Reptiles of Greek mainland  (excl. Peloponnese and all Greek islands) Related checklists:  Greece (incl. Peloponnese and all Greek islands) Bibliography of the reptiles of Greek mainland Taxonomic history of the reptiles of Greek mainland Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Anguidae (Slowworms, etc.) Genus Anguis (Slowworms)   Anguis cephallonica Peloponnese Slowworm Sindaco & Jeremchenko 2008   Anguis fragilis Western European Slowworm Gvozdik, Benkovsky, Crottini, Bellati, Moravec, Romano, Sacchi & Jandzik 2013 Gvozdik, Jandzik, Lymberakis, Jablonski & Moravec 2010   Anguis graeca Greek Slowworm Chondropoulos 1986 [as fragilis colchicus] Gvozdik, Benkovsky, Crottini, Bellati, Moravec, Romano, Sacchi & Jandzik 2013 Gvozdik, Jandzik, Lymberakis, Jablonski & Moravec 2010 Thanou, Giokas & Kornilios 2014 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 [as fragilis colchicus] Genus Pseudopus (Eurasian Glass Lizard)   Pseudopus apodus Eurasian Glass Lizard Chondropoulos 1986 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008  Family Gekkonidae (Typical Geckos) Genus Hemidactylus (Half-toed Geckos)   Hemidactylus turcicus Turkish Half-toed Gecko Chondropoulos 1986 Salvador in Boehme (ed.) 1981 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Mediodactylus (Mediterranean Bent-toed Geckos)   Mediodactylus danilewskii Northern Bent-toed Gecko Chondropoulos 1986   Mediodactylus kotschyi Balkan Bent-toed Gecko Beutler in Boehme (ed.) 1981 Chondropoulos 1986 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008  Family Lacertidae (Typical Lizards) Genus Algyroides (Keeled Lizards)   Algyroides moreoticus Greek Keeled Lizard Sindaco & Jeremchenko 2008   Algyroides nigropunctatus Blue-throated Keeled Lizard Andriopoulos & Pafilis 2016 Bischoff in Boehme (ed.) 1981 Chondropoulos 1986 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Darevskia (Caucasian Rock Lizards)   Darevskia hungarica Balkan Meadow Lizard Sindaco & Jeremchenko 2008 [as praticola] Sos, Kecskes, Hegyeli & Marosi 2012 [as pontica] Speybroeck, Beukema, Bok & Voort 2016 [as praticola] Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 [as pontica] Genus Lacerta (Eurasian Green Lizards)   Lacerta agilis Northern Eurasian Sand Lizard Chondropoulos 1986 Sagonas, Poulakakis, Lymberakis, Parmakelis, Pafilis & Valakos 2014 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Lacerta bilineata Western Green Lizard Boehme, Fritz, Kotenko, Dzukic, Ljubisavljevic, Tzankov & Berendonk 2007 Sagonas, Poulakakis, Lymberakis, Parmakelis, Pafilis & Valakos 2014 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Lacerta diplochondrodes Eastern Balkan Green Lizard Kornilios, Thanou, Lymberakis, Ilgaz, Kumlutas & Leache 2020 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Lacerta trilineata Western Balkan Green Lizard Chondropoulos 1986 Kornilios, Thanou, Lymberakis, Ilgaz, Kumlutas & Leache 2020 Nettmann & Rykena in Boehme (ed.) 1984 Sagonas, Poulakakis, Lymberakis, Parmakelis, Pafilis & Valakos 2014 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Lacerta viridis Eastern Green Lizard Boehme, Fritz, Kotenko, Dzukic, Ljubisavljevic, Tzankov & Berendonk 2007 Chondropoulos 1986 Marzahn, Mayer, Joger, Ilgaz, Jablonski, Kindler, Kumlutas, Nistri, Schneewiess, Vamberger, Zagar & Fritz 2015 Nettmann & Rykena in Boehme (ed.) 1984 Sagonas, Poulakakis, Lymberakis, Parmakelis, Pafilis & Valakos 2014 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Ophisops (Snake-eyed Lizards)   Ophisops elegans Elegant Snake-eyed Lizard Chondropoulos 1986 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Podarcis (Wall Lizards)   Podarcis erhardii Aegean Wall Lizard Chondropoulos 1986 Gruber in Boehme (ed.) 1986 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Podarcis ionicus Ionian Wall Lizard Chondropoulos 1986 Kabisch in Boehme (ed.) 1986 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Podarcis muralis Common European Wall Lizard Chondropoulos 1986 Gruschwitz & Boehme in Boehme (ed.) 1986 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Podarcis peloponnesiacus Peloponnese Wall Lizard Hedman, Kapsalas, Karameta, Psonis, Poulakakis, Foufopoulos & Pafilis 2017   Podarcis tauricus Balkan Wall Lizard Chondropoulos 1986 Kabisch in Boehme (ed.) 1986 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Zootoca (Eurasian Forest Lizards)   Zootoca vivipara Viviparous Lizard Strachinis, Karagianni, Stanchev & Stanchev 2019  Family Scincidae (Skinks) Genus Ablepharus (Eurasian Snake-eyed Skinks)   Ablepharus kitaibelii European Snake-eyed Skink Chondropoulos 1986 Skourtanioti, Kapli, Ilgaz, Kumlutas, Avci, Ahmadzadeh, Crnobrnja-Isailovic, Ghergel, Lymberakis & Poulakakis 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Chalcides (Cylindrical Skinks)   Chalcides ocellatus Mediterranean Ocellated Skink Chondropoulos 1986 Schneider in Boehme (ed.) 1981 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Ophiomorus (South West Asian Snake Skinks)   Ophiomorus punctatissimus Greek Snake Skink Bischoff in Boehme (ed.) 1981 Chondropoulos 1986 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx jaculus Western Sand Boa Chondropoulos 1989 Speybroeck, Beukema, Bok & Voort 2016 Tokar & Obst in Boehme (ed.) 1993 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008  Family Colubridae (Typical Snakes or Colubrids) Genus Coronella (Palearctic Smooth Snakes)   Coronella austriaca Northern Smooth Snake Chondropoulos 1989 Engelmann in Boehme (ed.) 1993 Jablonski, Nagy, Avci, Olgun, Kukushkin, Safaei-Mahroo & Jandzik 2019 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Dolichophis (Greater Whip Snakes)   Dolichophis caspius Caspian Whip Snake Chondropoulos 1989 Nagy, Bellaagh, Wink, Paunovic & Korsos 2010 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Eirenis (Middle Eastern Dwarf Snakes)   Eirenis modestus Ring-headed Dwarf Snake Mahlow, Tillack, Schmidtler & Muller 2013 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe quatuorlineata Western Four-lined Rat Snake Boehme & Szczerbak in Boehme (ed.) 1993 Chondropoulos 1989 Schulz 1996 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Elaphe sauromates Sarmatian Rat Snake Boehme & Szczerbak in Boehme (ed.) 1993 Chondropoulos 1989 Schulz 1996 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Hierophis (Eurasian Whip Snakes)   Hierophis gemonensis Balkan Whip Snake Chondropoulos 1989 Henle in Boehme (ed.) 1993 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Natrix (Eurasian Keelbacks)   Natrix natrix Common European Grass Snake Chondropoulos 1989 Kabisch in Boehme (ed.) 1999 Kindler, Boehme, Corti, Gvozdik, Jablonski, Jandzik, Metallinou, Siroky & Fritz 2013 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Natrix tessellata Dice Snake Chondropoulos 1989 Gruschwitz, Lenz, Mebert & Lanka in Boehme (ed.) 1999 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Platyceps (Flat-headed Whip Snakes)   Platyceps najadum European Slender Whip Snake Chondropoulos 1989 Darevsky & Szczerbak in Boehme (ed.) 1993 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus fallax European Cat Snake Chondropoulos 1989 Grillitsch & Grillitsch in Boehme (ed.) 1999 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Zamenis (Western Palearctic Rat Snakes)   Zamenis longissimus Common Aesculapian Snake Boehme in Boehme (ed.) 1993 Chondropoulos 1989 Nilson & Andren 1984 Schulz 1996 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Zamenis situla Leopard Snake Chondropoulos 1989 Obst, Szczerbak & Boehme in Boehme (ed.) 1993 Schulz 1996 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008  Family Lamprophiidae (House Snakes, etc.) Genus Malpolon (Montpellier Snakes)   Malpolon insignitus Eastern Montpellier Snake Carranza, Arnold & Pleguezuelos 2006 Chondropoulos 1989 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008  Family Typhlopidae (Typical Blind Snakes) Genus Xerotyphlops (Desert Blind Snakes)   Xerotyphlops vermicularis European Worm Snake Chondropoulos 1989 Grillitsch & Grillitsch in Boehme (ed.) 1993 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008  Family Viperidae (Vipers) Genus Montivipera (Middle East Mountain Vipers)   Montivipera xanthina Ottoman Viper Cattaneo 2017 Chondropoulos 1989 David & Vogel 2010 Joger & Nilson in Joger & Stumpel 2005 McDiarmid, Campbell & Toure 1999 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Genus Vipera (Typical Vipers or Adders)   Vipera ammodytes European Nose-horned Viper Chondropoulos 1989 David & Vogel 2010 Heckes, Gruber & Stumpel in Joger & Stumpel 2005 McDiarmid, Campbell & Toure 1999 Speybroeck, Beukema, Bok & Voort 2016 Ursenbacher, Schweiger, Tomovic, Crnobrnja-Isailovic, Fumagalli & Mayer 2008 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Vipera berus Northern Eurasian Viper Chondropoulos 1989 David & Vogel 2010 Nilson, Andren & Voelkl in Joger & Stumpel 2005 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Vipera graeca Greek Meadow Viper Chondropoulos 1989 David & Vogel 2010 Dely & Joger in Joger & Stumpel 2005 Nilson & Andren 1988 Nilson & Andren 2001 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Gramentz in Fritz (ed.) 2005 Speybroeck, Beukema, Bok & Voort 2016  Family Emydidae (Western Pond Turtles) Genus Emys (European Pond Turtles)   Emys orbicularis Common European Pond Turtle Fritz in Fritz (ed.) 2001 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008  Family Geoemydidae (Eastern Pond Turtles) Genus Mauremys (Stripe-necked Turtles)   Mauremys rivulata Balkan Pond Turtle Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008 Wischuf & Busack in Fritz (ed.) 2001  Family Testudinidae (Tortoises) Genus Testudo (Palearctic Tortoises)   Testudo graeca Spur-thighed Tortoise Buskirk, Keller & Andreu in Fritz (ed.) 2001 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Testudo hermanni Greek Tortoise Cheylan in Fritz (ed.) 2001 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008   Testudo marginata Marginated Tortoise Bringsøe, Buskirk & Willemsen in Fritz (ed.) 2001 Speybroeck, Beukema, Bok & Voort 2016 Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou & Foufopoulos 2008