Reptiles of Italy  Related checklists:  Italian mainland - Aegadian Islands - Aeolian Islands - Capraia - Capri - Elba - Montecristo - Pantelleria - Pelagie Islands - Pontine Islands - Sardinia - Sicily Bibliography of the reptiles of Italy Taxonomic history of the reptiles of Italy Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Anguidae (Slowworms, etc.) Genus Anguis (Slowworms)   Anguis fragilis Western European Slowworm Corsetti & Capula 1992 Gvozdik, Benkovsky, Crottini, Bellati, Moravec, Romano, Sacchi & Jandzik 2013 Gvozdik, Jandzik, Lymberakis, Jablonski & Moravec 2010 Sindaco, Doria, Razzetti & Bernini (eds.) 2006   Anguis veronensis Italian Slowworm Gvozdik, Benkovsky, Crottini, Bellati, Moravec, Romano, Sacchi & Jandzik 2013 Mikulicek, Jablonski, Palenik, Gvozdik & Jandzik 2018 Sindaco & Razzetti 2021 Sindaco, Venchi & Grieco 2013 [as Anguis sp.]  Family Gekkonidae (Typical Geckos) Genus Hemidactylus (Half-toed Geckos)   Hemidactylus turcicus Turkish Half-toed Gecko Salvador in Boehme (ed.) 1981 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016 Genus Mediodactylus (Mediterranean Bent-toed Geckos)   Mediodactylus kotschyi Balkan Bent-toed Gecko Beutler in Boehme (ed.) 1981 Kotsakiozi, Jablonski, Ilgaz, Kumlutas, Avci, Meiri, Itescu, Kukushkin, Gvozdik, Scillitani, Roussos, Jandzik, Kasapidis, Lymberakis & Poulakakis 2018 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016  Family Lacertidae (Typical Lizards) Genus Algyroides (Keeled Lizards)   Algyroides fitzingeri Pygmy Keeled Lizard Corti, Biaggini, Nulchis, Cogoni, Cossu, Frau, Mulargia, Lunghi & Bassu 2022 Pous, Speybroeck, Bogaerts, Pasmans & Beukema 2013 Salvi & Bombi 2010 Schneider in Boehme (ed.) 1981 Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016   Algyroides nigropunctatus Blue-throated Keeled Lizard Bischoff in Boehme (ed.) 1981 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016 Genus Archaeolacerta (Tyrrhenian Mountain Lizard)   Archaeolacerta bedriagae Tyrrhenian Mountain Lizard Corti, Biaggini, Nulchis, Cogoni, Cossu, Frau, Mulargia, Lunghi & Bassu 2022 Pous, Speybroeck, Bogaerts, Pasmans & Beukema 2013 Salvi & Bombi 2010 Schneider in Boehme (ed.) 1984 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016 Genus Iberolacerta (Southern European Rock Lizards)   Iberolacerta horvathi Croatian Rock Lizard Bischoff in Boehme (ed.) 1984 Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016 Genus Lacerta (Eurasian Green Lizards)   Lacerta agilis Northern Eurasian Sand Lizard Necas, Modry & Zavadil 1997 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006   Lacerta bilineata Western Green Lizard Amann, Razetti & Joger 2001 Amann, Rykena, Joger, Nettmann & Veith 1997 Boehme, Fritz, Kotenko, Dzukic, Ljubisavljevic, Tzankov & Berendonk 2007 Marzahn, Mayer, Joger, Ilgaz, Jablonski, Kindler, Kumlutas, Nistri, Schneewiess, Vamberger, Zagar & Fritz 2015 Nettmann & Rykena in Boehme (ed.) 1984 [as L. viridis] Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016   Lacerta viridis Eastern Green Lizard Amann, Razetti & Joger 2001 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Genus Podarcis (Wall Lizards)   Podarcis filfolensis Maltese Wall Lizard Bischoff in Boehme (ed.) 1986 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016   * Podarcis latastei Western Pontine Islands Wall Lizard Bedriaga 1879 Corti & Cascio 2002 Senczuk, Castiglia, Boehme & Corti 2019   Podarcis melisellensis Dalmatian Wall Lizard Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016 Tiedemann & Henle in Boehme (ed.) 1986   Podarcis muralis Common European Wall Lizard Gruschwitz & Boehme in Boehme (ed.) 1986 Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016   * Podarcis raffoneae Aeolian Wall Lizard Palma 1980 [as P. sicula cucchiarai] Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016   Podarcis siculus Italian Wall Lizard Corti, Biaggini, Nulchis, Cogoni, Cossu, Frau, Mulargia, Lunghi & Bassu 2022 Henle & Klaver in Boehme (ed.) 1986 Podnar, Mayer & Tvrtkovic 2005 Pous, Speybroeck, Bogaerts, Pasmans & Beukema 2013 Salvi & Bombi 2010 Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016   Podarcis tiliguerta Tyrrhenian Wall Lizard Corti, Biaggini, Nulchis, Cogoni, Cossu, Frau, Mulargia, Lunghi & Bassu 2022 Pous, Speybroeck, Bogaerts, Pasmans & Beukema 2013 Salvi & Bombi 2010 Schneider in Boehme (ed.) 1986 Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016   * Podarcis waglerianus Sicilian Wall Lizard Boehme in Boehme (ed.) 1986 Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016 Genus Timon (Ocellated Lizards)   Timon lepidus Northern Ocellated Lizard Bischoff 1996 Honegger 1981 Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016 Genus Zootoca (Eurasian Forest Lizards)   Zootoca carniolica Carniolian Lizard Ghielmi, Giovine, Menegon, Lapini, Surget T-Groba & Heulin 2006 Mayer, Boehme, Tiedemann & Bischoff 2000 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006   Zootoca vivipara Viviparous Lizard Dely & Boehme in Boehme (ed.) 1984 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016 Surget-Groba, Heulin, Guillaume, Thorpe, Kupriyanova, Vogrin, Maslak, Mazzotti, Venczel, Ghira, Odierna, Leontyeva, Monney & Smith 2001  Family Phyllodactylidae (Leaf-toed Geckos) Genus Tarentola (Wall Geckos)   Tarentola fascicularis Libyan Wall Gecko Harris, Carretero, Corti & Lo-Cascio 2009 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 [as mauritanica] Speybroeck, Beukema, Bok & Voort 2016 [as mauritanica]   Tarentola mauritanica Moorish Wall Gecko Rieppel in Boehme (ed.) 1981 Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016  Family Scincidae (Skinks) Genus Chalcides (Cylindrical Skinks)   Chalcides chalcides Italian Three-toed Cylindrical Skink Busack 1986 Caputo, Lanza & Palmieri 1995 Carranza, Arnold, Geniez, Roca & Mateo 2008 Orsini & Cheylan in Boehme (ed.) 1981 Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016   Chalcides ocellatus Mediterranean Ocellated Skink Pasteur 1981 Schneider in Boehme (ed.) 1981 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016   Chalcides striatus Western Three-toed Cylindrical Skink Caputo & Odierna 1992 Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006  Family Sphaerodactylidae (Least Geckos, etc.) Genus Euleptes (Mediterranean Leaf-toed Gecko)   Euleptes europaea Mediterranean Leaf-toed Gecko Bauer, Good & Branch 1997 Corti, Biaggini, Nulchis, Cogoni, Cossu, Frau, Mulargia, Lunghi & Bassu 2022 Delaugerre, Ouni & Nouira 2011 Pous, Speybroeck, Bogaerts, Pasmans & Beukema 2013 Rieppel & Schneider 1981 Rieppel & Schneider in Boehme (ed.) 1981 Salvi & Bombi 2010 Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx jaculus Western Sand Boa Insacco, Spadola, Russotto & Scaravelli 2015 Sindaco & Razzetti 2021 Speybroeck, Beukema, Bok & Voort 2016  Family Colubridae (Typical Snakes or Colubrids) Genus Coronella (Palearctic Smooth Snakes)   Coronella austriaca Northern Smooth Snake Engelmann in Boehme (ed.) 1993 Geniez 2018 Jablonski, Nagy, Avci, Olgun, Kukushkin, Safaei-Mahroo & Jandzik 2019 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini [eds.] 2006 Speybroeck, Beukema, Bok & Voort 2016   Coronella girondica Southern Smooth Snake Dusej in Boehme (ed.) 1993 Geniez 2018 Iversen, Fatis, Romano, Pedrini & Nicola 2020 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini [eds.] 2006 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016 Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe quatuorlineata Western Four-lined Rat Snake Boehme & Szczerbak in Boehme (ed.) 1993 Geniez 2018 Schulz 1996 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini [eds.] 2006 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016 Genus Hierophis (Eurasian Whip Snakes)   Hierophis carbonarius European Black Whip Snake Avella, Castiglia & Senczuk 2017 Heimes in Boehme (ed.) 1993 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 [as viridiflavus] Sindaco, Doria, Razzetti & Bernini [eds.] 2006 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016   Hierophis viridiflavus Western Whip Snake Avella, Castiglia & Senczuk 2017 Corti, Biaggini, Nulchis, Cogoni, Cossu, Frau, Mulargia, Lunghi & Bassu 2022 Geniez 2018 Heimes in Boehme (ed.) 1993 Pous, Speybroeck, Bogaerts, Pasmans & Beukema 2013 Salvi & Bombi 2010 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Doria, Razzetti & Bernini [eds.] 2006 Speybroeck, Beukema, Bok & Voort 2016 Genus Macroprotodon (False Smooth Snakes)   Macroprotodon cucullatus Eastern False Smooth Snake Busack & McCoy 1990 [as mauritanicus] Joger in Boehme (ed.) 1999 Sindaco & Razzetti 2021 [as "M. cf. cucullatus"] Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013 Venchi & Sindaco 2007 Wade 1988 Wade 2001 Genus Natrix (Eurasian Keelbacks)   Natrix helvetica Barred Grass Snake Arnold & Ovenden 2002 Corti, Biaggini, Nulchis, Cogoni, Cossu, Frau, Mulargia, Lunghi & Bassu 2022 Geniez 2018 Gruber 1989 Kabisch in Boehme (ed.) 1999 Kindler, Boehme, Corti, Gvozdik, Jablonski, Jandzik, Metallinou, Siroky & Fritz 2013 Pous, Speybroeck, Bogaerts, Pasmans & Beukema 2013 Salvi & Bombi 2010 [as natrix] Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016   Natrix maura Viperine Grass Snake Geniez 2018 Gruber 1989 Schatti in Boehme (ed.) 1999 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016   Natrix natrix Common European Grass Snake Arnold & Ovenden 2002 Geniez 2018 [as N. natrix [part]] Kabisch in Boehme (ed.) 1999 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013   Natrix tessellata Dice Snake Geniez 2018 Gruber 1989 Gruschwitz, Lenz, Mebert & Lanka in Boehme (ed.) 1999 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus fallax European Cat Snake Geniez 2018 Grillitsch & Grillitsch in Boehme (ed.) 1999 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016 Venchi & Sindaco 2007 Genus Zamenis (Western Palearctic Rat Snakes)   * Zamenis lineatus South Italian Aesculapian Snake Boehme in Boehme (ed.) 1993 [as Elaphe longissima romana] Esposito & Romano 2011 Geniez 2018 Schulz 1996 [as romana] Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016 Venchi & Sindaco 2007   Zamenis longissimus Common Aesculapian Snake Boehme in Boehme (ed.) 1993 Geniez 2018 Schulz 1996 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016 Vaccaro & Turrisi 2007   Zamenis situla Leopard Snake Geniez 2018 Obst, Szczerbak & Boehme in Boehme (ed.) 1993 Schulz 1996 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016 Venchi & Sindaco 2007  Family Lamprophiidae (House Snakes, etc.) Genus Malpolon (Montpellier Snakes)   Malpolon insignitus Eastern Montpellier Snake Faraone, Melfi, Nicola, Giacalone & Valvo 2022 Haan in Boehme (ed.) 1999 Kramer & Schnurrenberger 1963 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016 Venchi & Sindaco 2007   Malpolon monspessulanus Western Montpellier Snake Geniez 2018 Haan in Boehme (ed.) 1999 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013 Sindaco, Venchi, Carpaneto & Bologna 2000  Family Viperidae (Vipers) Genus Vipera (Typical Vipers or Adders)   Vipera ammodytes European Nose-horned Viper David & Vogel 2010 Geniez 2018 Gruber 1989 Heckes, Gruber & Stumpel in Joger & Stumpel 2005 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016 Ursenbacher, Schweiger, Tomovic, Crnobrnja-Isailovic, Fumagalli & Mayer 2008   Vipera aspis Asp Viper David & Vogel 2010 Geniez 2018 Gruber 1989 McDiarmid, Campbell & Toure 1999 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016 Trutnau, Boehme & Joger in Joger & Stumpel 2005 Venchi & Sindaco 2007   Vipera berus Northern Eurasian Viper David & Vogel 2010 Geniez 2018 Honegger 1981 Nilson, Andren & Voelkl in Joger & Stumpel 2005 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006   Vipera ursinii Meadow and Karst Vipers David & Ineich 1999 David & Vogel 2010 Dely & Joger in Joger & Stumpel 2005 Geniez 2018 Gruber 1989 Gvozdik, Jandzik, Cordos, Rehak & Kotlik 2012 Nilson & Andren 2001 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016   * Vipera walser Northwest Italian Viper Ghielmi, Menegon, Marsden, Laddaga & Ursenbacher 2016 Sindaco & Razzetti 2021 Speybroeck, Beukema, Bok & Voort 2016

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Blasi, Hochscheid, Bardelli, Bruno, Melodia, Salzeri, Rosa & Madonia 2022 Cambie 2011 Casale, Palilla, Salemi, Napoli, Prinzi, Genco, Bonaviri, Mastrogiacomo, Oliveiro & Valvo 2012 Casale, Simone, Conoscitore, Conoscitore & Salvemini 2012 Freggi, Fornari, Lo Conte & Longo 2003 King & Burke 1989 Prazzi, Nicolini, Piovano & Giacoma 2010 Pritchard 1979 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Sindaco, Venchi, Carpaneto & Bologna 2000 Speybroeck, Beukema, Bok & Voort 2016 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Freggi, Fornari, Lo Conte & Longo 2003 Gramentz & Fritz in Fritz (ed.) 2005 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys kempii Atlantic Ridley Sea Turtle Insacco & Spadola 2010 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sindaco & Razzetti 2021  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Frazier, Gramentz & Fritz in Fritz (ed.) 2005 Honegger 1981 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006  Family Emydidae (Western Pond Turtles) Genus Emys (European Pond Turtles)   Emys orbicularis Common European Pond Turtle Fritz in Fritz (ed.) 2001 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sindaco & Jeremchenko 2008 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 [as "cf. orbicularis hellenica"] Speybroeck, Beukema, Bok & Voort 2016   * Emys trinacris Sicilian Pond Turtle Fritz & Havas 2006 [as orbicularis] Fritz, Fattizzo, Guicking, Tripepi, Pennisi, Lenk, Joger & Wink 2005 Ottonello, D'Angelo, Marrone, Oneto, Spadola, Zuffi & Fritz 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016  Family Testudinidae (Tortoises) Genus Testudo (Palearctic Tortoises)   Testudo hermanni Greek Tortoise Bertolero, Cheylan, Hailey, Livoreil & Willemsen 2011 Cheylan in Fritz (ed.) 2001 Fritz & Havas 2006 Honegger 1981 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sindaco & Razzetti 2021 Sindaco, Doria, Razzetti & Bernini (eds.) 2006 Speybroeck, Beukema, Bok & Voort 2016