Reptiles of Halmahera [Indonesia] Related checklists:  Indonesia Bibliography of the reptiles of Halmahera Taxonomic history of the reptiles of Halmahera Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus porosus Saltwater Crocodile Rooij 1915

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama Hallermann 2005 Rooij 1915 Setiadi, Hamidy, Abidin, Susanto, Brown, Peterson, Li & Evans 2009 Genus Hydrosaurus (Sail-tailed Agamas)   Hydrosaurus weberi Halmahera Sail-tailed Agama Rooij 1915 Setiadi, Hamidy, Abidin, Susanto, Brown, Peterson, Li & Evans 2009 Wermuth 1967  Family Dibamidae (Blind Skinks) Genus Dibamus (Oriental Blind Skinks)   Dibamus novaeguineae Indo-Australian Blind Skink Greer 1985  Family Gekkonidae (Typical Geckos) Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus deveti Moluccan Bent-toed Gecko Oliver, Edgar, Mumpuni, Iskandar & Lilley 2009 Oliver, Karkkainen, Roesler & Richards 2019   Cyrtodactylus halmahericus Halmahera Bent-toed Gecko Mecke, Hartmann, Mader, Kieckbusch & Kaiser 2016 Oliver, Edgar, Mumpuni, Iskandar & Lilley 2009 Rooij 1915 [as fumosus] Wermuth 1965 Genus Gehyra (Four-clawed Geckos)   Gehyra marginata Giant Moluccan Dtella Flecks, Schmitz, Boehme, Henkel & Ineich 2012 Oliver, Clegg, Fisher, Richards, Taylor & Jocque 2016 Rooij 1915 Wermuth 1965   Gehyra mutilata Common Four-clawed Gecko Rooij 1915   Gehyra oceanica Oceanic Gecko Rooij 1915 Genus Gekko (Tokays)   Gekko vittatus White-striped Gecko Rooij 1915 Setiadi, Hamidy, Abidin, Susanto, Brown, Peterson, Li & Evans 2009 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus frenatus Asian House Gecko Rooij 1915   Hemidactylus platyurus Flat-tailed Gecko Rooij 1915 Genus Lepidodactylus (Scaly-toed Geckos)   Lepidodactylus lugubris Common Mourning Gecko Rooij 1915  Family Scincidae (Skinks) Genus Carlia (Australian-Pacific Rainbow Skinks)   Carlia tutela Moluccan Rainbow Skink Zug 2004 Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Brown 1991 Koch, Arida, Riyanto & Boehme 2009   Emoia caeruleocauda Pacific Blue-tailed Skink Koch, Arida, Riyanto & Boehme 2009   Emoia kuekenthali East Indonesian Slender Skink Brown 1991   Emoia reimschisseli Morotai Slender Skink Brown 1991   Emoia sorex Halmahera Slender Skink Brown 1991 Genus Eugongylus (Short-legged Giant Skinks)   Eugongylus mentovarius Moluccan Recluse Skink Brongersma 1948 Rooij 1915 Shea 2018   Eugongylus rufescens Reddish Giant Skink Koch, Arida, Riyanto & Boehme 2009 Genus Eutropis (South Asian Sun Skinks)   Eutropis multifasciata Many-lined Sun Skink Koch, Arida, Riyanto & Boehme 2009 O'Shea, Kusuma & Kaiser 2018 Rooij 1915 Setiadi, Hamidy, Abidin, Susanto, Brown, Peterson, Li & Evans 2009 Genus Lamprolepis (Malay Archipelago Tree Skinks)   Lamprolepis smaragdina Emerald Tree Skink Koch, Arida, Riyanto & Boehme 2009 Mertens 1929 Rooij 1915 Setiadi, Hamidy, Abidin, Susanto, Brown, Peterson, Li & Evans 2009 Genus Ornithuroscincus (Smooth-eared Skinks)   Ornithuroscincus noctua Moth Skink Koch, Arida, Riyanto & Boehme 2009 Rooij 1915 Zweifel 1979 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus consobrinus Bacan Forest Skink Rooij 1915 Shea 2017   Sphenomorphus solomonis Solomon Forest Skink Greer & Parker 1974 Rooij 1915 Genus Tiliqua (Blue-tongued Skinks)   Tiliqua gigas Giant Blue-tongued Skink Rooij 1915 Shea 1990  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus caerulivirens Turquoise Monitor Boehme 2003 Weijola 2010 Ziegler, Boehme & Philipp 1999   Varanus rainerguentheri Moluccan Mangrove Monitor Boehme 2003 [as indicus] Boehme, Horn & Ziegler 1994 [as indicus] Dryden & Ziegler in Pianka, King & King (eds.) 2004 [as indicus] Koch, Auliya & Ziegler 2010 Somma & Koch 2012 Weijola 2010 Ziegler, Boehme & Philipp 1999 [as indicus] Ziegler, Boehme & Schmitz 2007 Ziegler, Schmitz, Koch & Boehme 2007 [as indicus]   * Varanus yuwonoi Black-backed Mangrove Monitor Boehme 2003 Harvey & Barker 1998 Philipp, Ziegler & Boehme in Pianka, King & King (eds.) 2004 Weijola 2010 Weijola, Donnellan & Lindquist 2016   * Varanus zugorum Halmahera Mangrove Monitor Boehme & Ziegler 2005 Weijola 2010

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Candoia (Pacific Boas)   Candoia paulsoni Pacific Ground Boa Haas 1950 [as Enygrus carinata] Iskandar & Colijn 2001 [as C. carinata] Koch, Arida, Riyanto & Boehme 2009 Lang 2013 O'Shea 1996 Rooij 1917 [as Enygrus carinatus] Setiadi, Hamidy, Abidin, Susanto, Brown, Peterson, Li & Evans 2009 Smith, Chiszar, Tepedelen & Breukelen 2001  Family Colubridae (Typical Snakes or Colubrids) Genus Boiga (Australasian Cat Snakes)   Boiga irregularis Brown Cat Snake Haas 1950 Iskandar & Colijn 2001 Lang 2013 Orlov & Ryabov 2002 Rooij 1917 Setiadi, Hamidy, Abidin, Susanto, Brown, Peterson, Li & Evans 2009 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis modestus Northern Moluccan Bronzeback Haas 1950 Iskandar & Colijn 2001 Lang 2013 Rooij 1917 Rooijen & Vogel 2012   Dendrelaphis pictus Southern Painted Bronzeback Iskandar & Colijn 2001 Genus Stegonotus (Indo-Australian Ground Snakes)   Stegonotus batjanensis Bacan Ground Snake Haas 1950 [as S. modestus] Iskandar & Colijn 2001 Kaiser, Kaiser & O'Shea 2018 Lang 2013 Rooij 1917 Setiadi, Hamidy, Abidin, Susanto, Brown, Peterson, Li & Evans 2009 Genus Tropidonophis (Indo-Australian Keelbacks)   Tropidonophis elongatus Moluccan Keelback Brongersma 1933 Haas 1950 Iskandar & Colijn 2001 Lang 2013 Malnate & Underwood 1988 O'Shea 1996   Tropidonophis halmahericus Red-spotted Keelback Haas 1950 Iskandar & Colijn 2001 Lang 2013 Malnate & Underwood 1988 Rooij 1917   * Tropidonophis punctiventris Spot-bellied Keelback Haas 1950 Iskandar & Colijn 2001 Lang 2013 Malnate & Underwood 1988 Petri & Doria 2012 Rooij 1917   Tropidonophis truncatus Halmahera Keelback Haas 1950 Iskandar & Colijn 2001 Lang 2013 Malnate & Underwood 1988 Rooij 1917  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Laticauda (Sea Kraits)   Laticauda colubrina Yellow-lipped Sea Krait Heatwole, Busack & Cogger 2005 Haas 1950 Kharin, Roedel & Hallermann 2010 Rooij 1917   Laticauda laticaudata Blue-lipped Sea Krait Haas 1950 Kharin, Roedel & Hallermann 2010 Rooij 1917 Genus Thalassophis (Scale-snouted Sea Snake)   Thalassophis anomalus Scale-snouted Sea Snake Rasmussen, Sanders & Lobo 2010  Family Gerrhopilidae (Indo-Malayan Blind Snakes) Genus Gerrhopilus (Indo-Malayan Blind Snakes)   Gerrhopilus suturalis Andai Blind Snake Bosch 1985 [as ater] Brongersma 1934 Hahn 1980 [as ater] Lang 2013 [as ater] McDiarmid, Campbell & Toure 1999 [as ater] McDowell 1974 [as ater] Wallach 1996 Wynn 2021  Family Homalopsidae (Australasian Mud Snakes) Genus Brachyorrhos (Moluccan Short-tailed Snakes)   Brachyorrhos wallacei Halmahera Short-tailed Snake Iskandar & Colijn 2001 [as albus] Lang 2013 Murphy & Voris 2014 Murphy & Voris 2021 Murphy, Mumpuni, Lang, Gower & Sanders 2012 Rooij 1917 [as albus] Genus Cerberus (Dog-faced Water Snakes)   Cerberus schneiderii South East Asian Dog-faced Water Snake Iskandar & Colijn 2001 [as C. rynchops] Koch, Arida, Riyanto & Boehme 2009 [as C. rynchops] Lang 2013 Murphy & Voris 2014 Rooij 1917 [as C. rynchops]  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Iskandar & Colijn 2001 Koch, Arida, Riyanto & Boehme 2009 Lang 2013 Mertens 1930 Rooij 1917 Genus Simalia (Scrub Pythons)   Simalia tracyae Halmahera Scrub Python Harvey, Barker, Ammerman & Chippindale 2000 Iskandar & Colijn 2001 Lang 2013 McDowell 1975 [as Python amethistinus] Rooij 1917 [as Python amethystinus]  Family Typhlopidae (Typical Blind Snakes) Genus Ramphotyphlops (Western Pacific Blind Snakes)   Ramphotyphlops flaviventer Yellow-bellied Blind Snake Brongersma 1934 Hahn 1980 Iskandar & Colijn 2001 Lang 2013 McDiarmid, Campbell & Toure 1999 McDowell 1974 Genus Virgotyphlops (Brahminy Blind Snake)   Virgotyphlops braminus Brahminy Blind Snake Hahn 1980 Iskandar & Colijn 2001 Lang 2013 McDiarmid, Campbell & Toure 1999 Mertens 1930 Rooij 1917 Wallach 2009

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Koch, Arida, Riyanto & Boehme 2009  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle Mertens 1930 Rooij 1915 Wermuth & Mertens 1977