Reptiles of
North Macedonia  

 

Bibliography of the reptiles of North Macedonia Taxonomic history of the reptiles of North Macedonia Introduced species
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© Rune Midtgaard

Reptile biodiversity of
North Macedonia
excl. introduced, extinct, unconfirmed, and unnamed species
See complete statistics for families and genera
Families
Endemic
families
Endemism
(families)
Genera
Endemic
genera
Endemism
(genera)
Species
Endemic
species
Endemism
(species)
Species in %
of total
Lizards (Sauria) 4 0 0.0 % 8 0 0.0 % 13 0 0.0 % 39.4 %
Snakes (Serpentes) 5 0 0.0 % 12 0 0.0 % 16 0 0.0 % 48.5 %
Turtles (Testudines) 3 0 0.0 % 3 0 0.0 % 4 0 0.0 % 12.1 %
Total 12 0 0.0 % 23 0 0.0 % 33 0 0.0 % 100.0 %

Checklist
General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable.
Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:
Lizards (Sauria)
= endemicEX = extinct or assumed extinct

Species
References
 Family Anguidae (Slowworms, etc.)
Genus Anguis (Slowworms)
 
Anguis colchica
Eastern Slowworm
Gvozdik, Jandzik, Lymberakis, Jablonski & Moravec 2010 Pozzi 1966 [as fragilis] Sterijovski, Tomovic & Ajtic 2014 [as fragilis] Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016 [as fragilis]
 
Anguis graeca
Greek Slowworm
Mikulicek, Jablonski, Palenik, Gvozdik & Jandzik 2018
Genus Pseudopus (Eurasian Glass Lizard)
 
Pseudopus apodus
Eurasian Glass Lizard
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Mikulicek, Jablonski, Palenik, Gvozdik & Jandzik 2018 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 Family Gekkonidae (Typical Geckos)
Genus Mediodactylus (Mediterranean Bent-toed Geckos)
 
Mediodactylus kotschyi
Balkan Bent-toed Gecko
Arnold & Ovenden 2002 Beutler in Böhme 1981 Brelih & Dzukic 1974 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 Family Lacertidae (Typical Lizards)
Genus Algyroides (Keeled Lizards)
 
Algyroides nigropunctatus
Blue-throated Keeled Lizard
Arnold & Ovenden 2002 Bischoff in Böhme 1981 Brelih & Dzukic 1974 Dimovski 1964 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
Genus Lacerta (Eurasian Green Lizards)
 
Lacerta agilis
Northern Eurasian Sand Lizard
Bischoff 1984a Brelih & Dzukic 1974 Dimovski 1959 Dimovski 1964 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 
Lacerta trilineata
Balkan Green Lizard
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Nettmann & Rykena in Böhme 1984 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 
Lacerta viridis
Eastern Green Lizard
Böhme, Fritz, Kotenko, Dzukic, Ljubisavljevic, Tzankov & Berendonk 2007 Dimovski 1959 Marzahn, Mayer, Joger, Ilgaz, Jablonski, Kindler, Kumlutas, Nistri, Schneewiess, Vamberger, Zagar & Fritz 2015 Necas, Modry & Zavadil 1997 Pafilis & Maragou 2013 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016 [as "viridis complex"]
Genus Podarcis (Wall Lizards)
 
Podarcis erhardii
Aegean Wall Lizard
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Gruber in Böhme (ed.) 1986 Sindaco & Jeremchenko 2008 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 
Podarcis muralis
Common European Wall Lizard
Brelih & Dzukic 1974 Dimovski 1959 Gomard 2011 Gruschwitz & Böhme in Böhme (ed.) 1986 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 
Podarcis tauricus
Balkan Wall Lizard
Brelih & Dzukic 1974 Dimovski 1959 Gomard 2011 Pozzi 1966 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
Genus Zootoca (Viviparous Lizard)
 
Zootoca vivipara
Viviparous Lizard
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Dimovski 1964 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 Family Scincidae (Skinks)
Genus Ablepharus (Eurasian Snake-eyed Skinks)
 
Ablepharus kitaibelii
European Snake-eyed Skink
Brelih & Dzukic 1974 Dimovski 1959 Pozzi 1966 Sindaco & Jeremchenko 2008 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016

Snakes (Serpentes)
= endemicEX = extinct or assumed extinct

Species
References
 Family Boidae (Typical Boas)
Genus Eryx (Sand Boas)
 
Eryx jaculus
Western Sand Boa
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Sterijovski, Tomovic & Ajtic 2014 Tokar & Obst in Böhme (ed.) 1993 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 Family Colubridae (Typical Snakes or Colubrids)
Genus Coronella (Palearctic Smooth Snakes)
 
Coronella austriaca
Northern Smooth Snake
Brelih & Dzukic 1974 Engelmann in Böhme (ed.) 1993 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
Genus Dolichophis (Greater Whip Snakes)
 
Dolichophis caspius
Caspian Whip Snake
Brelih & Dzukic 1974 Nagy, Bellaagh, Wink, Paunovic & Korsós 2010 Pozzi 1966 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
Genus Elaphe (Palearctic-Oriental Rat Snakes)
 
Elaphe quatuorlineata
Western Four-lined Rat Snake
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Böhme & Szczerbak in Böhme (ed.) 1993 Gomard 2011 Schulz 1996 Sindaco, Venchi & Grieco 2013 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
Genus Hierophis (Eurasian Whip Snakes)
 
Hierophis gemonensis
Balkan Whip Snake
Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
Genus Natrix (Eurasian Keelbacks)
 
Natrix natrix
Common European Grass Snake
Brelih & Dzukic 1974 Gomard 2011 Kabisch in Böhme (ed.) 1999 Kindler, Böhme, Corti, Gvozdik, Jablonski, Jandzik, Metallinou, Siroký & Fritz 2013 Pozzi 1966 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 
Natrix tessellata
Dice Snake
Brelih & Dzukic 1974 Gomard 2011 Necas, Modry & Zavadil 1997 Sterijovski, Ajtic, Tomovic, Djordjevic, Djurakic, Golubovic, Crnobrnja-Isailovic, Ballouard, Desmont, Groumpf & Bonnet 2011 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
Genus Platyceps (Flat-headed Whip Snakes)
 
Platyceps najadum
European Slender Whip Snake
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Darevsky & Szczerbak in Böhme (ed.) 1993 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
Genus Telescopus (Cat Snakes & Tiger Snakes)
 
Telescopus fallax
European Cat Snake
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Grillitsch & Grillitsch in Böhme (ed.) 1999 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
Genus Zamenis (Western Palearctic Rat Snakes)
 
Zamenis longissimus
Common Aesculapian Snake
Brelih & Dzukic 1974 Gomard 2011 Necas, Modry & Zavadil 1997 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 
Zamenis situla
Leopard Snake
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Obst, Szczerbak & Böhme in Böhme (ed.) 1993 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 Family Lamprophiidae (House Snakes, etc.)
Genus Malpolon (Montpellier Snakes)
 
Malpolon insignitus
Eastern Montpellier Snake
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Haan in Böhme 1999 Sterijovski, Tomovic & Ajtic 2014 [as monspessulanus] Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 Family Typhlopidae (Typical Blind Snakes)
Genus Xerotyphlops (Desert Blind Snakes)
 
Xerotyphlops vermicularis
European Worm Snake
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Dimovski 1959 Grillitsch & Grillitsch in Böhme (ed.) 1993. Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 Family Viperidae (Vipers)
Genus Vipera (Typical Vipers or Adders)
 
Vipera ammodytes
European Nose-horned Viper
Brelih & Dzukic 1974 Dimovski 1959 Gomard 2011 Heckes, Gruber & Stümpel in Joger & Stümpel 2005 Jelic, Ajtic, Sterijovski, Crnobrnja-Isailovic, Lelo & Tomovic 2013 Sterijovski, Tomovic & Ajtic 2014 Tomovic & Dzukic 2003 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016 Ursenbacher, Schweiger, Tomovic, Crnobrnja-Isailovic, Fumagalli & Mayer 2008
 
Vipera berus
Northern Eurasian Viper
Brelih & Dzukic 1974 David & Ineich 1999 Dimovski 1964 Nilson, Andrén & Völkl in Joger & Stümpel 2005 Pozzi 1966 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 
Vipera ursinii
Meadow and Karst Vipers
Arnold & Ovenden 2002 Brelih & Dzukic 1974 David & Ineich 1999 Dely & Joger in Joger & Stümpel 2005 Dimovski 1964 Jelic, Ajtic, Sterijovski, Crnobrnja-Isailovic, Lelo & Tomovic 2013 Nilson & Andrén 2001 Sterijovski 2006 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016 Venchi & Sindaco 2007

Turtles (Testudines)
= endemicEX = extinct or assumed extinct

Species
References
 Family Emydidae (Western Pond Turtles)
Genus Emys (European Pond Turtles)
 
Emys orbicularis
Common European Pond Turtle
Brelih & Dzukic 1974 Fritz in Fritz (ed.) 2001 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2017 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 Family Geoemydidae (Eastern Pond Turtles)
Genus Mauremys (Stripe-necked Turtles)
 
Mauremys rivulata
Balkan Pond Turtle
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Mantziou & Rifai 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2017 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016 Wischuf & Busack in Fritz (ed.) 2001
 Family Testudinidae (Tortoises)
Genus Testudo (Palearctic Tortoises)
 
Testudo graeca
Spur-thighed Tortoise
Arnold & Ovenden 2002 Brelih & Dzukic 1974 Buskirk, Keller & Andreu in Fritz (ed.) 2001 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2017 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016
 
Testudo hermanni
Greek Tortoise
Bertolero, Cheylan, Hailey, Livoreil & Willemsen 2011 Brelih & Dzukic 1974 Cheylan in Fritz (ed.) 2001 Gomard 2011 Ljubisavljevic, Dzukic, Vukov & Kalezic 2014 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2017 Sterijovski, Tomovic & Ajtic 2014 Uhrin, Havas, Minarik, Kodejs, Bugos, Danko, Husák, Koleska & Jablonski 2016