Reptiles of Croatia    Bibliography of the reptiles of Croatia Taxonomic history of the reptiles of Croatia Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Anguidae (Slowworms, etc.) Genus Anguis (Slowworms)   Anguis fragilis Western European Slowworm Gvozdik, Benkovsky, Crottini, Bellati, Moravec, Romano, Sacchi & Jandzik 2013 Gvozdik, Jandzik, Lymberakis, Jablonski & Moravec 2010 Jelic 2014 Mikulicek, Jablonski, Palenik, Gvozdik & Jandzik 2018 Pawlowski & Kramer 2009 Pozzi 1966 Genus Pseudopus (Eurasian Glass Lizard)   Pseudopus apodus Eurasian Glass Lizard Arnold & Ovenden 2002 Brelih & Dzukic 1974 Jelic 2014 Obst in Boehme (ed.) 1981 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020 Speybroeck, Beukema, Bok & Voort 2016  Family Gekkonidae (Typical Geckos) Genus Hemidactylus (Half-toed Geckos)   Hemidactylus turcicus Turkish Half-toed Gecko Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Salvador in Boehme (ed.) 1981 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020  Family Lacertidae (Typical Lizards) Genus Algyroides (Keeled Lizards)   Algyroides nigropunctatus Blue-throated Keeled Lizard Arnold & Ovenden 2002 Bischoff in Boehme (ed.) 1981 Brelih & Dzukic 1974 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Mayer & Podnar 2002 Speybroeck, Beukema, Bok & Voort 2016 Genus Dalmatolacerta (Sharp-snouted Rock Lizard)   Dalmatolacerta oxycephala Sharp-snouted Rock Lizard Arnold & Ovenden 2002 Bischoff in Boehme (ed.) 1984 Brelih & Dzukic 1974 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Mayer & Podnar 2002 Sindaco & Jeremchenko 2008 Speybroeck, Beukema, Bok & Voort 2016 Genus Dinarolacerta (Montenegro Rock Lizards)   Dinarolacerta mosorensis Mosor Rock Lizard Arnold & Ovenden 2002 Bischoff in Boehme (ed.) 1984 Brelih & Dzukic 1974 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Mayer & Podnar 2002 Sindaco & Jeremchenko 2008 Speybroeck, Beukema, Bok & Voort 2016 Genus Iberolacerta (Southern European Rock Lizards)   Iberolacerta horvathi Croatian Rock Lizard Arnold & Ovenden 2002 Bischoff in Boehme (ed.) 1984 Brelih & Dzukic 1974 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Pawlowski & Kramer 2009 Sindaco & Jeremchenko 2008 Speybroeck, Beukema, Bok & Voort 2016 Genus Lacerta (Eurasian Green Lizards)   Lacerta agilis Northern Eurasian Sand Lizard Bischoff in Boehme (ed.) 1984 Brelih & Dzukic 1974 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014   Lacerta bilineata Western Green Lizard Boehme, Fritz, Kotenko, Dzukic, Ljubisavljevic, Tzankov & Berendonk 2007 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Marzahn, Mayer, Joger, Ilgaz, Jablonski, Kindler, Kumlutas, Nistri, Schneewiess, Vamberger, Zagar & Fritz 2015 Mayer & Podnar 2002 Tvrtkovic, Lazar, Tome & Grbac 1998   Lacerta trilineata Western Balkan Green Lizard Arnold & Ovenden 2002 Brelih & Dzukic 1974 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Kornilios, Thanou, Lymberakis, Ilgaz, Kumlutas & Leache 2020 Mayer & Podnar 2002 Nettmann & Rykena in Boehme (ed.) 1984 Pawlowski & Kramer 2009 Speybroeck, Beukema, Bok & Voort 2016   Lacerta viridis Eastern Green Lizard Jelic 2014 Mayer & Podnar 2002 Nettmann & Rykena in Boehme (ed.) 1984 Pawlowski & Kramer 2009 Tvrtkovic, Lazar, Tome & Grbac 1998 Genus Podarcis (Wall Lizards)   Podarcis melisellensis Dalmatian Wall Lizard Arnold & Ovenden 2002 Brelih & Dzukic 1974 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Mayer & Podnar 2002 Pawlowski & Kramer 2009 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020 Speybroeck, Beukema, Bok & Voort 2016 Tiedemann & Henle in Boehme (ed.) 1986   Podarcis muralis Common European Wall Lizard Brelih & Dzukic 1974 Gruschwitz & Boehme in Boehme (ed.) 1986 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Mayer & Podnar 2002 Pawlowski & Kramer 2009 Speybroeck, Beukema, Bok & Voort 2016   Podarcis siculus Italian Wall Lizard Arnold & Ovenden 2002 Brelih & Dzukic 1974 Henle & Klaver in Boehme (ed.) 1986 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Mayer & Podnar 2002 Oskyrko, Sreelatha, Silva-Rocha, Vlad, Cogalniceanu, Stanescu, Iskenderov, Doronin, Lisicic & Carretero 2022 Pawlowski & Kramer 2009 Podnar, Mayer & Tvrtkovic 2005 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020 Sindaco & Jeremchenko 2008 Speybroeck, Beukema, Bok & Voort 2016 Genus Zootoca (Eurasian Forest Lizards)   Zootoca vivipara Viviparous Lizard Anonymous 2005 Brelih & Dzukic 1974 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014  Family Phyllodactylidae (Leaf-toed Geckos) Genus Tarentola (Wall Geckos)   Tarentola mauritanica Moorish Wall Gecko Arnold & Ovenden 2002 Brelih & Dzukic 1974 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Rieppel in Boehme (ed.) 1981 Speybroeck, Beukema, Bok & Voort 2016  Family Scincidae (Skinks) Genus Ablepharus (Eurasian Snake-eyed Skinks)   Ablepharus kitaibelii European Snake-eyed Skink Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Speybroeck, Beukema, Bok & Voort 2016 Szoevenyi & Jelic 2011

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Coronella (Palearctic Smooth Snakes)   Coronella austriaca Northern Smooth Snake Bonte 2012 Brelih & Dzukic 1974 Engelmann in Boehme (ed.) 1993 Geniez 2018 Jablonski, Jandzik & Gvozdik 2012 Jablonski, Nagy, Avci, Olgun, Kukushkin, Safaei-Mahroo & Jandzik 2019 Jelic 2014 Genus Dolichophis (Greater Whip Snakes)   Dolichophis caspius Caspian Whip Snake Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Krcmar, Mikuska & Kletecki 2007 Nagy, Bellaagh, Wink, Paunovic & Korsos 2010 Szczerbak & Boehme in Boehme (ed.) 1993 Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe quatuorlineata Western Four-lined Rat Snake Boehme & Szczerbak in Boehme (ed.) 1993 Brelih & Dzukic 1974 Geniez 2018 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Pawlowski & Kramer 2009 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020 Schulz 1996 Genus Hierophis (Eurasian Whip Snakes)   Hierophis carbonarius European Black Whip Snake Arnold & Ovenden 2002 [as viridiflavus] Brelih & Dzukic 1974 Geniez 2018 [as viridiflavus] Heimes in Boehme (ed.) 1993 Jablonski, Jandzik & Gvozdik 2012 [as viridiflavus] Jelic 2014 [as viridiflavus] Pawlowski & Kramer 2009 [as viridiflavus] Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016 Venchi & Sindaco 2007   Hierophis gemonensis Balkan Whip Snake Arnold & Ovenden 2002 Brelih & Dzukic 1974 Geniez 2018 Henle in Boehme (ed.) 1993 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Pawlowski & Kramer 2009 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020 Speybroeck, Beukema, Bok & Voort 2016 Genus Natrix (Eurasian Keelbacks)   Natrix natrix Common European Grass Snake Brelih & Dzukic 1974 Geniez 2018 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Kabisch in Boehme (ed.) 1999 Kindler, Boehme, Corti, Gvozdik, Jablonski, Jandzik, Metallinou, Siroky & Fritz 2013 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020   Natrix tessellata Dice Snake Brelih & Dzukic 1974 Geniez 2018 Gruschwitz, Lenz, Mebert & Lanka in Boehme (ed.) 1999 Jablonski, Jandzik & Gvozdik 2012 Jelic & Lelo 2011 Jelic 2014 Pawlowski & Kramer 2009 Genus Platyceps (Flat-headed Whip Snakes)   Platyceps najadum European Slender Whip Snake Arnold & Ovenden 2002 Brelih & Dzukic 1974 Darevsky & Szczerbak in Boehme (ed.) 1993 Geniez 2018 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Speybroeck, Beukema, Bok & Voort 2016 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus fallax European Cat Snake Brelih & Dzukic 1974 Geniez 2018 Grillitsch & Grillitsch in Boehme (ed.) 1999 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020 Speybroeck, Beukema, Bok & Voort 2016 Genus Zamenis (Western Palearctic Rat Snakes)   Zamenis longissimus Common Aesculapian Snake Boehme in Boehme (ed.) 1993 Brelih & Dzukic 1974 Geniez 2018 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Schulz 1996   Zamenis situla Leopard Snake Brelih & Dzukic 1974 Geniez 2018 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Obst, Szczerbak & Boehme in Boehme (ed.) 1993 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016  Family Lamprophiidae (House Snakes, etc.) Genus Malpolon (Montpellier Snakes)   Malpolon insignitus Eastern Montpellier Snake Brelih & Dzukic 1974 Geniez 2018 Haan in Boehme (ed.) 1999 Haan in Boehme 1999 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020  Family Typhlopidae (Typical Blind Snakes) Genus Xerotyphlops (Desert Blind Snakes)   Xerotyphlops vermicularis European Worm Snake Anonymous 2005 Arnold & Ovenden 2002 Grillitsch, Weish & Tiedemann 1999 Jelic 2014 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020 Sindaco, Venchi & Grieco 2013  Family Viperidae (Vipers) Genus Vipera (Typical Vipers or Adders)   Vipera ammodytes European Nose-horned Viper Brelih & Dzukic 1974 Geniez 2018 Heckes, Gruber & Stumpel in Joger & Stumpel 2005 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Jelic, Ajtic, Sterijovski, Crnobrnja-Isailovic, Lelo & Tomovic 2013 McDiarmid, Campbell & Toure 1999 Pawlowski & Kramer 2009 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020 Ursenbacher, Schweiger, Tomovic, Crnobrnja-Isailovic, Fumagalli & Mayer 2008   Vipera berus Northern Eurasian Viper Brelih & Dzukic 1974 David & Vogel 2010 Geniez 2018 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Jelic, Ajtic, Sterijovski, Crnobrnja-Isailovic, Lelo & Tomovic 2013 Nilson, Andren & Voelkl in Joger & Stumpel 2005 Pawlowski & Kramer 2009   Vipera ursinii Meadow and Karst Vipers Arnold & Ovenden 2002 Brelih & Dzukic 1974 David & Ineich 1999 David & Vogel 2010 Dely & Joger in Joger & Stumpel 2005 Geniez 2018 Gvozdik, Jandzik, Cordos, Rehak & Kotlik 2012 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Jelic, Ajtic, Sterijovski, Crnobrnja-Isailovic, Lelo & Tomovic 2013 Speybroeck, Beukema, Bok & Voort 2016

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Gramentz in Fritz (ed.) 2005 Jelic 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Gramentz & Fritz in Fritz (ed.) 2005 Jelic 2014  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Frazier, Gramentz & Fritz in Fritz (ed.) 2005 Jelic 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Emydidae (Western Pond Turtles) Genus Emys (European Pond Turtles)   Emys orbicularis Common European Pond Turtle Brelih & Dzukic 1974 Fritz in Fritz (ed.) 2001 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Pawlowski & Kramer 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sindaco & Jeremchenko 2008  Family Geoemydidae (Eastern Pond Turtles) Genus Mauremys (Stripe-necked Turtles)   Mauremys rivulata Balkan Pond Turtle Arnold & Ovenden 2002 Brelih & Dzukic 1974 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Mantziou & Rifai 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Speybroeck, Beukema, Bok & Voort 2016 Wischuf & Busack in Fritz (ed.) 2001  Family Testudinidae (Tortoises) Genus Testudo (Palearctic Tortoises)   Testudo hermanni Greek Tortoise Bertolero, Cheylan, Hailey, Livoreil & Willemsen 2011 Bonin, Devaux & Dupre 2006 Brelih & Dzukic 1974 Cheylan in Fritz (ed.) 2001 Jablonski, Jandzik & Gvozdik 2012 Jelic 2014 Ljubisavljevic, Dzukic, Vukov & Kalezic 2014 Pawlowski & Kramer 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Schmidt, Kranzelic, Majetic, Laus, Stih & Koren 2020 Speybroeck, Beukema, Bok & Voort 2016