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© Rune Midtgaard


Taxonomy of the family Colubridae
Bibliography of the genus Coelognathus
Biodiversity of the family Colubridae








Genus
Coelognathus

Oriental Rat Snakes, (Racers)

Orientalische Rattennattern

Orientalske Rottesnoge

1843 Coelognathus Fitzinger (type species: Coluber radiatus Boie 1827)
1849 Cynophis Gray (type species: Cynophis bistrigatus Gray 1849)
1853 Compsosoma Dumeril [not Compsosoma Audinet-Serville 1835 (Coleoptera)] (type species: Coluber radiatus Boie 1827)
1853 Plagiodon Dumeril (type species: Coluber helena Daudin 1803)
Contents: 7 species, of which 2 (28.6%) are endemic.
Endemism: 0% 100%
Remarks: Previously included in Elaphe (e.g., Smith 1943; Schulz 1996). Revalidated by Helfenberger (2001).
Distribution: S. Asia, Malay Archipelago.
Reported from: Bangladesh, Bhutan, Brunei, Cambodia, China (Fujian, Guangdong [incl. Nan Ao Island], Guangxi, Guizhou, Hong Kong [incl. Shek Kwu Chau Island], Yunnan), East Timor (incl. Atauro Island), India (Andaman & Nicobar Islands [Andaman Islands, Nicobar Islands], Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Chhattisgarh, Delhi, Goa, Gujarat, Haryana, Himachal Pradesh, Jammu & Kashmir, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Punjab, Rajasthan, Sikkim, Tamil Nadu, Telangana, Tripura, Uttar Pradesh, Uttarakhand, West Bengal), Indonesia (Adonara, Alor, Anambas Islands, Bali, Banggai Islands, Bangka, Belitung, Buton, Ende, Enggano, Flores, Java, Kalimantan, Komodo, Lembata, Lombok, Nias, Penida, Riau Islands, Rinca, Rote, Sangir Islands, Semau, Siberut, Simeulue, Sipura, Sulawesi, Sumatra, Sumba, Sumbawa, Weh, West Timor, Wetar), Laos, Malaysia (Sabah, Sarawak, West Malaysia [incl. Langkawi Archipelago, Seribuat Archipelago (incl. Tioman)]), Myanmar, Nepal, Philippines (Babuyan Islands [Barit, Calayan, Dalupiri], Balabac, Batan Islands [Batan], Bohol, Calamian Islands [Busuanga, Calauit, Culion], Camiguin Sur, Camotes Islands [Pacijan, Ponson, Poro], Catanduanes, Cebu, Dinagat, Guimaras, Leyte, Luzon, Masbate, Mindanao, Mindoro, Negros, Palawan, Panay, Samar, Siquijor, Sulu Islands [Basilan, Bongao, Jolo, Sangasanga, Siasi, Sibutu, Tawi-Tawi], Tablas), Singapore, Sri Lanka, Thailand, Vietnam (incl. Cat Ba Island).

Coelognathus enganensis

Enggano Rat Snake

Enggano-Rattennatter

Enggano-rottesnog

1892 Coluber enganensis Vinciguerra
Elaphe enganensis Barbour 1912
Elaphe subradiata enganensis Mertens 1930
Gonyosoma enganensis Dowling 1958
Coelognathus enganensis Das 2012

Remarks: Status uncertain. Listed as a valid species by Iskandar & Colijn (2001), but regarded by some authors as a subspecies of subradiatus (e.g., Schulz 1996). Utiger, Schätti & Helfenberger (2005) stated that the affinities with this form may be with the erythrurus-philippinus complex.
Distribution: Indonesia (Enggano).


Coelognathus erythrura

Philippine Rat Snake, Philippine Trinket Snake, Reddish Rat Snake, Red-fronted Racer

Philippinische Rattennatter

Filippinsk Rottesnog

1854 Plagiodon erythrurus Dumeril, Bibron & Dumeril
Compsosoma melanurum erythrurus Fischer 1885
Coluber erythrurus Boulenger 1894
Elaphe erythrura Griffin 1910
Coelognathus erythrurus Helfenberger 2001
1863 Elaphis melanurus celebensis Jan
Elaphe erythrura celebensis Leviton 1979
1863 Elaphis melanurus manillensis Jan
Elaphe erythrura manilliensis Leviton 1979
Coelognathus erythrura manilliensis David, Pauwels, Lays & Lenglet 2006
1863 Elaphis nyctenurus Jan (Iskandar & Colijn 2001)
1924 Coluber walli Werner (Leviton 1979)
1977 Elaphe erythrura psephenoura Leviton
Coelognathus erythrura psephenoura David, Pauwels, Lays & Lenglet 2006

Other common names:
celebensis: Sulawesi Rat Snake
erythrura: Southern Philippine Rat Snake, Red-tailed Philippine Rat Snake
manillensis: Northern Philippine Rat Snake, Manila Rat Snake
psephenoura: Western Visayan Rat Snake, Grey-tailed Philippine Rat Snake
Remarks: Previously included philippinus. David, Pauwels, Lays & Lenglet (2006) pointed out that the scientific species name is a noun in aposition, not an adjective, for which reason its correct form is erythrura, not erythrurus. Erroneously recorded from Java (Leviton 1979). There is a single record from Banggi Island, north of Sabah, Malaysia, but the record is suspected to represent a misidentified flavolineatus (Stuebing, Inger & Lardner 2014).
Distribution: Indonesia (Banggai Islands, Buton, Sangir Islands, Sulawesi), Philippines (Babuyan Islands [Barit, Calayan, Dalupiri), Batan Islands [Batan], Bohol, Camiguin Sur, Camotes Islands [Pacijan, Ponson, Poro], Catanduanes, Cebu, Dinagat, Guimaras, Leyte, Luzon, Masbate, Mindanao, Mindoro, Negros, Panay, Samar, Siquijor, Sulu Islands [Basilan, Jolo, Siasi], Tablas).


Coelognathus flavolineatus

Yellow-striped Rat Snake, Yellow-striped Trinket Snake, Sunda Rat Snake, Sunda Racer, Black-tailed Trinket Snake, Common Malayan Racer, Black Copper Rat Snake

Gelbstreifen-Kletternatter, Schwarzschwanz-Kletternatter

Gulstribet Rottesnog

1827 Coluber melanurus Boie [not Coluber melanurus Shaw 1802]
Elaphe melanura Barbour 1912
1837 Coluber flavolineata Schlegel
Elaphe flavolineata Smith 1930
Coelognathus flavolineatus Helfenberger 2001

Remarks: Reported from the Nicobar Islands by Daniel & David (1996), but without reference to voucher specimens. Presence in these islands needs confirmation (Vijayakumar & David 2006). According to Iskandar & Colijn (2001), records from Sulawesi and Buton refer to C. erythrurus celebensis, but Lang & Vogel (2005) continued to list the species from Sulawesi (but not Buton).
Distribution: Brunei, Cambodia, India (Andaman Islands), Indonesia (Anambas Islands, Bali, Banggai Islands, Bangka, Belitung, Java, Kalimantan, Nias, Riau Islands, Siberut, Simeulue, Sipura, Sulawesi, Sumatra, Weh), Malaysia (Sabah, Sarawak, West Malaysia [incl. Langkawi Archipelago, Seribuat Archipelago (incl. Tioman)]), Myanmar, Singapore, Thailand, Vietnam.


Coelognathus flavolineatus
© Henrik Bringsøe

Coelognathus helena

Indian Trinket Snake, Common Trinket Snake

Indische Schmucknatter

Indisk Pragtrottesnog

1803 Coluber helena Daudin
Elaphe helena Shaw, Shebbeare & Barker 1939
Coelognathus helena Helfenberger 2001
1849 Cynophis bistrigatus Gray (Smith 1943)
1854 Herpetodryas malabaricus Jerdon (Smith 1943)
1878 Herpetodryas malabaricus carinata Müller (Smith 1943)
1992 Elaphe helena monticollaris Schulz
2016 Coelognathus helena nigriangularis Mohapatra, Schulz, Helfenberger, Hofmann & Dutta

Other common names:
monticollaris: Collared Trinket Snake, Montane Trinket Snake
Distribution: Bangladesh, Bhutan, India (Andhra Pradesh, Assam, Bihar, Chhattisgarh, Delhi, Goa, Gujarat, Haryana, Himachal Pradesh, Jammu & Kashmir, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Nagaland, Odisha, Punjab, Rajasthan, Sikkim, Tamil Nadu, Telangana, Uttar Pradesh, Uttarakhand, West Bengal), Nepal, Sri Lanka.


Coelognathus helena
© Henrik Bringsøe

Coelognathus philippinus

Palawan Rat Snake

Palawan-Rattennatter

Palawan-rottesnog

1909 Elaphe philippina Griffin
Elaphe erythrura philippina Leviton 1979
Coelognathus philippinus Helfenberger 2001
Coelognathus erythrura philippinus David, Pauwels, Lays & Lenglet 2006

Remarks: Previously regarded as a subspecies of erythrura. Helfenberger (2001) revalidated philippinus as a separate species, followed by Wallach, Williams & Boundy (2014), however, not by David, Pauwels, Lays & Lenglet (2006).
Distribution: Philippines (Balabac, Calamian Islands [Busuanga, Calauit, Culion], Palawan, Sulu Islands [Bongao, Sangasanga, Sibutu, Tawi-Tawi]).


Coelognathus radiatus

Copper-headed Rat Snake, Radiated Rat Snake, Copper-headed Trinket Snake, Copperhead Racer

Strahlennatter, Asiatische Kupferkopfnatter

Strålesnog

1827 Coluber radiatus Boie
Compsosoma radiatum Duméril & Bibron 1854
Spilotes radiatus Günther 1858
Elaphis radiatus Jan 1863
Coelognathus radiatus Cochran 1930
Elaphe radiata Pope 1929
1839 Coluber quadrifasciatus Cantor (Smith 1943)
1839 Tropidonotus quinque Cantor (Smith 1943)

Remarks: Present in the past, but not confirmed recently from Singapore (Baker & Lim 2008).
Distribution: Bangladesh, Bhutan, Brunei, Cambodia, China (Fujian, Guangdong [incl. Nan Ao Island], Guangxi, Guizhou, Hong Kong [incl. Shek Kwu Chau Island], Yunnan), India (Andhra Pradesh, Arunachal Pradesh, Assam, Bihar, Chhattisgarh, Himachal Pradesh, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Sikkim, Tripura, Uttarakhand, West Bengal), Indonesia (Bali, Bangka, Java, Kalimantan, Penida, Sumatra), Laos, Malaysia (Sarawak, West Malaysia), Myanmar, Nepal, Singapore, Thailand, Vietnam (incl. Cat Ba Island).


Coelognathus radiatus
© Rune Midtgaard

Coelognathus subradiatus

Lesser Sundas Rat Snake, Indonesian Rat Snake, Indonesian Trinket Snake, Timor Rat Snake

Indonesische Kletternatter

Sunda-rottesnog

1837 Coluber subradiatus Schlegel
Elaphis subradiatus Günther 1858
Elaphe subradiata Barbour 1912
Coelognathus subradiatus Helfenberger 2001
1898 Coluber melanurus timorensis Bethencourt-Ferreira (Iskandar & Colijn 2001)

Remarks: Some authors regard enganensis as a subspecies of subradiatus (e.g., Schulz 1996).
Distribution: East Timor (incl. Atauro Island), Indonesia (Adonara, Alor, Ende, Flores, Komodo, Lembata, Lombok, Rinca, Rote, Semau, Sumba, Sumbawa, West Timor, Wetar).