Reptiles of Goa [India] Related checklists:  India Bibliography of the reptiles of Goa Taxonomic history of the reptiles of Goa Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus palustris Mugger Pradhan 2008 Whitaker & Andrews 2003

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Calotes (South Asian Forest Agamas)   Calotes versicolor South Indian Crested Agama Manacas 1961 Pradhan 2008 Sharma 1976 Genus Draco (Gliding Dragons)   Draco dussumieri South Indian Gliding Dragon Daniel 1983 Murthy 1986 Pradhan 2008 Sharma 1976 Tikader & Sharma 1992 Wermuth 1967 Genus Monilesaurus (Necklace Forest Agamas)   Monilesaurus rouxii Central Indian Forest Agama Pal, Vijayakumar, Shanker, Jayarajan & Deepak 2018 Pradhan 2008 Sharma 1976 Sharma 1982 Tikader & Sharma 1992 Genus Psammophilus (Indian Rock Agamas)   Psammophilus dorsalis South Indian Rock Agama Pradhan 2008 Genus Sitana (Lesser Fan-throated Agamas)   Sitana sp. (Fan-throated Agama) Pradhan 2008 [as ponticeriana]  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo zeylanicus Asian Chameleon Pradhan 2008  Family Gekkonidae (Typical Geckos) Genus Cnemaspis (Asian Rock and Forest Geckos)   Cnemaspis flaviventralis Yellow-bellied Day Gecko Pal, Mirza, Dsouza & Shanker 2021   Cnemaspis goaensis Goa Forest Gecko Murthy 1985 Pal, Mirza, Dsouza & Shanker 2021 Pradhan 2008 [as kandiana] Sayyed, Pyron & Dahanukar 2016 Sayyed, Pyron & Dileepkumar 2018 Sharma 1976 Tikader & Sharma 1992   Cnemaspis gracilis Gracile Rock Gecko Sayyed, Pyron & Dahanukar 2016   * Cnemaspis heteropholis Strange-scaled Rock Gecko Khandekar, Thackeray, Pal & Agarwal 2020 Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus albofasciatus White-banded Rock Gecko Bauer & Giri 2004 Pradhan 2008 Sharma 1976 Tikader & Sharma 1992   Cyrtodactylus deccanensis Deccan Bent-toed Gecko Lewis, Piggott, Griffin, Greig-Smith, Martin, Barretto, Bajibab, Thorpe-Dixon, Prodromou, Fordham, Willis, Turner, Radovanovic & Holloway 2010 Pradhan 2008   Cyrtodactylus varadgirii Mumbai Bent-toed Gecko Pradhan 2008 [as collegalensis] Genus Hemidactylus (Half-toed Geckos)   Hemidactylus brookii Spotted Half-toed Gecko Pradhan 2008 Sharma 1976   Hemidactylus flaviviridis Yellow-bellied House Gecko Pradhan 2008   Hemidactylus frenatus Asian House Gecko Pradhan 2008 Sharma 1976   Hemidactylus leschenaultii Tamarind Tree Gecko Pradhan 2008   Hemidactylus murrayi Pimpri Half-toed Gecko Lajmi, Giri & Karanth 2016   Hemidactylus parvimaculatus Small-spotted Half-toed Gecko Lajmi, Giri & Karanth 2016   Hemidactylus prashadi Prashad Half-toed Gecko Giri & Bauer 2006 Pradhan 2008 Sharma 1976 Tikader & Sharma 1992   Hemidactylus reticulatus Reticulated Half-toed Gecko Pradhan 2008   Hemidactylus whitakeri Karnataka Termite Hill Gecko Pradhan 2008 [as triedrus] Genus Hemiphyllodactylus (Slender Geckos)   * Hemiphyllodactylus goaensis Goa Slender Gecko Khandekar, Parmar, Sawant & Agarwal 2021  Family Lacertidae (Typical Lizards) Genus Ophisops (Snake-eyed Lizards)   Ophisops beddomii Kerala Snake-eyed Lizard Pradhan 2008 Sharma 1976  Family Scincidae (Skinks) Genus Eutropis (South Asian Sun Skinks)   Eutropis allapallensis Allapalli Sun Skink Bag, Raha & Deuti 2020 Ganesh & Arumugam 2016 Pradhan 2008 Sharma 1976 Srinivasulu & Srinivasulu 2013 Tikader & Sharma 1992   Eutropis beddomii South Indian Sun Skink Pradhan 2008   Eutropis carinata Brahminy Skink Amarasinghe, Ganesh, Patel, Deuti, Campbell, Palot, Karunarathna, Abinawanto & Supriatna 2021 Pradhan 2008 Sharma 1976   Eutropis macularia Speckled Forest Skink Pradhan 2008 Sharma 1976   Eutropis vertebralis Striped Ground Skink Pradhan 2008 [as >trivittata] Genus Riopa (Asian Gracile Skinks)   Riopa goaensis Goa Gracile Skink Murthy 1985 Pradhan 2008 Sharma 1976 Tikader & Sharma 1992   Riopa guentheri Malabar Gracile Skink Bhilala, Ashaharraza, Ingle, Das, Sambath & Mohapatra 2021 Pradhan 2008 Sharma 1976 Tikader & Sharma 1992 Vyas 2014   Riopa lineata Indian Striped Gracile Skink Pradhan 2008   Riopa punctata Spotted Gracile Skink Pradhan 2008 Sharma 1976  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus bengalensis Bengal Monitor Pradhan 2008 Sharma 1976

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Acrochordidae (Australasian File Snakes) Genus Acrochordus (Australasian File Snakes)   Acrochordus granulatus Lesser File Snake Pradhan 2008 Sharma 1976  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx conicus Rough-scaled Sand Boa Pradhan 2008 Sharma 1976   Eryx johnii Red Sand Boa Pradhan 2008   Eryx whitakeri Kerala Sand Boa Pradhan 2008 Whitaker & Captain 2004  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla borealis Northwest Indian Whip Snake Pradhan 2008 [as nasuta] Sharma 1976 [as nasuta] Wallach, Williams & Boundy 2014 [as nasuta]   Ahaetulla sahyadrensis Sahyadri Hills Whip Snake Pradhan 2008 [as pulverulenta] Sharma 1976 [as pulverulenta] Genus Amphiesma (South Asian Keelbacks)   Amphiesma monticolum Yellow-collared Forest Keelback Pradhan 2008   Amphiesma stolatum Buff-striped Keelback Pradhan 2008 Sharma 1976 Genus Boiga (Australasian Cat Snakes)   Boiga beddomei Yellow-bellied Cat Snake Ganesh, Mallik, Achyuthan, Shanker & Vogel 2021 Lewis, Piggott, Griffin, Greig-Smith, Martin, Barretto, Bajibab, Thorpe-Dixon, Prodromou, Fordham, Willis, Turner, Radovanovic & Holloway 2010   Boiga forsteni Speckled Cat Snake Pradhan 2008   Boiga trigonata Western Gamma Cat Snake Pradhan 2008 Sharma 1976 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea ornata Ornate Flying Tree Snake Pradhan 2008 Sharma 1976 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus helena Indian Trinket Snake Mohapatra, Schulz, Helfenberger, Hofmann & Dutta 2016 Pradhan 2008 Sharma 1976 Whitaker & Captain 2004 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis ashoki Anaimalai Bronzeback Pradhan 2008 [as pictus] Sharma 1976 [as pictus]   Dendrelaphis grandoculis Indian Large-eyed Bronzeback Pradhan 2008 Sharma 1976   Dendrelaphis tristis Common Indian Bronzeback Pradhan 2008 Genus Fowlea (Fishing Keelbacks)   Fowlea piscator Checkered Keelback Pradhan 2008 Purkayastha, Kalita, Brahma, Doley & Das 2018 Sharma 1976 Genus Hebius (Oriental Equatorial Keelbacks)   Hebius beddomei Nilgiri Keelback Lewis, Piggott, Griffin, Greig-Smith, Martin, Barretto, Bajibab, Thorpe-Dixon, Prodromou, Fordham, Willis, Turner, Radovanovic & Holloway 2010 Pradhan 2008 Sharma 1976 Genus Lycodon (Oriental Wolf Snakes)   Lycodon aulicus Indian Wolf Snake Pradhan 2008 Sharma 1976   Lycodon striatus Striped Wolf Snake Pradhan 2008   Lycodon travancoricus Travancore Wolf Snake Pradhan 2008 Genus Oligodon (Kukri Snakes)   Oligodon taeniolatus Variegated Kukri Snake Pradhan 2008 Seetharamaraju, Srinivasulu & Srinivasulu 2011 Sharma 1976   Oligodon tillacki Kurduvadi Kukri Snake Bandara, Ganesh, Kanishka, Danushka, Sharma, Campbell, Ineich, Vogel & Amarasinghe 2022 [based on map] Pradhan 2008 [as arnensis] Sharma 1976 [as arnensis] Genus Platyceps (Flat-headed Whip Snakes)   Platyceps plinii Banded Whip Snake Deepak, Narayanan, Mohapatra, Dutta, Melvinselvan, Khan, Mahlow & Tillack 2021 Pradhan 2008 [as fasciolata] Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas mucosa Common Oriental Rat Snake Pradhan 2008 Sharma 1976 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis plumbicolor Green Keelback Pradhan 2008 Sharma 1976 Genus Rhabdops (South Asian Forest Snakes)   Rhabdops aquaticus Northern Olive Forest Snake Giri, Deepak, Captain, Das, Das, Rajkumar, Rathish & Gower 2017 Genus Sibynophis (Oriental Black-headed Snakes)   Sibynophis subpunctatus Malabar Black-headed Snake Pradhan 2008  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Bungarus (Kraits)   Bungarus caeruleus Blue Krait Pradhan 2008 Sharma 1976 Vogel 2006 Genus Calliophis (South Asian Coral Snakes)   Calliophis castoe Amboli Coral Snake Jadhav, Asode, Kininge, Kurundwadkar & Yadav 2021 Smith, Ogale, Deepak & Giri 2012   Calliophis melanurus South Asian Slender Coral Snake Pradhan 2008   Calliophis nigrescens Black Slender Coral Snake Pradhan 2008 Smith, Ogale, Deepak & Giri 2012 Vogel 2006 Genus Naja (Typical Cobras)   Naja naja Indian Spectacled Cobra Pradhan 2008 Sharma 1976 Whitaker & Captain 2004 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Pradhan 2008 Whitaker & Captain 2004 Yadav & Yankanchi 2015  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis caerulescens Bluish Slender-necked Sea Snake Pradhan 2008 Sharma 1976 Sharma 1982   Hydrophis curtus Short Sea Snake Padate, Baragi & Rivonker 2009 Pradhan 2008 Sharma 1976 Sharma 1982   Hydrophis cyanocinctus Blue-banded Sea Snake Pradhan 2008   Hydrophis fasciatus Banded Slender-necked Sea Snake Pradhan 2008   Hydrophis mamillaris Bombay Gulf Sea Snake Pradhan 2008   Hydrophis ornatus Reef Sea Snake Pradhan 2008   Hydrophis platurus Yellow-bellied Sea Snake Pradhan 2008   Hydrophis schistosus Beaked Sea Snake Padate, Baragi & Rivonker 2009 Prachi & Ramesh 2016 Pradhan 2008 Sharma 1976 Sharma 1982   Hydrophis spiralis Narrow-banded Sea Snake Pradhan 2008 Genus Microcephalophis (Small-headed Sea Snakes)   Microcephalophis gracilis Graceful Small-headed Sea Snake Pradhan 2008  Family Homalopsidae (Australasian Mud Snakes) Genus Cerberus (Dog-faced Water Snakes)   Cerberus rynchops Western Dog-faced Water Snake Pradhan 2008 Genus Gerarda (Glossy Marsh Snake)   Gerarda prevostiana Glossy Marsh Snake Pradhan 2008  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python molurus Indian Python Pradhan 2008 Sharma 1976  Family Typhlopidae (Typical Blind Snakes) Genus Grypotyphlops (Indian Beaked Blind Snake)   Grypotyphlops acutus Indian Beaked Blind Snake Narayana, Venkateshwarlu, Swamy, Surender, Kumar & Rao 2018 Pradhan 2008 Sharma 1976 Wallach 1994 Genus Indotyphlops (South Asian Blind Snakes)   Indotyphlops porrectus Indian Slender Blind Snake Pradhan 2008 Genus Virgotyphlops (Brahminy Blind Snake)   Virgotyphlops braminus Brahminy Blind Snake Pradhan 2008 Sharma 1976  Family Uropeltidae (Shield-tailed Snakes) Genus Melanophidium (Black Shieldtails)   Melanophidium khairei Amboli Shieldtail Gower, Giri, Captain & Wilkinson 2016 Murthy 1985 [as punctatum] Murthy 1986 [as punctatum] Pradhan 2008 [as punctatum] Pyron, Ganesh, Sayyed, Sharma, Wallach & Somaweera 2016 Genus Uropeltis (Typical Shieldtails)   Uropeltis ceylanica Kerala Shieldtail Murthy 1985 Pradhan 2008   Uropeltis ellioti Madras Shieldtail Pradhan 2008   Uropeltis macrolepis Large-scaled Shieldtail Lobo 2006 Pradhan 2008  Family Viperidae (Vipers) Genus Craspedocephalus (Ridge-nosed Tree Pitvipers)   Craspedocephalus gramineus Indian Green Pitviper Pradhan 2008 Sawant, Jadhav & Shyama 2010 Sharma 1976   Craspedocephalus malabaricus Malabar Rock Pitviper Pradhan 2008 Sawant, Jadhav & Shyama 2010 Smith 1943 Vogel 2006 Genus Daboia (Daboias)   Daboia russelii Indian Chain Viper Pradhan 2008 Sharma 1976 Vogel 2006 Genus Echis (Carpet Vipers)   Echis carinatus Common Carpet Viper Auffenberg & Rehman 1991 Lewis, Piggott, Griffin, Greig-Smith, Martin, Barretto, Bajibab, Thorpe-Dixon, Prodromou, Fordham, Willis, Turner, Radovanovic & Holloway 2010 Pradhan 2008 Sharma 1976 Sharma 1982 Vogel 2006 Genus Hypnale (Hump-nosed Pitvipers)   Hypnale hypnale Indian Hump-nosed Pitviper Pradhan 2008 Sawant, Jadhav & Shyama 2010

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Pradhan 2008 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Giri & Chaturvedi (in Shanker & Choudhury) 2006 Pradhan 2008 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Pradhan 2008 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Das 2001 Giri & Chaturvedi (in Shanker & Choudhury) 2006 Kar & Bhaskar in Bjorndal (ed.) 1982 Pradhan 2008  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Das 2001 Giri & Chaturvedi (in Shanker & Choudhury) 2006 Kar & Bhaskar in Bjorndal (ed.) 1982 Pradhan 2008  Family Geoemydidae (Eastern Pond Turtles) Genus Melanochelys (Indian Black Turtles)   Melanochelys trijuga Indian Black Turtle Frazier & Das 1994 Pradhan 2008 Sharma 1976  Family Trionychidae (Softshell Turtles) Genus Lissemys (South Asian Flapshell Turtles)   Lissemys punctata Indian Flapshell Turtle Bhupathy, Webb & Praschag 2014 Das 2001 Pradhan 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sharma 1976 Genus Nilssonia (South Asian Softshell Turtles)   Nilssonia leithii Deccan Softshell Turtle Pradhan 2008