Reptiles of Bioko  (formerly Fernando Po or Masie Ngueme Biyogo Island) [Equatorial Guinea] Related checklists:  Equatorial Guinea Bibliography of the reptiles of Bioko Taxonomic history of the reptiles of Bioko Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama lebretoni Cameroon Rainbow Agama Eisentraut 1973 [as agama] Leache, Grummer, Miller, Krishnan, Fujita, Boehme, Schmitz, LeBreton, Ineich, Chirio, Ofori-boateng, Eniang, Greenbaum, Roedel & Wagner 2016 Mediannikov, Trape & Trape 2012 [as agama] Mertens 1964 [as agama] Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Wagner, Barej & Schmitz 2009  Family Chamaeleonidae (Chameleons) Genus Rhampholeon (African Leaf Chameleons)   Rhampholeon spectrum Spectral Leaf Chameleon Eisentraut 1973 Glaw 2015 Klaver & Boehme 1997 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Tilbury 2010 Genus Trioceros (Equatorial Chameleons)   Trioceros cristatus Crested Chameleon Eisentraut 1973 Glaw 2015 Klaver & Boehme 1997 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Tilbury 2010   * Trioceros feae Bioko Montane Chameleon Eisentraut 1973 Glaw 2015 Klaver & Boehme 1997 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Tilbury 2010   Trioceros oweni Central African Three-horned Chameleon Eisentraut 1973 Glaw 2015 Klaver & Boehme 1997 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Tilbury 2010  Family Gekkonidae (Typical Geckos) Genus Ancylodactylus (African Rock and Forest Geckos)   Ancylodactylus koehleri Cameroon Mountains Rock Gecko Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus angulatus West African House Gecko Eisentraut 1973 Sanchez-Vialas, Calvo-Revuelta & Riva 2022   * Hemidactylus biokoensis Bioko Half-toed Gecko Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Wagner, Leache & Fujita 2014 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus conraui Cameroon Dwarf Gecko Eisentraut 1973 Loveridge 1947 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Wermuth 1965  Family Lacertidae (Typical Lizards) Genus Poromera (African Swamp Lizard)   Poromera fordii African Swamp Lizard Bischoff 1992 Eisentraut 1973 Mertens 1968 Sanchez-Vialas, Calvo-Revuelta & Riva 2022  Family Scincidae (Skinks) Genus Lacertaspis (Central African Leaf-litter Skinks)   Lacertaspis gemmiventris Yellow-bellied Mountain Skink Eisentraut 1973 Fuhn 1972 Mertens 1964 Perret 1973 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Schmitz, Ineich & Chirio 2005   Lacertaspis reichenowii Cameroon Mountain Leaf-Litter Skink Eisentraut 1973 Fuhn 1972 Mertens 1964 Perret 1973 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Schmitz, Ineich & Chirio 2005 Genus Leptosiaphos (Slender Leaf-litter Skinks)   Leptosiaphos vigintiserierum Twenty-lined Mountain Skink Eisentraut 1973 Fuhn 1972 Mertens 1964 Perret 1973 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 [as "L. cf. vigintiserierum"] Schmitz, Ineich & Chirio 2005 Genus Melanoseps (Black Limbless Skinks)   Melanoseps occidentalis Western Limbless Skink Brygoo & Roux-Esteve 1981 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Mochlus (African Supple Skinks)   Mochlus fernandi Western Red-flanked Skink Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Wagner, Boehme, Pauwels & Schmitz 2009 Genus Panaspis (African Snake-eyed Skinks)   Panaspis breviceps Short-headed Snake-eyed Skink Eisentraut 1973 Fuhn 1972 Mertens 1964 Perret 1973 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Schmitz, Ineich & Chirio 2005 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis affinis Lesser Brown-flanked Skink Eisentraut 1973 Mertens 1964 [as blandingi] Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Schatti & Loumont 1992   Trachylepis polytropis Greater Forest Skink Eisentraut 1973 Hoogmoed 1974 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus stellatus African Forest Monitor Boehme & Ziegler 1997 [as ornatus] Boehme & Ziegler in Pianka, King & King (eds.) 2004 [as V. ornatus] Eisentraut 1973 [as ornatus] Mertens 1964 [as ornatus] Sanchez-Vialas, Calvo-Revuelta & Riva 2022 [as V. niloticus]

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Calabaria (Calabar Boa)   Calabaria reinhardtii Calabar Boa Chippaux & Jackson 2019 Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Stimson 1969  Family Colubridae (Typical Snakes or Colubrids) Genus Afronatrix (West African Brown Water Snake)   Afronatrix anoscopus West African Brown Water Snake Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Wallach, Williams & Boundy 2014 Genus Dipsadoboa (Nocturnal Tree Snakes)   Dipsadoboa unicolor Unicoloured Nocturnal Tree Snake Mertens 1964 Rasmussen 1993 Sanchez-Vialas, Calvo-Revuelta & Riva 2022   Dipsadoboa viridis Green Nocturnal Tree Snake Eisentraut 1973 [as elongata] Mertens 1964 [as elongata] Rasmussen 1993 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Grayia (Equatorial African Water Snakes)   Grayia caesar Long-tailed Water Snake Chippaux & Jackson 2019 Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Hapsidophrys (Emerald Snakes)   Hapsidophrys lineatus Black-lined Green Snake Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022   Hapsidophrys smaragdinus Common Emerald Snake Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Natriciteres (African Marsh Snakes)   Natriciteres fuliginoides Collared Marsh Snake Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Philothamnus (African Green Snakes)   Philothamnus carinatus Barred Green Snake Eisentraut 1973 Loveridge 1957 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022   Philothamnus nitidus Cameroon Green Snake Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Rhamnophis (Dagger-tooth Tree Snakes)   Rhamnophis aethiopissa Large-eyed Green Tree Snake Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Thelotornis (African Vine Snakes)   Thelotornis kirtlandii Northern Vine Snake Eisentraut 1973 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Thrasops (Black Tree Snakes)   Thrasops flavigularis Yellow-throated Black Tree Snake Broadley & Wallach 2002 Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Toxicodryas (African Tree Cat Snakes)   Toxicodryas adamanteus Eastern Powdered Cat Snake Eisentraut 1973 [as pulverulenta] Mertens 1964 [as pulverulenta] Sanchez-Vialas, Calvo-Revuelta & Riva 2022   Toxicodryas blandingii Western Black-and-Yellow Tree Snake Eisentraut 1973 Greenbaum, Allen, Vaughan, Pauwels, Wallach, Kusamba, Muninga, Aristote, Mali, Badjedjea, Penner, Roedel, Rivera, Sterkhova, Johnson, Tapondjou Nkonmeneck & Brown 2021 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Dendroaspis (Mambas)   Dendroaspis jamesonii Central African Green Mamba Eisentraut 1973 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Naja (Typical Cobras)   Naja melanoleuca Central African Forest Cobra Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Pseudohaje (Tree Cobras)   Pseudohaje goldii Central African Tree Cobra David & Ineich 1999 Eisentraut 1973 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022  Family Lamprophiidae (House Snakes, etc.) Genus Boaedon (Afro-Arabian House Snakes)   Boaedon olivaceus Olive House Snake Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022   Boaedon virgatus West African House Snake Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Bothrolycus (Black Loreal-pitted Snake)   Bothrolycus ater Black Loreal-pitted Snake Boulenger 1919 Chippaux & Jackson 2019 Eisentraut 1973 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Bothrophthalmus (Red-and-Black Striped Snakes)   Bothrophthalmus brunneus Brown-bellied Snake Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Buhoma (African Forest Snakes)   Buhoma depressiceps Pale-headed Forest Snake Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Chamaelycus (African Banded Snakes)   Chamaelycus fasciatus Ituri Banded Snake Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Hormonotus (Yellow Forest Snake)   Hormonotus modestus Yellow Forest Snake Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Lycophidion (African Wolf Snakes)   Lycophidion laterale Masked Wolf Snake Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Mehelya (Greater File Snakes)   Mehelya poensis Western Forest File Snake Chippaux & Jackson 2019 Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Polemon (African Snake Eaters)   Polemon collaris Fawn-headed Snake Eater Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022  Family Leptotyphlopidae (Thread Snakes) Genus Tricheilostoma (West African Forest Thread Snakes)   Tricheilostoma sundewalli Central African Thread Snake Eisentraut 1973 Hahn & Wallach 1998 Hahn 1980 McDiarmid, Campbell & Toure 1999 Mertens 1964 [as gestri] Sanchez-Vialas, Calvo-Revuelta & Riva 2022  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python sebae Northern Rock Python Broadley 1984 Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops angolensis Golden Blind Snake Sanchez-Vialas, Calvo-Revuelta & Riva 2022   Afrotyphlops congestus Marbled Blind Snake Broadley & Wallach 2009 Eisentraut 1973 Hahn 1980 McDiarmid, Campbell & Toure 1999 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Wallach, Williams & Boundy 2014   Afrotyphlops punctatus Spotted Blind Snake Wallach, Williams & Boundy 2014  Family Viperidae (Vipers) Genus Atheris (African Bush Vipers)   * Atheris hetfieldi Bioko Bush Viper Ceriaco, Marques & Bauer 2020 Sanchez-Vialas, Calvo-Revuelta & Riva 2022   Atheris squamigera Southern Green Bush Viper Broadley 1998 Capocaccia 1961 Ceriaco, Marques & Bauer 2020 Eisentraut 1973 Loveridge 1957 McDiarmid, Campbell & Toure 1999 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Wallach, Williams & Boundy 2014 Genus Bitis (African Adders)   Bitis nasicornis African Nose-horned Viper David & Ineich 1999 Eisentraut 1973 Mertens 1964 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Genus Causus (Night Adders)   Causus maculatus West African Night Adder Sanchez-Vialas, Calvo-Revuelta & Riva 2022

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Brongersma in Bjorndal (ed.) 1982 Butynski 1996 Eisentraut 1964 Mettler, Clyde-Brockway, Honarvar & Paladino 2019 Rader, Mba, Morra & Hearn 2006 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Stuart, Adams & Jenkins 1990 Tomas, Godley, Castroviejo & Raga 2010 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Brongersma in Bjorndal (ed.) 1982 Butynski 1996 Eisentraut 1964 Rader, Mba, Morra & Hearn 2006 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Stuart, Adams & Jenkins 1990 Tomas, Godley, Castroviejo & Raga 2010 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Butynski 1996 Rader, Mba, Morra & Hearn 2006 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Tomas, Fretey, Raga & Castroviejo 2001 Tomas, Godley, Castroviejo & Raga 2010  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Butynski 1996 Rader, Mba, Morra & Hearn 2006 Sanchez-Vialas, Calvo-Revuelta & Riva 2022 Tomas, Godley, Castroviejo & Raga 2010