Reptiles of Palawan [Philippines] Related checklists:  Philippines Bibliography of the reptiles of Palawan Taxonomic history of the reptiles of Palawan Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: April 14th, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Bronchocela (Long-tailed Agamas)   Bronchocela cristatella Malayan Green Crested Agama Gaulke 1996 Hallermann 2005 Manthey 2008 Schoppe & Cervancia 2009 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Taylor 1922 Genus Draco (Gliding Dragons)   Draco palawanensis Palawan Gliding Dragon Manthey 2008 McGuire & Alcala 2000 McGuire & Kiew 2001 Schoppe & Cervancia 2009 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020  Family Dibamidae (Blind Skinks) Genus Dibamus (Oriental Blind Skinks)   Dibamus leucurus White-tailed Blind Skink Alcala 1986   Dibamus novaeguineae Indo-Australian Blind Skink Greer 1985  Family Gekkonidae (Typical Geckos) Genus Cyrtodactylus (Australasian Bent-toed Geckos)   Cyrtodactylus annulatus Mindanao Banded Bent-toed Gecko Alcala 1986 Brown & Alcala 1978 Gaulke 1996   * Cyrtodactylus redimiculus Palawan Bent-toed Gecko Alcala 1986 Brown & Alcala 1978 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Welton, Siler, Linkem, Diesmos & Brown 2010   * Cyrtodactylus tautbatorum Mainit Bent-toed Gecko Welton, Siler, Diesmos & Brown 2009 Welton, Siler, Linkem, Diesmos & Brown 2010 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Brown & Alcala 1978 Schoppe & Cervancia 2009 Genus Gekko (Tokays)   * Gekko athymus Smooth-scaled Narrow-disked Gecko Alcala 1986 Brown & Alcala 1978 Linkem, Siler, Diesmos, Sy & Brown 2010 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Schoppe & Cervancia 2009 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020   Gekko gecko Tokay Brown & Alcala 1978 Schoppe & Cervancia 2009 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020   * Gekko gulat Mount Mantalingajan Flap-legged Gecko Brown, Diesmos, Duya, Garcia & Rico 2010 [as Luperosaurus gulat]   Gekko monarchus Spotted House Gecko Alcala 1986 Brown & Alcala 1978 Linkem, Siler, Diesmos, Sy & Brown 2010 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020   Gekko palawanensis Palawan Narrow-disked Gecko Alcala 1986 Brown & Alcala 1978 Linkem, Siler, Diesmos, Sy & Brown 2010 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus frenatus Asian House Gecko Brown & Alcala 1978 Schoppe & Cervancia 2009 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Genus Hemiphyllodactylus (Slender Geckos)   Hemiphyllodactylus insularis Philippine Slender Gecko Brown & Alcala 1978 [as typus] Zug 2010   Hemiphyllodactylus typus Indo-Pacific Slender Gecko Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Zug 2010 Genus Lepidodactylus (Scaly-toed Geckos)   Lepidodactylus lugubris Common Mourning Gecko Brown & Alcala 1978 Genus Luperosaurus (Flap-legged Geckos)   * Luperosaurus palawanensis Palawan Flap-legged Gecko Alcala 1986 Brown & Alcala 1978 Brown, Supriatna & Ota 2000  Family Scincidae (Skinks) Genus Dasia (Indo-Malayan Tree Skinks)   Dasia griffini Palawan Tree Skink Alcala 1986 Brown & Alcala 1980 Brown, Dimalibot & Rico 2009 Greer 1970 Inger & Brown 1980 Genus Emoia (Emo Skinks)   Emoia atrocostata Pacific Mangrove Skink Brown & Alcala 1980 Brown 1991 Gaulke 1996 Schoppe & Cervancia 2009   Emoia caeruleocauda Pacific Blue-tailed Skink Brown 1991 Genus Eutropis (South Asian Sun Skinks)   Eutropis indeprensa Mindoro Sun Skink Brown & Alcala 1980 Gaulke 1996 Schoppe & Cervancia 2009   Eutropis multifasciata Many-lined Sun Skink Brown & Alcala 1980 O'Shea, Kusuma & Kaiser 2018 Schoppe & Cervancia 2009 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020   * Eutropis sahulinghangganan Palawan Sun Skink Barley, Diesmos, Siler, Martinez & Brown 2020 Genus Insulasaurus (Island Forest Skinks)   * Insulasaurus traanorum Rizal Forest Skink Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Linkem, Diesmos & Brown 2010   * Insulasaurus victoria Victoria Range Forest Skink Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010   * Insulasaurus wrighti Palawan Forest Skink Alcala 1986 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Genus Lamprolepis (Malay Archipelago Tree Skinks)   Lamprolepis smaragdina Emerald Tree Skink Brown & Alcala 1980 Gaulke 1996 Greer 1970 Mertens 1929 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Genus Lipinia (Indo-Pacific Striped Skinks)   Lipinia quadrivittata Black-striped Slender Tree Skink Alcala 1986 Brown & Alcala 1980 Supsup, Redoblado & Guinto 2016 Genus Lygosoma (Southeast Asian Writhing Skinks)   Lygosoma tabonorum Palawan Writhing Skink Heitz, Diesmos, Freitas, Ellsworth, Grismer, Aowphol, Brown & Siler 2016 Siler, Heitz, Davis, Freitas, Aowphol, Termprayoon & Grismer 2018 Genus Parvoscincus (Philippine Dwarf Forest Skinks)   * Parvoscincus palawanensis Palawan Earless Dwarf Forest Skink Alcala 1986 Brown & Alcala 1961 Brown & Alcala 1980 Brown, Linkem, Diesmos, Balete, Duya & Ferner 2010 Ferner, Brown & Greer 1997 Genus Subdoluseps (Asian Agile Skinks)   Subdoluseps bowringii Short-legged Agile Skink Siler, Heitz, Davis, Freitas, Aowphol, Termprayoon & Grismer 2018 Taylor 1963  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus palawanensis Palawan Monitor Brown & Alcala 1970 [as salvator] Gaulke 1996 [as marmoratus] Koch, Auliya, Schmitz, Kuch & Boehme 2007 [as marmoratus] Koch, Gaulke & Boehme 2010 Lisle 1996 [as marmoratus] Schoppe & Cervancia 2009 [as marmoratus] Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Welton, Travers, Siler & Brown 2014

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Acrochordidae (Australasian File Snakes) Genus Acrochordus (Australasian File Snakes)   Acrochordus granulatus Lesser File Snake Leviton, Siler, Weinell & Brown 2018 Wallach, Williams & Boundy 2014  Family Colubridae (Typical Snakes or Colubrids) Genus Ahaetulla (Oriental Whip Snakes)   Ahaetulla prasina Short-nosed Whip Snake Alcala 1986 Haas 1950 Iskandar & Colijn 2001 Leviton 1967 Leviton, Siler, Weinell & Brown 2018 Schoppe & Cervancia 2009 Wallach, Williams & Boundy 2014 Genus Boiga (Australasian Cat Snakes)   Boiga cynodon Dog-toothed Cat Snake Alcala 1986 Ganesh, Mallik, Achyuthan, Shanker & Vogel 2021 Gaulke 1996 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Orlov & Ryabov 2002   Boiga dendrophila Eastern Mangrove Cat Snake Alcala 1986 Brongersma 1934 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Orlov & Ryabov 2002 Schoppe & Cervancia 2009   * Boiga schultzei Palawan Cat Snake Ganesh, Mallik, Achyuthan, Shanker & Vogel 2021 Iskandar & Colijn 2001 Leviton 1968 Leviton, Siler, Weinell & Brown 2018 Orlov & Ryabov 2002 Schoppe & Cervancia 2009 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Genus Calamaria (Oriental Reed Snakes)   * Calamaria palavanensis Palawan Reed Snake Alcala 1986 Inger & Marx 1965 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018   Calamaria virgulata Blunt-tailed Reed Snake Alcala 1986 Inger & Marx 1965 Iskandar & Colijn 2001 Leviton, Siler, Weinell & Brown 2018 Genus Chrysopelea (Flying Tree Snakes)   Chrysopelea paradisi Paradise Flying Tree Snake Iskandar & Colijn 2001 Leviton 1964 Leviton, Siler, Weinell & Brown 2018 Genus Coelognathus (Oriental Rat Snakes)   Coelognathus philippinus Palawan Rat Snake Alcala 1986 Gaulke 1996 [as erythrura] Iskandar & Colijn 2001 Leviton 1979 Leviton, Siler, Weinell & Brown 2018 Schulz 1996 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis levitoni Palawan Stripe-tailed Bronzeback Alcala 1986 [as caudolineatus] Gaulke 1996 [as caudolineatus] Haas 1950 [as caudolineatus] Iskandar & Colijn 2001 [as caudolineatus] Leviton 1968 [as caudolineatus] Leviton, Siler, Weinell & Brown 2018 Rooijen & Vogel 2012   Dendrelaphis marenae Philippine Painted Bronzeback Iskandar & Colijn 2001 [as pictus] Leviton 1968 [as pictus] Leviton, Siler, Weinell & Brown 2018 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Vogel & Rooijen 2008 Genus Dryophiops (Malayan Vine Snakes)   Dryophiops rubescens Keel-bellied Vine Snake Iskandar & Colijn 2001 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Taylor 1965 Genus Gonyosoma (Oriental Green Rat Snakes and Rainbow Tree Snake)   Gonyosoma oxycephalum Red-tailed Green Rat Snake Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Schulz 1996 Genus Liopeltis (Ringneck Snakes)   Liopeltis philippina Culion Ringneck Snake Alcala 1986 Iskandar & Colijn 2001 Lalremsanga, Bal, Vogel & Biakzuala 2022 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Som, Grismer, Wood, Quah, Brown, Diesmos, Weinell & Stuart 2020   Liopeltis tricolor Malayan Ringneck Snake Alcala 1986 Haas 1950 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Genus Lycodon (Oriental Wolf Snakes)   Lycodon capucinus Island Wolf Snake Iskandar & Colijn 2001 Leviton 1965   Lycodon philippinus Philippine Bridle Snake Alcala 1986 [as tristrigatus] Griffin 1909 Iskandar & Colijn 2001 [as tristrigatus] Leviton 1963 [as tristrigatus] Leviton, Siler, Weinell & Brown 2018 Taylor 1922   Lycodon sealei Palawan Wolf Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1955 Leviton 1965 Leviton, Siler, Weinell & Brown 2018 Schoppe & Cervancia 2009 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020   Lycodon subannulatus Malayan Bridle Snake Iskandar & Colijn 2001 Lee, Miller, Connette, Oo, Zug & Mulcahy 2018 Leviton 1963 Genus Oligodon (Kukri Snakes)   Oligodon notospilus Palawan Kukri Snake Alcala 1986 [as vertebralis] Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Genus Opisthotropis (Mountain Stream Keelbacks)   Opisthotropis typica Dwarf Mountain Stream Keelback Alcala 1986 David, Pauwels, Nguyen & Vogel 2015 Iskandar & Colijn 2001 Iskandar & Kamsi 2009 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Schoppe & Cervancia 2009 Genus Ptyas (Giant Rat Snakes and Oriental Green Snakes)   Ptyas carinata Malayan Keeled Rat Snake Alcala 1986 Haas 1950 Iskandar & Colijn 2001 Leviton 1963 Leviton 1983 Leviton, Siler, Weinell & Brown 2018 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Taylor 1965 Genus Rhabdophis (Asian Keelbacks)   Rhabdophis chrysargos Speckled-bellied Keelback Alcala 1986 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Haas 1950 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Schoppe & Cervancia 2009 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Genus Sibynophis (Oriental Black-headed Snakes)   Sibynophis bivittatus Palawan Black-headed Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Calliophis (South Asian Coral Snakes)   Calliophis bilineatus Two-striped Coral Snake Alcala 1986 Brown, Smart, Leviton & Smith 2018 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Iskandar & Colijn 2001 Leviton 1963 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Genus Naja (Typical Cobras)   Naja sumatrana Sunda Spitting Cobra Alcala 1986 [as N. miolepis] Iskandar & Colijn 2001 Leviton 1964 [as Naja naja miolepis] Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Schoppe & Cervancia 2009 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Wuster & Thorpe 1990 [as Naja naja miolepis] Wuster & Thorpe 1991 Genus Ophiophagus (King Cobra)   Ophiophagus hannah King Cobra Alcala 1986 Iskandar & Colijn 2001 Leviton 1964 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Laticauda (Sea Kraits)   Laticauda colubrina Yellow-lipped Sea Krait Heatwole, Busack & Cogger 2005 Iskandar & Colijn 2001   Laticauda semifasciata Erabu Sea Krait Leviton, Siler, Weinell & Brown 2018  Family Homalopsidae (Australasian Mud Snakes) Genus Cerberus (Dog-faced Water Snakes)   Cerberus schneiderii South East Asian Dog-faced Water Snake Iskandar & Colijn 2001 [as C. rynchops] Leviton, Siler, Weinell & Brown 2018 Murphy & Voris 2014 Murphy, Voris & Karns 2012 Genus Gerarda (Glossy Marsh Snake)   Gerarda prevostiana Glossy Marsh Snake Auth, Auffenberg & Dorman 1990 Leviton, Siler, Weinell & Brown 2018  Family Lamprophiidae (House Snakes, etc.) Genus Psammodynastes (Asian Mock Vipers)   Psammodynastes pulverulentus Dusky Mock Viper Iskandar & Colijn 2001 Leviton 1963 Leviton 1983 Leviton, Siler, Weinell & Brown 2018 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020  Family Pareidae (Oriental Slug Eaters) Genus Aplopeltura (Blunt-headed Slug Eater)   Aplopeltura boa Blunt-headed Slug Eater Alcala 1986 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 Poyarkov, Nguyen, Pawangkhanant, Yushchenko, Brakels, Nguyen, Nguyen, Suwannapoom, Orlov & Vogel 2022 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Taylor 1965 Wallach, Williams & Boundy 2014  Family Pythonidae (Pythons) Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Gaulke 1996 Iskandar & Colijn 2001 Leviton 1963 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 Schoppe & Cervancia 2009  Family Viperidae (Vipers) Genus Parias (Malay Archipelago Pitvipers)   Parias flavomaculatus Yellow-spotted Pitviper Leviton, Siler, Weinell & Brown 2018   Parias schultzei Palawan Pitviper Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Iskandar & Colijn 2001 Leviton 1964 Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 Schoppe & Cervancia 2009 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Vogel 2006 Genus Tropidolaemus (Temple Pitvipers)   Tropidolaemus subannulatus North Philippine Temple Pitviper Gaulke 1996 [as wagleri] Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 [as wagleri] Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 [as wagleri] Hoge & Romano Hoge 1981 [as wagleri] Haas 1950 [as wagleri] Iskandar & Colijn 2001 [as wagleri] Leviton 1964 [as wagleri] Leviton, Brown & Siler 2014 Leviton, Siler, Weinell & Brown 2018 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Vogel 2006 Vogel, David, Lutz, Rooijen & Vidal 2007  Family Xenopeltidae (Asian Sunbeam Snakes) Genus Xenopeltis (Asian Sunbeam Snakes)   Xenopeltis unicolor Common Asian Sunbeam Snake Alcala 1986 Iskandar & Colijn 2001 Leviton 1963 Leviton 1983 Leviton, Siler, Weinell & Brown 2018 McDiarmid, Campbell & Toure 1999 Orlov 2000 Taylor 1965

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Sagun, Ramoso & Mejino 2005  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Salinas, Ramoso & Rodriguez 2009  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle David, Pauwels, Lays & Lenglet 2006 Diesmos, Brown, Alcala & Sison 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Schoppe & Cervancia 2009 Schoppe & Das 2011 Genus Cyclemys (Asian Leaf Turtles)   Cyclemys dentata Common Leaf Turtle Alcala 1986 Diesmos, Brown, Alcala & Sison 2008 Fritz & Havas 2006 Fritz, Gaulke & Lehr 1997 Fritz, Guicking, Auer, Sommer, Wink & Hundsdoerfer 2008 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Schoppe & Cervancia 2009 Supsup, Asis, Carestia, Diesmos, Mallari & Brown 2020 Genus Siebenrockiella (Black Marsh Turtle and Philippine Pond Turtle)   Siebenrockiella leytensis Philippine Pond Turtle Diesmos, Brown, Alcala & Sison 2008 Diesmos, Buskirk, Schoppe, Diesmos, Sy & Brown 2012 Diesmos, Gee, Diesmos, Brown, Widman & Dimalibot 2004 Fidenci 2004 Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Schoppe & Cervancia 2009 Timmerman & Auth 1988  Family Trionychidae (Softshell Turtles) Genus Dogania (Malayan Softshell Turtle)   Dogania subplana Malayan Softshell Turtle Diesmos, Brown, Alcala & Sison 2008 Fidenci 2005 Fritz & Havas 2006 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Pelochelys (Indomalayan Giant Softshell Turtles)   Pelochelys cantorii Asian Giant Softshell Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021