Turtles of China (incl. Tibet)   Related checklists:  China (main menu) - Anhui - Beijing - Chongqing - Fujian - Gansu - Guangdong - Guangxi - Guizhou - Hainan - Hebei - Heilongjiang - Henan - Hong Kong - Hubei - Hunan - Inner Mongolia (Neimenggu) - Jiangsu - Jiangxi - Jilin - Liaoning - Macau - Ningxia - Qinghai - Shaanxi - Shandong - Shanghai - Shanxi - Sichuan - Tianjin - Tibet (Xizang) - Xinjiang - Xisha Islands - Yunnan - Zhejiang Bibliography of the reptiles of China

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Chen, Zhang & Xu 2009 Huang in Bjorndal (ed.) 1982 Lu, Wang & Li 1999 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Wen 1984 Zhao & Adler 1993 Zheng 1985 Zhou & Chen 2002 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Chen, Zhang & Xu 2009 Huang in Bjorndal (ed.) 1982 Jia, Wang, Balazs & Liu 2019 Karsen, Lau & Bogadek 1998 King & Burke 1989 Lu, Wang & Li 1999 Ng & Wan 2011 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Wen 1984 Zhao & Adler 1993 Zheng 1985 Zhou & Chen 2002 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Chen, Zhang & Xu 2009 Huang in Bjorndal (ed.) 1982 Lu, Wang & Li 1999 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Wen 1984 Zhao & Adler 1993 Zhao 2005 Zheng 1985 Zhou & Chen 2002 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Chen, Zhang & Xu 2009 Karsen, Lau & Bogadek 1998 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Wen 1984 Zhao & Adler 1993 Zheng 1985 Zhou & Chen 2002  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Chen, Zhang & Xu 2009 Huang in Bjorndal (ed.) 1982 Karsen, Lau & Bogadek 1998 King & Burke 1989 Lu, Wang & Li 1999 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhao & Adler 1993 Zheng 1985 Zhou & Chen 2002  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   * Cuora aurocapitata Yellow-headed Box Turtle Blanck, Ting, Protiva, Crow & Tiedemann 2017 Fritz & Havas 2006 Iverson 1992 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhang & Chen 2002 Zhao & Adler 1993   Cuora cyclornata Vietnamese Three-striped Box Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Cuora flavomarginata Chinese Yellow-margined Box Turtle Chen, Zhang & Xu 2009 Easton & Leung-Va 1993 Iverson 1992 Karsen, Lau & Bogadek 1998 Lau & Shi 2000 Lau, Chan, Crow & Ades 2000 Luo, Gao & Zhou 2004 McCord & Joseph-Ouni 2002 Ota, Yasukawa, Fu & Chen 2009 Qu, Lu, Chen, Niu, Lu & Niu 2002 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhang & Chen 2002 Zhao & Adler 1993 Zhou & Chen 2002   Cuora galbinifrons Northern Indochinese Box Turtle Iverson 1992 Lau & Shi 2000 McCord & Joseph-Ouni 2002 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhao & Adler 1993 Zhao 2005   * Cuora mccordi Guangxi Box Turtle Bonin, Devaux & Dupre 2006 Iverson 1992 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhao & Adler 1993   Cuora mouhotii Keeled Box Turtle Das 1987 Das, McCormack, Dijk, Hoang & Struijk 2016 He & Zhou 2002 Iverson 1992 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhao & Adler 1993 Zhao 2005   * Cuora pani Shaanxi Box Turtle Fritz & Havas 2006 Huang & Huang 2011 Iverson 1992 Lau & Shi 2000 Li 2000 [reported from "Sichuan", but may refer to Chongqing] Luo, Liu, Gao, Luo, Huang, Xiao, Wang, Zhou, Luo & Tang 2012 McCord & Joseph-Ouni 2002 Parham & Li 1999 [reported from "Sichuan", but may refer to Chongqing] Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Song 2002 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhao & Adler 1993   * Cuora trifasciata Northern Three-striped Box Turtle Blanck, Ting, Protiva, Crow & Tiedemann 2017 Chen, Zhang & Xu 2009 Easton & Leung-Va 1993 Fritz & Havas 2006 Iverson 1992 Karsen, Lau & Bogadek 1998 Lau & Shi 2000 Lau, Chan, Crow & Ades 2000 Li, Xiao, Qing, Lu & Lazell 2009 Nguyen, Ho & Nguyen 2009 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Xu 2007 Zhao & Adler 1993 Zhao 2005   * Cuora yunnanensis Yunnan Box Turtle Fritz & Havas 2006 He & Zhou 2002 Iverson 1992 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhao & Adler 1993   Cuora zhoui Southern Chinese Box Turtle He & Zhou 2002 Iverson 1992 Lau & Shi 2000 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhao & Adler 1993 Genus Geoemyda (Black-breasted Leaf Turtles)   Geoemyda spengleri Common Black-breasted Leaf Turtle Dawson, Gaillard, Gong, Lin, McCormack, Nanthavong, Nguyen, Nguyen, Pham & Shi 2022 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Xu 2007 Yasukawa & Ota 2010 Zhao & Adler 1993 Genus Mauremys (Stripe-necked Turtles)   Mauremys mutica Yellow Pond Turtle Chen, Zhang & Xu 2009 Iverson 1992 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Xu 2007 Zhang & Chen 2002 Zhao & Adler 1993 Zhao 2005 Zhou & Chen 2002   Mauremys nigricans Guangdong River Turtle Iverson 1992 Lau & Shi 2000 Li, Xiao, Qing, Lu & Lazell 2009 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Xu 2007 Zhao & Adler 1993   Mauremys reevesii Chinese Three-keeled Pond Turtle Chen, Zhang & Xu 2009 Easton & Leung-Va 1993 Iverson 1992 Karsen, Lau & Bogadek 1998 Lau & Shi 2000 [as megalocephala] Lazell 2002 Lovich, Yasukawa & Ota 2011 Lu, Wang & Li 1999 Luo, Gao & Zhou 2004 Luo, Gao, Wei, Yan, Peng, Zeng, Zhao & Chen 2003 Luo, Liu, Gao, Luo, Huang, Xiao, Wang, Zhou, Luo & Tang 2012 McCord & Joseph-Ouni 2004 Ng 2010 [as megalocephala] Qu, Lu, Chen, Niu, Lu & Niu 2002 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Song 2002 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Xu 2007 Yao 2004 Zhang & Chen 2002 [as megalocephala] Zhao & Adler 1993 Zhong 2004 Zhou & Chen 2002 [as megalocephala]   Mauremys sinensis Chinese Stripe-necked Turtle Anonymous 2013 Chen, Zhang & Xu 2009 Iverson 1992 Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Xu 2007 Zhao & Adler 1993 Zhao 2005 Zhou & Chen 2002 Genus Sacalia (Eastern Eyed Turtles)   * Sacalia bealei Eastern Chinese Eyed Turtle Chen, Zhang & Xu 2009 Iverson 1992 Karsen, Lau & Bogadek 1998 Lau & Shi 2000 Lau, Chan, Crow & Ades 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Xu 2007 Zhao & Adler 1993 Zhong 2004   Sacalia quadriocellata Four-eyed Turtle Chen, Zhang & Xu 2009 Iverson 1992 Lau & Shi 2000 McCord & Joseph-Ouni 2004 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Xu 2007 Zhao & Adler 1993 Zhao 2005  Family Platysternidae (Big-headed Turtle) Genus Platysternon (Big-headed Turtle)   Platysternon megacephalum Big-headed Turtle Chen & Li 1994 [reported from "Sichuan", but may refer to Chongqing] Chen, Zhang & Xu 2009 He & Zhou 2002 Iverson 1992 Karsen, Lau & Bogadek 1998 Lau & Shi 2000 Lau, Chan, Crow & Ades 2000 Li, Xiao, Qing, Lu & Lazell 2009 Luo, Gao & Zhou 2004 Luo, Liu, Gao, Luo, Huang, Xiao, Wang, Zhou, Luo & Tang 2012 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Xu 2007 Zhang & Chen 2002 Zhao & Adler 1993 Zhao 2005 Zhong 2004 Zhou & Chen 2002  Family Testudinidae (Tortoises) Genus Indotestudo (Asian Tortoises)   Indotestudo elongata Elongated Tortoise Ihlow, Dawson, Hartmann & Som 2016 Iverson 1992 Lau & Shi 2000 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhao & Adler 1993 Genus Manouria (Indochinese Tortoises)   Manouria impressa Impressed Tortoise He & Zhou 2002 Iverson 1992 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2017 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhao & Adler 1993 Genus Testudo (Palearctic Tortoises)   Testudo horsfieldii Steppe Tortoise Fritz & Havas 2006 Iverson 1992 Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Shi, Zhou & Yuan 2002 Sindaco & Jeremchenko 2008 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhao & Adler 1993  Family Trionychidae (Softshell Turtles) Genus Nilssonia (South Asian Softshell Turtles)   Nilssonia formosa Burmese Peacock Softshell Turtle Liebing, Praschag, Gosh, Vasudevan, Rashid, Rao, Stuckas & Fritz 2012 Genus Palea (Wattle-necked Softshell Turtle)   Palea steindachneri Wattle-necked Softshell Turtle He & Zhou 2002 Iverson 1992 Lau & Shi 2000 Parham & Wang 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Xu 2007 Zhao & Adler 1993 Genus Pelochelys (Indomalayan Giant Softshell Turtles)   Pelochelys cantorii Asian Giant Softshell Turtle Bonin, Devaux & Dupre 2006 Chen, Zhang & Xu 2009 [as bibroni] Das 2008 Fritz & Havas 2006 Iverson 1992 [as bibroni] Lau & Shi 2000 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Xu 2007 Zhao & Adler 1993 [as bibroni] Zhao 2005 Genus Pelodiscus (East Asian Softshell Turtles)   * Pelodiscus axenaria Hunan Softshell Turtle Fritz & Havas 2006 Gong, Fritz, Vamberger, Gao & Farkas 2022 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Yang, Tang, Ding, Guo & Wang 2011   * Pelodiscus huangshanensis Huangshan Softshell Turtle Gong, Peng, Huang, Lin, Huang, Xu, Yang & Nie 2021 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021   Pelodiscus maackii Amur Softshell Turtle Chen, Liu & Zhao 2007 [as sinensis] Fritz & Havas 2006 Jiang 2002 [as sinensis] Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Schneeweiss & Mix 2011 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhao & Adler 1993 [as sinensis]   Pelodiscus parviformis Lesser Chinese Softshell Turtle Fritz & Havas 2006 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020   * Pelodiscus shipian Chinese Stone Slab Soft-shelled Turtle Gong, Fritz, Vamberger, Gao & Farkas 2022   Pelodiscus sinensis Chinese Softshell Turtle Chen, Zhang & Xu 2009 Easton & Leung-Va 1993 Guo, Wang & Lian 2002 Heimes 1996 Iverson 1992 Karsen, Lau & Bogadek 1998 Lau, Chan, Crow & Ades 2000 Li, Xiao, Qing, Lu & Lazell 2009 Lu, Wang & Li 1999 Luo, Gao & Zhou 2004 Luo, Gao, Wei, Yan, Peng, Zeng, Zhao & Chen 2003 Luo, Liu, Gao, Luo, Huang, Xiao, Wang, Zhou, Luo & Tang 2012 Qu, Lu, Chen, Niu, Lu & Niu 2002 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Song 2002 Tian, Chen, Bao & Liu 2007 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Xu 2007 Zhang & Chen 2002 Zhang & Yu 2002 Zhao & Adler 1993 Zhao 2002 Zhong 2004 Zhou & Chen 2002   Pelodiscus variegatus Large-spotted Softshell Turtle Farkas, Ziegler, Pham, Ong & Fritz 2019 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Rafetus (Bicallosite Softshell Turtles)   Rafetus swinhoei Indochinese Giant Softshell Turtle Fritz & Havas 2006 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang & Che 2020 Zhao & Adler 1993