Reptiles of
Hong Kong [China]

Related checklists:  China

Bibliography of the reptiles of Hong Kong Taxonomic history of the reptiles of Hong Kong Introduced species
Home - Taxonomy - Geography - Biodiversity - Literature - Purchase RepFocus Recent updates
Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist
General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial.
Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:
Lizards (Sauria)
= endemicEX = extinct or assumed extinct

Species
References
 Family Agamidae (Agamas and Dragons)
Genus Calotes (South Asian Forest Agamas)
 
Calotes versicolor

South Indian Crested Agama

Ananjeva, Guo & Wang 2011 Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Karsen, Lau & Bogadek 1998 Lazell 1999 Smith 1935 Taylor 1963 Wermuth 1967 Zhao & Adler 1993
 Family Dibamidae (Blind Skinks)
Genus Dibamus (Oriental Blind Skinks)
  *
Dibamus bogadeki

Hong Kong Blind Skink

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Ng, Yeung, Cheung, Lam, Fung & Chan 2012 Darevsky 1992 Karsen, Lau & Bogadek 1998 Lazell 1999 Lazell 2002 Zhao & Adler 1993
 
Dibamus bourreti

Indochinese Blind Skink

Ineich 1999
 Family Gekkonidae (Typical Geckos)
Genus Gehyra (Four-clawed Geckos)
 
Gehyra mutilata

Common Four-clawed Gecko

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Cheung, Ho, Lam & Tang 2006 Karsen, Lau & Bogadek 1998 Lazell 1999 Zhao & Adler 1993
Genus Gekko (Tokays)
 
Gekko chinensis

Chinese Narrow-disked Gecko

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Cheung, Ho, Lam & Tang 2006 Karsen, Lau & Bogadek 1998 Lazell 1999 Lazell 2002 Lyu, Lin, Ren, Jiang, Zhang, Qi & Wang 2021 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Zhao & Adler 1993
 
Gekko gecko

Tokay

Bogadek & Lau 1997 Chan, Cheung, Ho, Lam & Tang 2006 Karsen, Lau & Bogadek 1998 Lazell 1999 Zhao & Adler 1993
 
Gekko reevesii

Chinese Black Tokay

Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021
Genus Hemidactylus (Half-toed Geckos)
 
Hemidactylus bowringii

Hong Kong House Gecko

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Cheung, Ho, Lam & Tang 2006 Karsen, Lau & Bogadek 1998 Lazell 1999 Lazell 2002 McMahan & Zug 2007 Smith 1935 Zhao & Adler 1993
 
Hemidactylus frenatus

Asian House Gecko

Bogadek & Lau 1997
 
Hemidactylus garnotii

Fox Gecko

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Cheung, Ho, Lam & Tang 2006 Karsen, Lau & Bogadek 1998 Lazell 1999 Lazell 2002 Zhao & Adler 1993
Genus Hemiphyllodactylus (Slender Geckos)
  *
Hemiphyllodactylus hongkongensis

Hong Kong Slender Gecko

Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Cheung, Ho, Lam & Tang 2006 [as H. sp.] Lazell 2002 [as chapaensis] Sung, Lee, Ng, Zhang & Yang 2018
 Family Lacertidae (Typical Lizards)
Genus Takydromus (Oriental Grass Lizards)
 
Takydromus sexlineatus

Six-lined Grass Lizard

Arnold 1997 Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Karsen, Lau & Bogadek 1998 Lazell 1999 Zhao & Adler 1993
 Family Scincidae (Skinks)
Genus Ateuchosaurus (East Asian Short-legged Skinks)
 
Ateuchosaurus chinensis

Chinese Short-Legged Skink

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Chan, Cheung, Ho, Ng & Tang 2009 Karsen, Lau & Bogadek 1998 Lazell 1999 Zhao & Adler 1993
Genus Eutropis (South Asian Sun Skinks)
 
Eutropis longicaudata

Long-tailed Sun Skink

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Chan, Cheung, Ho, Ng & Tang 2009 Karsen, Lau & Bogadek 1998 Lazell 1999 Zhao & Adler 1993
Genus Plestiodon (Blue-tailed Skinks)
 
Plestiodon chinensis

Eastern Chinese Long Skink

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Chan, Cheung, Ho, Ng & Tang 2009 Karsen, Lau & Bogadek 1998 Lazell 1999 Zhao & Adler 1993
 
Plestiodon elegans

Elegant Five-lined Skink

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Chan, Cheung, Ho, Ng & Tang 2009 Karsen, Lau & Bogadek 1998 Lazell 1999 Zhao & Adler 1993
 
Plestiodon quadrilineatus

Four-striped Long Skink

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Chan, Cheung, Ho, Ng & Tang 2009 Karsen, Lau & Bogadek 1998 Lazell 1999 Lazell 2002 Zhao & Adler 1993
 
Plestiodon tamdaoensis

Tamdao Blue-tailed Skink

Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Chan, Cheung, Ho, Ng & Tang 2009 Hikida, Lau & Ota 2001
Genus Scincella (South Asian and American Ground Skinks)
 
Scincella reevesii

Speckled Ground Skink

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Chan, Cheung, Ho, Ng & Tang 2009 Chen, Hikida, Han, Shim, Oh & Ota 2001 Lazell 1999 Lazell 2002 Ouboter 1986 Zhao & Adler 1993
Genus Sphenomorphus (Indo-Pacific Forest Skinks)
 
Sphenomorphus incognitus

Koshun Forest Skink

Chan, Chan, Cheung, Ho, Ng & Tang 2009
 
Sphenomorphus indicus

Common Indian Forest Skink

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Chan, Cheung, Ho, Ng & Tang 2009 Karsen, Lau & Bogadek 1998 Lazell 1999 Zhao & Adler 1993
Genus Tropidophorus (Stream Skinks)
 
Tropidophorus sinicus

South Chinese Stream Skink

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Chan, Chan, Cheung, Ho, Ng & Tang 2009 Guo, Shu, Wu, Chen, Hou, Shi & Deng 2021 Karsen, Lau & Bogadek 1998 Lazell 1999 Zhao & Adler 1993
 Family Varanidae (Monitor Lizards)
Genus Varanus (Monitor Lizards)
 
Varanus salvator

Asian Water Monitor

Bogadek & Lau 1997 Cai, Ji, Wangs, Rao, Huang, Wang, Song, Guo & Jiang 2021 Karsen, Lau & Bogadek 1998 Lazell 1999 Zhao & Adler 1993

Snakes (Serpentes)
= endemicEX = extinct or assumed extinct

Species
References
 Family Colubridae (Typical Snakes or Colubrids)
Genus Ahaetulla (Oriental Whip Snakes)
 
Ahaetulla prasina

Short-nosed Whip Snake

Karsen, Lau & Bogadek 1998 Lazell 2002 Wallach, Williams & Boundy 2014
Genus Amphiesma (South Asian Keelbacks)
 
Amphiesma stolatum

Buff-striped Keelback

Karsen, Lau & Bogadek 1998 Wallach, Williams & Boundy 2014 Zhao & Adler 1993
Genus Boiga (Australasian Cat Snakes)
 
Boiga multomaculata

Marbled Cat Snake

Karsen, Lau & Bogadek 1998 Lazell 2002 Orlov & Ryabov 2002 Zhao & Adler 1993
Genus Calamaria (Oriental Reed Snakes)
 
Calamaria septentrionalis

Northern Reed Snake

Chan, Fung, Ng & Wong 2011 Inger & Marx 1965 Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
Genus Chrysopelea (Flying Tree Snakes)
 
Chrysopelea ornata

Ornate Flying Tree Snake

Zhao & Adler 1993
Genus Coelognathus (Oriental Rat Snakes)
 
Coelognathus radiatus

Copper-headed Rat Snake

Karsen, Lau & Bogadek 1998 Lazell 2002 Zhao & Adler 1993
Genus Dendrelaphis (Bronzebacks)
  *
Dendrelaphis hollinrakei

Hong Kong Bronzeback

Lazell 2002
 
Dendrelaphis proarchos

Northern Painted Bronzeback

Karsen, Lau & Bogadek 1998 Zhao & Adler 1993 [as pictus]
Genus Fowlea (Fishing Keelbacks)
 
Fowlea flavipunctata

Yellow-spotted Keelback

Karsen, Lau & Bogadek 1998 [as piscator] Smith 1943 Vogel & David 2012 Zhao & Adler 1993 [as piscator]
Genus Hebius (Oriental Equatorial Keelbacks)
 
Hebius atemporalis

Tonkin Keelback

Chan, Fung, Ng & Wong 2011 Karsen, Lau & Bogadek 1998 Wallach, Williams & Boundy 2014 Zhao & Adler 1993
 
Hebius boulengeri

White-browed Keelback

Karsen, Lau & Bogadek 1998
Genus Lycodon (Oriental Wolf Snakes)
 
Lycodon capucinus

Island Wolf Snake

Karsen, Lau & Bogadek 1998 [as aulicus] Lazell 2002 Leviton 1965 O'Shea, Kusuma & Kaiser 2018 Smith 1943 Zhao & Adler 1993 [as Ophites aulicus]
 
Lycodon futsingensis

Fujian Wolf Snake

Karsen, Lau & Bogadek 1998 [as ruhstrati] Vogel, David, Pauwels, Sumontha, Norval, Hendrix, Vu & Ziegler 2009 Vogel, Nguyen, Kingsada & Ziegler 2012 Zhang, Jiang, Vogel & Rao 2011 Zhao & Adler 1993 [as ruhstrati]
 
Lycodon subcinctus

Malayan Banded Wolf Snake

Karsen, Lau & Bogadek 1998 Zhang, Jiang, Vogel & Rao 2011 Zhao & Adler 1993
Genus Oligodon (Kukri Snakes)
 
Oligodon cinereus

Black-barred Kukri Snake

David, Nguyen, Nguyen, Jiang, Chen, Teynie & Ziegler 2012 Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
 
Oligodon formosanus

Taiwanese Kukri Snake

Karsen, Lau & Bogadek 1998 Lazell 2002 Zhao & Adler 1993
Genus Opisthotropis (Mountain Stream Keelbacks)
 
Opisthotropis andersonii

Hong Kong Mountain Stream Keelback

David, Pauwels, Nguyen & Vogel 2015 Karsen, Lau & Bogadek 1998 Ren, Wang, Guo, Wang, Nguyen & Li 2019 Ren, Wang, Jiang, Guo & Li 2017 Smith 1943 Wang, Guo, Liu, Lyu, Wang, Luo, Sun & Zhang 2017 Yang, Wang, Zhang, Lau & Chou 2011 Zhao & Adler 1993
 
Opisthotropis kuatunensis

Striped Mountain Stream Keelback

Karsen, Lau & Bogadek 1998 Ren, Wang, Guo, Wang, Nguyen & Li 2019 Zhao & Adler 1993
 
Opisthotropis lateralis

Bicoloured Mountain Stream Keelback

Karsen, Lau & Bogadek 1998 Ren, Wang, Guo, Wang, Nguyen & Li 2019 Zhao & Adler 1993
Genus Oreocryptophis (Red Mountain Rat Snake)
 
Oreocryptophis porphyraceus

Red Mountain Rat Snake

Karsen, Lau & Bogadek 1998 Schulz & Helfenberger 1998 Zhao & Adler 1993
Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)
 
Ptyas korros

Indochinese Rat Snake

Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
 
Ptyas major

Greater Green Snake

Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
 
Ptyas mucosa

Common Oriental Rat Snake

Karsen, Lau & Bogadek 1998 Lazell 2002 Zhao & Adler 1993
Genus Rhabdophis (Asian Keelbacks)
 
Rhabdophis callichroma

Bavi Keelback

Karsen, Lau & Bogadek 1998 [as chrysargos]
 
Rhabdophis helleri

Northern Red-necked Keelback

David & Vogel 2021 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
Genus Sibynophis (Oriental Black-headed Snakes)
 
Sibynophis chinensis

Chinese Black-headed Snake

Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
Genus Trimerodytes (Asian Water Snakes)
 
Trimerodytes aequifasciatus

Equal-banded Water Snake

Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
 
Trimerodytes balteatus

Banded Mountain Stream Keelback

Karsen, Lau & Bogadek 1998 Ren, Wang, Guo, Wang, Nguyen & Li 2019 Zhao & Adler 1993
 
Trimerodytes percarinatus

Chinese Keelback

Guo, Zhu, Liu, Wang, Che & Nguyen 2020 Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
 Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.)
Genus Bungarus (Kraits)
 
Bungarus fasciatus

Banded Krait

Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
 
Bungarus multicinctus

Many-banded Krait

Karsen, Lau & Bogadek 1998 Lazell 2002 Vogel 2006 Zhao & Adler 1993
Genus Naja (Typical Cobras)
 
Naja atra

Chinese Spitting Cobra

Karsen, Lau & Bogadek 1998 Lazell 2002 Vogel 2006 Zhao & Adler 1993
Genus Ophiophagus (King Cobra)
 
Ophiophagus hannah

King Cobra

Karsen, Lau & Bogadek 1998 Vogel 2006 Zhao & Adler 1993
Genus Sinomicrurus (East Asian Coral Snakes)
 
Sinomicrurus annularis

Hong Kong Coral Snake

Smart, Ingrasci, Sarker, Lalremsanga, Murphy, Ota, Tu, Shouche, Orlov & Smith 2021
 Family Elapidae [part 2: marine species] (Sea Snakes)
Genus Hydrophis (Typical Sea Snakes)
 
Hydrophis curtus

Short Sea Snake

Karsen, Lau & Bogadek 1998 Smith 1926 [as Lapemis hardwickii] Zhao & Adler 1993
 
Hydrophis cyanocinctus

Blue-banded Sea Snake

Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
 
Hydrophis ornatus

Reef Sea Snake

Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
 
Hydrophis platurus

Yellow-bellied Sea Snake

Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
 
Hydrophis viperinus

Viperine Sea Snake

Karsen, Lau & Bogadek 1998 Smith 1926 Zhao & Adler 1993
Genus Microcephalophis (Small-headed Sea Snakes)
 
Microcephalophis gracilis

Graceful Small-headed Sea Snake

Karsen, Lau & Bogadek 1998 Smith 1926 Zhao & Adler 1993
 Family Homalopsidae (Australasian Mud Snakes)
Genus Hypsiscopus (Orange-bellied Mud Snakes and Matanna Mud Snake)
 
Hypsiscopus murphyi

Northern Orange-bellied Mud Snake

Gyi 1970 [as plumbea] Karsen, Lau & Bogadek 1998 [as plumbea] Lazell 2002 [as plumbea] Zhao & Adler 1993 [as plumbea]
Genus Myrrophis (Indochinese Mud Snakes)
 
Myrrophis bennetti

Mangrove Mud Snake

Gyi 1970 Karsen, Lau & Bogadek 1998 Murphy & Voris 2014 Zhao & Adler 1993
 
Myrrophis chinensis

Chinese Mud Snake

Karsen, Lau & Bogadek 1998 Murphy & Voris 2014 Murphy 2007 Zhao & Adler 1993
 Family Lamprophiidae (House Snakes, etc.)
Genus Psammodynastes (Asian Mock Vipers)
 
Psammodynastes pulverulentus

Dusky Mock Viper

Karsen, Lau & Bogadek 1998 Lazell 2002 Zhao & Adler 1993
 Family Pareidae (Oriental Slug Eaters)
Genus Pareas (Typical Oriental Slug Eaters)
 
Pareas chinensis

Chinese Slug Eater

Karsen, Lau & Bogadek 1998 Ota, Bogadek & Lau 1997 Zhao & Adler 1993 [as hamptoni]
 
Pareas margaritophorus

White-spotted Slug Eater

Hauser 2017 Karsen, Lau & Bogadek 1998 Vogel, Nguyen, Lalremsanga, Biakzuala, Hrima & Poyarkov 2020 Yang, Yeung, Huang & Yang 2021 Zhao & Adler 1993
 Family Pythonidae (Pythons)
Genus Python (Typical Pythons)
 
Python bivittatus

Burmese Python

Barker & Barker 2010 Karsen, Lau & Bogadek 1998 Lazell 2002 Zhao & Adler 1993
 Family Typhlopidae (Typical Blind Snakes)
Genus Indotyphlops (South Asian Blind Snakes)
  *
Indotyphlops lazelli

Hong Kong Blind Snake

Wallach & Pauwels 2004
Genus Ramphotyphlops (Western Pacific Blind Snakes)
 
Ramphotyphlops albiceps

Southeast Asian White-headed Blind Snake

Karsen, Lau & Bogadek 1998 McDiarmid, Campbell & Toure 1999 Zhao & Adler 1993
Genus Virgotyphlops (Brahminy Blind Snake)
 
Virgotyphlops braminus

Brahminy Blind Snake

Karsen, Lau & Bogadek 1998 Lazell 2002 McDiarmid, Campbell & Toure 1999 Wallach 2020 Zhao & Adler 1993
 Family Viperidae (Vipers)
Genus Ovophis (Asian Mountain Pitvipers)
 
Ovophis makazayazaya

Eastern Mountain Pitviper

Karsen, Lau & Bogadek 1998 Vogel 2006 [as orientalis] Zhao & Adler 1993
 
Ovophis tonkinensis

Tonkin Pitviper

Fong, Grioni, Crow & Cheung 2017
Genus Trimeresurus (Oriental Pitvipers)
 
Trimeresurus albolabris

Southeast Asian White-lipped Tree Pitviper

Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Karsen, Lau & Bogadek 1998 Lazell 2002 Regenass & Kramer 1981 Vogel 2006 Zhao & Adler 1993
 Family Xenodermidae (Asian Litter Snakes)
Genus Achalinus (East Asian Burrowing Snakes)
 
Achalinus rufescens

Rufous Burrowing Snake

Karsen, Lau & Bogadek 1998 Li, Yu, Wu, Liao, Tang, Liu & Guo 2020 Wallach, Williams & Boundy 2014 Zhao & Adler 1993

Turtles (Testudines)
= endemicEX = extinct or assumed extinct

Species
References
 Family Cheloniidae (Hard-shelled Sea Turtles)
Genus Chelonia (Green Sea Turtle)
 
Chelonia mydas

Green Sea Turtle

Karsen, Lau & Bogadek 1998 King & Burke 1989 Ng & Wan 2011 Zhao & Adler 1993
Genus Lepidochelys (Ridley Sea Turtles)
 
Lepidochelys olivacea

Olive Ridley Sea Turtle

Karsen, Lau & Bogadek 1998 Zhao & Adler 1993
 Family Dermochelyidae (Leatherback Sea Turtle)
Genus Dermochelys (Leatherback Sea Turtle)
 
Dermochelys coriacea

Leatherback Sea Turtle

Karsen, Lau & Bogadek 1998 King & Burke 1989 Zhao & Adler 1993
 Family Geoemydidae (Eastern Pond Turtles)
Genus Cuora (Asian Box Turtles)
 
Cuora flavomarginata

Chinese Yellow-margined Box Turtle

Karsen, Lau & Bogadek 1998 Lau, Chan, Crow & Ades 2000 Ota, Yasukawa, Fu & Chen 2009 Zhao & Adler 1993
 
Cuora trifasciata

Northern Three-striped Box Turtle

Fritz & Havas 2006 Karsen, Lau & Bogadek 1998 Lau, Chan, Crow & Ades 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993
Genus Mauremys (Stripe-necked Turtles)
 
Mauremys reevesii

Chinese Three-keeled Pond Turtle

Karsen, Lau & Bogadek 1998 Lazell 2002 Ng 2010 [as megalocephala] Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993
Genus Sacalia (Eastern Eyed Turtles)
 
Sacalia bealei

Eastern Chinese Eyed Turtle

Karsen, Lau & Bogadek 1998 Lau, Chan, Crow & Ades 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993
 Family Platysternidae (Big-headed Turtle)
Genus Platysternon (Big-headed Turtle)
 
Platysternon megacephalum

Big-headed Turtle

Karsen, Lau & Bogadek 1998 Lau, Chan, Crow & Ades 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993
 Family Trionychidae (Softshell Turtles)
Genus Pelodiscus (East Asian Softshell Turtles)
 
Pelodiscus sinensis

Chinese Softshell Turtle

Karsen, Lau & Bogadek 1998 Lau, Chan, Crow & Ades 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Zhao & Adler 1993