Reptiles of North Macedonia [formerly FYR Macedonia]   Bibliography of the reptiles of North Macedonia Taxonomic history of the reptiles of North Macedonia Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Anguidae (Slowworms, etc.) Genus Anguis (Slowworms)   Anguis colchica Eastern Slowworm Gvozdik, Jandzik, Lymberakis, Jablonski & Moravec 2010 Pozzi 1966 [as A. fragilis] Sterijovski, Tomovic & Ajtic 2014 [as A. fragilis] Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 [as A. fragilis]   Anguis graeca Greek Slowworm Mikulicek, Jablonski, Palenik, Gvozdik & Jandzik 2018 Genus Pseudopus (Eurasian Glass Lizard)   Pseudopus apodus Eurasian Glass Lizard Arnold & Ovenden 2002 Brelih & Dzukic 1974 Mikulicek, Jablonski, Palenik, Gvozdik & Jandzik 2018 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016  Family Gekkonidae (Typical Geckos) Genus Mediodactylus (Mediterranean Bent-toed Geckos)   Mediodactylus kotschyi Balkan Bent-toed Gecko Arnold & Ovenden 2002 Beutler in Boehme (ed.) 1981 Brelih & Dzukic 1974 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016  Family Lacertidae (Typical Lizards) Genus Algyroides (Keeled Lizards)   Algyroides nigropunctatus Blue-throated Keeled Lizard Arnold & Ovenden 2002 Bischoff in Boehme (ed.) 1981 Brelih & Dzukic 1974 Dimovski 1964 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Genus Lacerta (Eurasian Green Lizards)   Lacerta agilis Northern Eurasian Sand Lizard Bischoff in Boehme (ed.) 1984 Brelih & Dzukic 1974 Dimovski 1959 Dimovski 1964 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016   Lacerta trilineata Western Balkan Green Lizard Arnold & Ovenden 2002 Brelih & Dzukic 1974 Kornilios, Thanou, Lymberakis, Ilgaz, Kumlutas & Leache 2020 Nettmann & Rykena in Boehme (ed.) 1984 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016   Lacerta viridis Eastern Green Lizard Boehme, Fritz, Kotenko, Dzukic, Ljubisavljevic, Tzankov & Berendonk 2007 Dimovski 1959 Marzahn, Mayer, Joger, Ilgaz, Jablonski, Kindler, Kumlutas, Nistri, Schneewiess, Vamberger, Zagar & Fritz 2015 Necas, Modry & Zavadil 1997 Pafilis & Maragou 2013 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 [as "viridis complex"] Genus Podarcis (Wall Lizards)   Podarcis erhardii Aegean Wall Lizard Arnold & Ovenden 2002 Brelih & Dzukic 1974 Gruber in Boehme (ed.) 1986 Sindaco & Jeremchenko 2008 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016   Podarcis muralis Common European Wall Lizard Brelih & Dzukic 1974 Dimovski 1959 Gruschwitz & Boehme in Boehme (ed.) 1986 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016   Podarcis tauricus Balkan Wall Lizard Brelih & Dzukic 1974 Dimovski 1959 Pozzi 1966 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Genus Zootoca (Eurasian Forest Lizards)   Zootoca vivipara Viviparous Lizard Arnold & Ovenden 2002 Brelih & Dzukic 1974 Dimovski 1964 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016  Family Scincidae (Skinks) Genus Ablepharus (Eurasian Snake-eyed Skinks)   Ablepharus kitaibelii European Snake-eyed Skink Brelih & Dzukic 1974 Dimovski 1959 Pozzi 1966 Sindaco & Jeremchenko 2008 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx jaculus Western Sand Boa Arnold & Ovenden 2002 Brelih & Dzukic 1974 Geniez 2018 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Tokar & Obst in Boehme (ed.) 1993 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016  Family Colubridae (Typical Snakes or Colubrids) Genus Coronella (Palearctic Smooth Snakes)   Coronella austriaca Northern Smooth Snake Brelih & Dzukic 1974 Engelmann in Boehme (ed.) 1993 Geniez 2018 Jablonski, Nagy, Avci, Olgun, Kukushkin, Safaei-Mahroo & Jandzik 2019 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Genus Dolichophis (Greater Whip Snakes)   Dolichophis caspius Caspian Whip Snake Brelih & Dzukic 1974 Geniez 2018 Nagy, Bellaagh, Wink, Paunovic & Korsos 2010 Pozzi 1966 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Genus Elaphe (Palearctic-Oriental Rat Snakes)   Elaphe quatuorlineata Western Four-lined Rat Snake Arnold & Ovenden 2002 Boehme & Szczerbak in Boehme (ed.) 1993 Brelih & Dzukic 1974 Geniez 2018 Qi, Shi, Ma, Gao, Bu, Grismer, Li & Wang 2021 Schulz 1996 Sindaco, Venchi & Grieco 2013 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Genus Hierophis (Eurasian Whip Snakes)   Hierophis gemonensis Balkan Whip Snake Geniez 2018 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Genus Natrix (Eurasian Keelbacks)   Natrix natrix Common European Grass Snake Brelih & Dzukic 1974 Geniez 2018 Kabisch in Boehme (ed.) 1999 Kindler, Boehme, Corti, Gvozdik, Jablonski, Jandzik, Metallinou, Siroky & Fritz 2013 Pozzi 1966 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016   Natrix tessellata Dice Snake Brelih & Dzukic 1974 Geniez 2018 Necas, Modry & Zavadil 1997 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Ajtic, Tomovic, Djordjevic, Djurakic, Golubovic, Crnobrnja-Isailovic, Ballouard, Desmont, Groumpf & Bonnet 2011 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Genus Platyceps (Flat-headed Whip Snakes)   Platyceps najadum European Slender Whip Snake Arnold & Ovenden 2002 Brelih & Dzukic 1974 Darevsky & Szczerbak in Boehme (ed.) 1993 Geniez 2018 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus fallax European Cat Snake Arnold & Ovenden 2002 Brelih & Dzukic 1974 Geniez 2018 Grillitsch & Grillitsch in Boehme (ed.) 1999 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Genus Zamenis (Western Palearctic Rat Snakes)   Zamenis longissimus Common Aesculapian Snake Brelih & Dzukic 1974 Geniez 2018 Necas, Modry & Zavadil 1997 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016   Zamenis situla Leopard Snake Arnold & Ovenden 2002 Brelih & Dzukic 1974 Geniez 2018 Obst, Szczerbak & Boehme in Boehme (ed.) 1993 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016  Family Lamprophiidae (House Snakes, etc.) Genus Malpolon (Montpellier Snakes)   Malpolon insignitus Eastern Montpellier Snake Arnold & Ovenden 2002 Brelih & Dzukic 1974 Geniez 2018 Haan in Boehme 1999 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 [as M. monspessulanus] Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016  Family Typhlopidae (Typical Blind Snakes) Genus Xerotyphlops (Desert Blind Snakes)   Xerotyphlops vermicularis European Worm Snake Arnold & Ovenden 2002 Brelih & Dzukic 1974 Dimovski 1959 Geniez 2018 Grillitsch & Grillitsch in Boehme (ed.) 1993 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016  Family Viperidae (Vipers) Genus Vipera (Typical Vipers or Adders)   Vipera ammodytes European Nose-horned Viper Brelih & Dzukic 1974 Dimovski 1959 Geniez 2018 Heckes, Gruber & Stumpel in Joger & Stumpel 2005 Jelic, Ajtic, Sterijovski, Crnobrnja-Isailovic, Lelo & Tomovic 2013 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Tomovic & Dzukic 2003 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Ursenbacher, Schweiger, Tomovic, Crnobrnja-Isailovic, Fumagalli & Mayer 2008   Vipera berus Northern Eurasian Viper Brelih & Dzukic 1974 David & Ineich 1999 Dimovski 1964 Geniez 2018 Nilson, Andren & Voelkl in Joger & Stumpel 2005 Pozzi 1966 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016   Vipera ursinii Meadow and Karst Vipers Arnold & Ovenden 2002 Brelih & Dzukic 1974 David & Ineich 1999 Dely & Joger in Joger & Stumpel 2005 Dimovski 1964 Geniez 2018 Jelic, Ajtic, Sterijovski, Crnobrnja-Isailovic, Lelo & Tomovic 2013 Nilson & Andren 2001 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski 2006 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Venchi & Sindaco 2007

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Emydidae (Western Pond Turtles) Genus Emys (European Pond Turtles)   Emys orbicularis Common European Pond Turtle Brelih & Dzukic 1974 Fritz in Fritz (ed.) 2001 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016  Family Geoemydidae (Eastern Pond Turtles) Genus Mauremys (Stripe-necked Turtles)   Mauremys rivulata Balkan Pond Turtle Arnold & Ovenden 2002 Brelih & Dzukic 1974 Mantziou & Rifai 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016 Wischuf & Busack in Fritz (ed.) 2001  Family Testudinidae (Tortoises) Genus Testudo (Palearctic Tortoises)   Testudo graeca Spur-thighed Tortoise Arnold & Ovenden 2002 Brelih & Dzukic 1974 Buskirk, Keller & Andreu in Fritz (ed.) 2001 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016   Testudo hermanni Greek Tortoise Bertolero, Cheylan, Hailey, Livoreil & Willemsen 2011 Brelih & Dzukic 1974 Cheylan in Fritz (ed.) 2001 Ljubisavljevic, Dzukic, Vukov & Kalezic 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Speybroeck, Beukema, Bok & Voort 2016 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husak, Koleska & Jablonski 2016