 2002), presumably referring to flavipunctatus, needs confirmation. Records from Sri Lanka (apart from asperrimus) probably represents an undescribed species, which may be conspecific with southern Indian populations of piscator. The latter species, as currently understood, may even represent more than two different species (Vogel & David 2012). The taxon sanctijohannis was regarded as a synonym of piscator (e.g., Wall 1923; Pope 1923; Smith 1943), revalidated as a separate species by Khan (1984), but most recently considered a montane colour morph of, and placed in the synonymy of, piscator (Vogel & David 2012). Nevertheless, some authors (e.g.; Das & Das 2017; Purkayastha, Kalita, Brahma, Doley & Das 2018) continue to list sanctijohannis as a valid species, but without discussion.
Introduced to: Singapore.
Distribution: Afghanistan, Bangladesh, Bhutan, China (Guangxi, Yunnan), India (Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Chhattisgarh, Delhi, Goa, Gujarat, Himachal Pradesh, Jammu & Kashmir, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Punjab, Rajasthan, Sikkim, Tamil Nadu, Telangana, Tripura, Uttar Pradesh, Uttarakhand, West Bengal), Laos, Myanmar, Nepal, Pakistan, Sri Lanka, Thailand.


Fowlea punctulatus

Burmese Spotted Keelback, (Spotted Keelback)

Burmesische Gefleckte Fischnatter

Burmesisk Plettet Fiskesnog

1858 Tropidonotus punctulatus Günther
Nerodia punctulata Wall 1923
Natrix punctulata Smith 1943
Fowlea punctulata Malnate 1960
Xenochrophis punctulatus Welch 1988
1868 Fowlea peguensis Theobald (Smith 1943)

Remarks: Records from India (e.g., Borang, Bhatt, Chaudhury, Borkotoki & Bhutia 2005; Sanyal & Gayen 2006) appear to be undocumented.
Distribution: Myanmar, Thailand.


Fowlea schnurrenbergeri

Bar-necked Keelback, (Kramer's Keelback)

Nackenband-Fischnatter

Nordvestlig Fiskesnog

1977 Xenochrophis flavipunctatus schnurrenbergeri Kramer
Xenochrophis schnurrenbergeri Vogel & David 2006
Fowlea schnurrenbergeri Purkayastha, Kalita, Brahma, Doley & Das 2018

Distribution: India (Assam, Bihar, Odisha, Uttar Pradesh, West Bengal), Nepal, Pakistan.


Fowlea tytleri

Andaman Keelback

Andamanische Fischnatter

Andamansk Fiskesnog

1863 Tropidonotus tytleri Blyth
Xenochrophis tytleri Vogel & David 2006
Fowlea tytleri Purkayastha, Kalita, Brahma, Doley & Das 2018
1868 Tropidonotus striolatus Blyth (Vogel & David 2012)

Remarks: Although reported from the Nicobar Islands (Biswas 1984), Vogel & David (2012) regarded the presence of tytleri in these islands as doubtful.
Distribution: India (Andaman & Nicobar Islands: Andaman Islands).