Reptiles of Georgia    Bibliography of the reptiles of Georgia Taxonomic history of the reptiles of Georgia Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Laudakia (Asian Rock Agamas)   Laudakia caucasia Caucasian Rock Agama Ananjeva & Tuniev 1994 Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bosch & Bischoff 2004 Orlowa in Boehme (ed.) 1981 Sindaco & Jeremchenko 2008 Szczerbak 2003 Yousefkhani, Ficetola, Rastegar-Pouyani, Ananjeva, Rastegar-Pouyani & Masroor 2013  Family Anguidae (Slowworms, etc.) Genus Anguis (Slowworms)   Anguis colchica Eastern Slowworm Bakradze & Chkhikvadze 1993 [as A. fragilis] Bischoff 1978 [as A. fragilis] Bosch & Bischoff 2004 Dely in Boehme (ed.) 1981 [as A. fragilis] Gvozdik, Benkovsky, Crottini, Bellati, Moravec, Romano, Sacchi & Jandzik 2013 Gvozdik, Jandzik, Lymberakis, Jablonski & Moravec 2010 Genus Pseudopus (Eurasian Glass Lizard)   Pseudopus apodus Eurasian Glass Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bosch & Bischoff 2004 Obst in Boehme (ed.) 1981  Family Gekkonidae (Typical Geckos) Genus Tenuidactylus (Central Asian Thin-toed Geckos)   Tenuidactylus caspius Caspian Thin-toed Gecko Bakradze & Chkhikvadze 1993 Bischoff 1978 Bosch & Bischoff 2004 Szczerbak & Golubev 1996  Family Lacertidae (Typical Lizards) Genus Darevskia (Caucasian Rock Lizards)   Darevskia adjarica Northern Red-bellied Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Arribas, Ilgaz & Kumlutas 2018 Bakradze & Chkhikvadze 1993 Bischoff 1978 [as parvula] Bosch & Bischoff 2004 Darevsky & Eiselt 1980 Fu, Murphy & Darevsky 1997 [as parvula] Sindaco & Jeremchenko 2008 Szczerbak 2003 Tuniyev, Ananjeva, Agasyan, Orlov & Tuniyev 2009 Uzzell & Darevsky 1975 [as parvula]   Darevskia alpina Alpine Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008   Darevskia armeniaca Armenian Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 2003 Uzzell & Darevsky 1975   Darevskia brauneri Western Caucasian Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2001 Bosch & Bischoff 2004 Darevsky in Boehme (ed.) 1984 Doronin, Doronina & Bekoshvili 2019 Sindaco & Jeremchenko 2008 Szczerbak 2003   Darevskia caucasica Common Caucasian Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2001 Bischoff 2003 Bosch & Bischoff 2004 Darevsky in Boehme (ed.) 1984 Sindaco & Jeremchenko 2008   Darevskia clarkorum Borcka Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2003 Bosch & Bischoff 2004 Darevsky & Tuniyev 1997 Sindaco & Jeremchenko 2008   Darevskia daghestanica Dagestan Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008   Darevskia dahli Shagali Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2001 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 2003 Uzzell & Darevsky 1975   Darevskia derjugini Artvin Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Fu, Murphy & Darevsky 1997 Orlowa & Bischoff in Boehme (ed.) 1984 Sindaco & Jeremchenko 2008 Szczerbak 2003 Terentjev & Chernov 1965 [1949]   * Darevskia dryada Charnali Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Darevsky & Tuniyev 1997   * Darevskia mixta Adjar Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Fu, Murphy & Darevsky 1997 Sindaco & Jeremchenko 2008 Szczerbak 2003 Uzzell & Darevsky 1975   Darevskia nairensis Nairi Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 2003   Darevskia obscura Borjomi Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Arribas, Candan, Kornilios, Ayaz, Kumlutas, Gul, Yilmaz, Caynak & Ilgaz 2022 Bakradze & Chkhikvadze 1993 Bischoff 1978 Boehme & Bischoff in Boehme (ed.) 1984 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 2003   Darevskia portschinskii River Kura Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 2003 Uzzell & Darevsky 1975   Darevskia praticola Caucasian Meadow Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Arakelyan, Danielyan, Corti, Sindaco & Leviton 2011 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Stugren in Boehme (ed.) 1984   Darevskia raddei Azerbaijan Rock Lizard Arakelyan, Danielyan, Corti, Sindaco & Leviton 2011   Darevskia rudis Spiny-tailed Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Boehme & Bischoff in Boehme (ed.) 1984 Bosch & Bischoff 2004 Fu, Murphy & Darevsky 1997 Sindaco & Jeremchenko 2008 Szczerbak 2003 Terentjev & Chernov 1965 [1949]   Darevskia saxicola Russian Rock Lizard Darevsky in Boehme (ed.) 1984   Darevskia szczerbaki Anapa Rock Lizard Milto, Pestov & Bezman-Moseyko 2010   Darevskia unisexualis White-bellied Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Szczerbak 2003   Darevskia valentini Basarchay Rock Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Arribas, Candan, Kornilios, Ayaz, Kumlutas, Gul, Yilmaz, Caynak & Ilgaz 2022 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 2003 Genus Eremias (Desert and Steppe Runners)   Eremias arguta Kazakhstan Steppe Runner Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Szczerbak 1974   Eremias velox Rapid Steppe Runner Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2003 Bosch & Bischoff 2004 Sindaco & Jeremchenko 2008 Szczerbak 1974 Genus Lacerta (Eurasian Green Lizards)   Lacerta agilis Northern Eurasian Sand Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bischoff 2003 Bischoff in Boehme (ed.) 1984 Bosch & Bischoff 2004 Doronin, Doronina & Bekoshvili 2019 Sindaco & Jeremchenko 2008 Szczerbak 2003   Lacerta media Levant Emerald Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Nettmann & Rykena in Boehme (ed.) 1984 Sindaco & Jeremchenko 2008 Szczerbak 2003 Terentjev & Chernov 1965 [1949]   Lacerta strigata Caspian Green Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Darevsky in Boehme (ed.) 1984 Doronin, Doronina & Bekoshvili 2019 Sindaco & Jeremchenko 2008 Szczerbak 2003 Genus Ophisops (Snake-eyed Lizards)   Ophisops elegans Elegant Snake-eyed Lizard Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 2003 Bosch & Bischoff 2004 Darevsky & Beutler in Boehme (ed.) 1981 Szczerbak 2003 Terentjev & Chernov 1965 [1949]  Family Scincidae (Skinks) Genus Ablepharus (Eurasian Snake-eyed Skinks)   Ablepharus pannonicus Southwest Asian Snake-eyed Skink Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bosch & Bischoff 2004 Grossmann, Kowalski & Schillert 2013 Szczerbak 2003 Genus Eumeces (Afro-Asian Long-legged Skinks)   Eumeces princeps Southwest Asian Long-legged Skink Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bakradze & Chkhikvadze 1993 Bischoff 1978 Bosch & Bischoff 2004 Terentjev & Chernov 1965 [1949]

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx jaculus Western Sand Boa Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Geniez 2018 Kudrjavtsev & Mamet 1989 Stimson 1969  Family Colubridae (Typical Snakes or Colubrids) Genus Coronella (Palearctic Smooth Snakes)   Coronella austriaca Northern Smooth Snake Bosch & Bischoff 2004 Engelmann in Boehme (ed.) 1993 Geniez 2018 Jablonski, Nagy, Avci, Olgun, Kukushkin, Safaei-Mahroo & Jandzik 2019 Steward 1971 Terentjev & Chernov 1965 [1949] Genus Dolichophis (Greater Whip Snakes)   Dolichophis schmidti Red-bellied Whip Snake Arakelyan, Danielyan, Corti, Sindaco & Leviton 2011 Bosch & Bischoff 2004 Geniez 2018 Steward 1971 Genus Eirenis (Middle Eastern Dwarf Snakes)   Eirenis collaris Collared Dwarf Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Geniez 2018 Kudrjavtsev & Mamet 1989 Mahlow, Tillack, Schmidtler & Muller 2013 Szczerbak 2003 Terentjev & Chernov 1965 [1949]   Eirenis modestus Ring-headed Dwarf Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Geniez 2018 Mahlow, Tillack, Schmidtler & Muller 2013 Schmidtler & Baran in Boehme (ed.) 1993 Sikmashvili 1975 Szczerbak 2003 Terentjev & Chernov 1965 [1949] Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe dione Dione Rat Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Obst & Szczerbak in Boehme (ed.) 1993 Schulz 1996 Steward 1971 Terentjev & Chernov 1965 [1949]   Elaphe urartica Urartian Rat Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 [as E. sauromates] Boehme & Szczerbak in Boehme (ed.) 1993 [as E. quatuorlineata] Bosch & Bischoff 2004 [as E. sauromates] Geniez 2018 [as E. sauromates] Jablonski, Kukushkin, Avci, Bunyatova, Kumlutas, Ilgaz, Polyakova, Shiryaev, Tuniyev & Jandzik 2019 Kudrjavtsev & Mamet 1989 [as E. sauromates] Schulz 1996 [as E. sauromates] Szczerbak 2003 [as E. sauromates] Genus Hemorrhois (Palearctic Whip Snakes)   Hemorrhois ravergieri Mountain Whip Snake Bosch & Bischoff 2004 Geniez 2018 Kudrjavtsev & Mamet 1989 Terentjev & Chernov 1965 [1949] Genus Natrix (Eurasian Keelbacks)   Natrix natrix Common European Grass Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 [as megalocephala] Anonymous 2005 Bosch & Bischoff 2004 Geniez 2018 Gruber 1989 [as megalocephala] Kindler, Boehme, Corti, Gvozdik, Jablonski, Jandzik, Metallinou, Siroky & Fritz 2013 Orlov & Tuniyev 1992 [as megalocephala] Orlov & Tuniyev in Boehme (ed.) 1999 [as megalocephala]   Natrix tessellata Dice Snake Anonymous 2005 Bosch & Bischoff 2004 Frotzler, Davitashvili & Mebert 2011 Gruschwitz, Lenz, Mebert & Lanka in Boehme (ed.) 1999 Tuniyev, Tuniyev, Kirschey & Mebert 2011 Genus Platyceps (Flat-headed Whip Snakes)   Platyceps najadum European Slender Whip Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Darevsky & Szczerbak in Boehme (ed.) 1993 Geniez 2018 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus fallax European Cat Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Geniez 2018 Grillitsch & Grillitsch in Boehme (ed.) 1999 Kudrjavtsev & Mamet 1989 Szczerbak 2003 Terentjev & Chernov 1965 [1949] Genus Zamenis (Western Palearctic Rat Snakes)   Zamenis hohenackeri Transcaucasian Rat Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bischoff in Boehme (ed.) 1993 Bosch & Bischoff 2004 Geniez 2018 Hofmann, Mebert, Schulz, Helfenberger, Goecmen & Boehme 2018 Kudrjavtsev & Mamet 1989 Nilson & Andren 1984 Schulz 1996 Sikmashvili 1975 Szczerbak 2003   Zamenis longissimus Common Aesculapian Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Boehme in Boehme (ed.) 1993 Bosch & Bischoff 2004 Geniez 2018 Nilson & Andren 1984 Schulz 1996 Sindaco, Venchi & Grieco 2013 Terentjev & Chernov 1965 [1949]  Family Lamprophiidae (House Snakes, etc.) Genus Malpolon (Montpellier Snakes)   Malpolon insignitus Eastern Montpellier Snake Bosch & Bischoff 2004 Geniez 2018 Terentjev & Chernov 1965 [1949] [as M. monspessulanus]  Family Typhlopidae (Typical Blind Snakes) Genus Xerotyphlops (Desert Blind Snakes)   Xerotyphlops vermicularis European Worm Snake Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Geniez 2018 Grillitsch & Grillitsch in Boehme (ed.) 1993 Kudrjavtsev & Mamet 1989 Szczerbak 2003  Family Viperidae (Vipers) Genus Macrovipera (Giant Vipers)   Macrovipera lebetinus Blunt-nosed Viper Bosch & Bischoff 2004 David & Ineich 1999 Geniez 2018 McDiarmid, Campbell & Toure 1999 Genus Vipera (Typical Vipers or Adders)   Vipera ammodytes European Nose-horned Viper Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 David & Vogel 2010 Geniez 2018 Heckes, Gruber & Stumpel in Joger & Stumpel 2005 Kudrjavtsev & Mamet 1989 McDiarmid, Campbell & Toure 1999 Sindaco, Venchi & Grieco 2013 Szczerbak 2003   Vipera darevskii South Caucasian Viper Tuniyev, Avci, Ilgaz, Olgun, Petrova, Bodrov, Geniez & Teynie 2018 Tuniyev, Iremashvili, Heras & Tuniyev 2014   Vipera dinniki West Caucasian Viper Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 David & Ineich 1999 Geniez 2018 McDiarmid, Campbell & Toure 1999 Orlov & Tuniyev 2005 Sindaco, Venchi & Grieco 2013 Sindaco, Venchi, Carpaneto & Bologna 2000 Tuniyev 2016   Vipera kaznakovi Caucasian Red Viper Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 David & Ineich 1999 David & Vogel 2010 Geniez 2018 Kudrjavtsev & Mamet 1989 McDiarmid, Campbell & Toure 1999 Orlov & Tuniyev 2005 Sindaco, Venchi & Grieco 2013 Sindaco, Venchi, Carpaneto & Bologna 2000 Szczerbak 2003 Tuniyev 2016   Vipera lotievi Caucasian Meadow Viper Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 David & Ineich 1999 McDiarmid, Campbell & Toure 1999 Nilson & Andren 2001 [Russia/Azerbaijan border] Tuniyev 2016 Venchi & Sindaco 2007   Vipera shemakhensis Shemakhan Steppe Viper Tuniyev 2016 [as schemakhensis [sic]] Tuniyev, Iremashvili, Petrova & Kravchenko 2018   * Vipera tuniyevi Georgian Viper Ananjeva, Gabaev, Iremashvili, Lotiev & Petrova 2021

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Emydidae (Western Pond Turtles) Genus Emys (European Pond Turtles)   Emys orbicularis Common European Pond Turtle Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Fritz & Havas 2006 Fritz in Fritz (ed.) 2001 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Geoemydidae (Eastern Pond Turtles) Genus Mauremys (Stripe-necked Turtles)   Mauremys caspica Caspian Turtle Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bosch & Bischoff 2004 Gasperetti, Stimson, Miller, Ross & Gasperetti 1993 Leviton, Anderson, Adler & Minton 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Terentjev & Chernov 1965 [1949] Wischuf & Fritz in Fritz (ed.) 2001  Family Testudinidae (Tortoises) Genus Testudo (Palearctic Tortoises)   Testudo graeca Spur-thighed Tortoise Ananjeva, Orlov, Khalikov, Darevsky, Ryabov & Barabanov 2006 Bonin, Devaux & Dupre 2006 Bosch & Bischoff 2004 Buskirk, Keller & Andreu in Fritz (ed.) 2001 Fritz & Havas 2006 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021