Reptiles of Timor [East Timor / Indonesia] Related checklists:  East Timor - West Timor [Indonesia] Bibliography of the reptiles of Timor Taxonomic history of the reptiles of Timor Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   Crocodylus porosus Saltwater Crocodile Auffenberg 1980 Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 McCoy 1980 Mertens 1930 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1915 Wermuth & Mertens 1977

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Draco (Gliding Dragons)   Draco timoriensis Timor Gliding Dragon Inger 1983 [as volans] Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Manthey 2008 McGuire & Kiew 2001 Musters 1983 O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1915 [as walkeri] Sanchez, Carvalho, Kathriner, O'Shea & Kaiser 2012 Wermuth 1967  Family Gekkonidae (Typical Geckos) Genus Cyrtodactylus (Australasian Bent-toed Geckos)   * Cyrtodactylus celatus Timor Bent-toed Gecko Kathriner, Bauer, O'Shea, Sanchez & Kaiser 2014 Mecke, Kieckbusch, Hartmann & Kaiser 2016   * Cyrtodactylus darmandvillei Flores Bent-toed Gecko Youmans & Grismer 2006   * Cyrtodactylus santana Napana Wei Bent-toed Gecko Chan, Grismer, Santana, Pinto, Loke & Conaboy 2023 Genus Gehyra (Four-clawed Geckos)   Gehyra mutilata Common Four-clawed Gecko Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 [as "cf. mutilata"] Kaiser, Sanchez, Heacox, Kathriner, Ribeiro, Soares, Araujo, Mecke & O'Shea 2013 Mertens 1930 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1915 Smith 1927 Genus Gekko (Tokays)   Gekko gecko Tokay Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Kaiser, Sanchez, Heacox, Kathriner, Ribeiro, Soares, Araujo, Mecke & O'Shea 2013 Mertens 1930 O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Reilly, Stubbs, Karin, Arida, Iskandar & McGuire 2019 Roesler, Bauer, Heinicke, Greenbaum, Jackman, Nguyen & Ziegler 2011 Rooij 1915 [as verticillatus] Sanchez, Carvalho, Kathriner, O'Shea & Kaiser 2012   Gekko vittatus White-striped Gecko Rooij 1915 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus frenatus Asian House Gecko Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Kaiser, Sanchez, Heacox, Kathriner, Ribeiro, Soares, Araujo, Mecke & O'Shea 2013 O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1915 Sanchez, Carvalho, Kathriner, O'Shea & Kaiser 2012   Hemidactylus garnotii Fox Gecko O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 [as "cf. garnotii"]   Hemidactylus murrayi Pimpri Half-toed Gecko Chaitanya, Agarwal, Lajmi & Khandekar 2019   Hemidactylus platyurus Flat-tailed Gecko Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Sanchez, Carvalho, Kathriner, O'Shea & Kaiser 2012   Hemidactylus tenkatei Rote Half-toed Gecko Kathriner, O'Shea & Kaiser 2014 Mahony 2011 Mertens 1930 [as brookii] O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 [as "cf. tenkatei"] O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Sanchez, Carvalho, Kathriner, O'Shea & Kaiser 2012 [as "cf. tenkatei"] Genus Hemiphyllodactylus (Slender Geckos)   Hemiphyllodactylus typus Indo-Pacific Slender Gecko O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 [as "cf. typus"]  Family Scincidae (Skinks) Genus Carlia (Australian-Pacific Rainbow Skinks)   Carlia peronii Timor Rainbow Skink Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Zug 2010   * Carlia spinauris Lelogama Rainbow Skink Brongersma 1942 Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Smith 1927 Zug 2010 Genus Cryptoblepharus (Indo-Pacific Snake-eyed Skinks)   Cryptoblepharus leschenault East Indonesian Snake-eyed Skink Brongersma 1942 Horner 2007 Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Kaiser, Sanchez, Heacox, Kathriner, Ribeiro, Soares, Araujo, Mecke & O'Shea 2013 Mertens 1964 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Genus Emoia (Emo Skinks)   Emoia cyanura White-bellied Copper-striped Skink Rooij 1915 Genus Eremiascincus (Glossy Night Skinks and Sand Swimmers)   * Eremiascincus antoniorum Lelogama Glossy Night Skink Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Mecke, Doughty & Donnellan 2009 Smith 1927   * Eremiascincus timorensis Timor Glossy Night Skink Greer 1990 Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Mecke, Doughty & Donnellan 2009 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 [as "cf. timorensis"] Genus Eutropis (South Asian Sun Skinks)   Eutropis multifasciata Many-lined Sun Skink Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 [as "cf. multifasciata"] Kaiser, Sanchez, Heacox, Kathriner, Ribeiro, Soares, Araujo, Mecke & O'Shea 2013 [as "cf. multifasciata"] Mertens 1930 O'Shea, Kusuma & Kaiser 2018 O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 [as "cf. multifasciata"] O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 [as "cf. multifasciata"] Reilly, Stubbs, Karin, Arida, Iskandar & McGuire 2019 Rooij 1915 Sanchez, Carvalho, Kathriner, O'Shea & Kaiser 2012 [as "cf. multifasciata"] Schmitt, How, Hisheh, Goldberg & Maryanto 2000 Genus Lamprolepis (Malay Archipelago Tree Skinks)   Lamprolepis smaragdina Emerald Tree Skink Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 [as "cf. smaragdina"] Kaiser, Sanchez, Heacox, Kathriner, Ribeiro, Soares, Araujo, Mecke & O'Shea 2013 [as "cf. smaragdina"] Mertens 1929 Mertens 1930 O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 [as "cf. smaragdina"] O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 [as "cf. smaragdina elberti"] Rooij 1915 Sanchez, Carvalho, Kathriner, O'Shea & Kaiser 2012 [as "cf. smaragdina"] Schmitt, How, Hisheh, Goldberg & Maryanto 2000 Genus Sphenomorphus (Indo-Pacific Forest Skinks)   Sphenomorphus melanopogon Black-bearded Forest Skink Auffenberg 1980 [as florense] Brongersma 1942 [as barbouri] Mertens 1930 [as florense] O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 [as melanopogon] Rooij 1915 [as florense] Shea 2012 Smith 1927 [as florense] Genus Tytthoscincus (Dwarf Leaf-litter Skinks)   Tytthoscincus temminckii Sunda Leaf-litter Skink Mertens 1930  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus salvator Asian Water Monitor O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 [as "cf. salvator"]   * Varanus timorensis Timor Spotted Tree Monitor Mertens 1963 O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1915 Smith 1927

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Acrochordidae (Australasian File Snakes) Genus Acrochordus (Australasian File Snakes)   Acrochordus granulatus Lesser File Snake Haas 1950 Iskandar & Colijn 2001 Lang 2011 McDiarmid, Campbell & Toure 1999 Mertens 1930 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1917 Wallach, Williams & Boundy 2014  Family Colubridae (Typical Snakes or Colubrids) Genus Coelognathus (Oriental Rat Snakes)   Coelognathus subradiatus Lesser Sundas Rat Snake Auffenberg 1980 Brongersma 1933 Haas 1950 Iskandar & Colijn 2001 Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Kaiser, Sanchez, Heacox, Kathriner, Ribeiro, Soares, Araujo, Mecke & O'Shea 2013 Lang 2011 Mertens 1930 O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1917 Schulz 1996 Smith 1927 Genus Dendrelaphis (Bronzebacks)   Dendrelaphis inornatus Lesser Sundas Painted Bronzeback Auffenberg 1980 [as pictus] Haas 1950 Iskandar & Colijn 2001 Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Lang 2011 Mertens 1930 [as pictus] O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1917 [as pictus] Smith 1927 Genus Lycodon (Oriental Wolf Snakes)   Lycodon capucinus Island Wolf Snake Haas 1950 Iskandar & Colijn 2001 Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Kaiser, Sanchez, Heacox, Kathriner, Ribeiro, Soares, Araujo, Mecke & O'Shea 2013 Lang 2011 Mertens 1930 O'Shea, Kusuma & Kaiser 2018 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Reilly, Stubbs, Karin, Arida, Iskandar & McGuire 2019 Rooij 1917 [as aulicus] Smith 1927 [as L. aulicus] Vogel & Harikrishnan 2013   Lycodon subcinctus Malayan Banded Wolf Snake Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Lang 2011 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Genus Stegonotus (Indo-Australian Ground Snakes)   * Stegonotus nancuro Timor Ground Snake Kaiser, Kaiser, Mecke & O'Shea 2021  Family Cylindrophiidae (Asian Pipe Snakes) Genus Cylindrophis (Oriental Pipe Snakes)   Cylindrophis boulengeri Wetar Pipe Snake Bernstein, Bauer, McGuire, Arida, Kaiser, Kieckbusch & Mecke 2020 Lang 2011 Lang 2013 McDiarmid, Campbell & Toure 1999 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis atriceps Black-headed Sea Snake Iskandar & Colijn 2001 Genus Laticauda (Sea Kraits)   Laticauda colubrina Yellow-lipped Sea Krait Heatwole, Busack & Cogger 2005 Haas 1950 Kharin, Roedel & Hallermann 2010 Mertens 1930 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1917 Sanchez, Carvalho, Kathriner, O'Shea & Kaiser 2012   Laticauda laticaudata Blue-lipped Sea Krait Haas 1950 Kharin, Roedel & Hallermann 2010 Mertens 1930 Rooij 1917  Family Homalopsidae (Australasian Mud Snakes) Genus Cantoria (Yellow-banded Mangrove Snake)   Cantoria violacea Yellow-banded Mangrove Snake Gyi 1970 Lang 2011 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1917 Genus Cerberus (Dog-faced Water Snakes)   Cerberus schneiderii South East Asian Dog-faced Water Snake Iskandar & Colijn 2001 [as C. rynchops] Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 [as C. rynchops] Lang 2011 [as C. rynchops] Mertens 1930 [as C. rynchops] Murphy & Voris 2014 Murphy, Voris & Karns 2012 O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 [as C. rynchops] O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 [as C. rynchops] Rooij 1917 [as C. rynchops] Genus Fordonia (White-bellied Mangrove Snake)   Fordonia leucobalia White-bellied Mangrove Snake Gyi 1970 Iskandar & Colijn 2001 Lang 2011 Lang 2013 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1917  Family Pythonidae (Pythons) Genus Liasis (Water Pythons and Australian Olive Python)   Liasis mackloti Timor Water Python Iskandar & Colijn 2001 Lang 2011 McDiarmid, Campbell & Toure 1999 O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1917 Stimson 1969 Genus Malayopython (Malayan Pythons)   Malayopython reticulatus Reticulated Python Iskandar & Colijn 2001 Lang 2011 McDiarmid, Campbell & Toure 1999 Mertens 1930 O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1917 Sanchez, Carvalho, Kathriner, O'Shea & Kaiser 2012 Stimson 1969  Family Typhlopidae (Typical Blind Snakes) Genus Sundatyphlops (Lesser Sunda Blind Snakes)   Sundatyphlops polygrammicus Many-lined Blind Snake Hahn 1980 Iskandar & Colijn 2001 Lang 2011 McDiarmid, Campbell & Toure 1999 McDowell 1974 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1917 Smith 1927 Wallach, Williams & Boundy 2014 Genus Virgotyphlops (Brahminy Blind Snake)   Virgotyphlops braminus Brahminy Blind Snake Hahn 1980 Iskandar & Colijn 2001 Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Kramer 1977 Lang 2011 McDiarmid, Campbell & Toure 1999 Mertens 1930 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rooij 1917 Wallach 2009 Wallach 2020  Family Viperidae (Vipers) Genus Trimeresurus (Oriental Pitvipers)   Trimeresurus insularis Lesser Sundas White-lipped Pitviper Auffenberg 1980 [as albolabris] Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Gumprecht, Tillack, Orlov, Captain & Ryabov 2004 Haas 1950 [as albolabris] Iskandar & Colijn 2001 Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Kaiser, Sanchez, Heacox, Kathriner, Ribeiro, Soares, Araujo, Mecke & O'Shea 2013 Kramer 1977 Lang 2011 O'Shea, Sanchez, Heacox, Kathriner, Carvalho, Varela-Ribeiro, Afrianio-Soares, Araujo & Kaiser 2012 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Regenass & Kramer 1981 Reilly, Stubbs, Karin, Arida, Iskandar & McGuire 2019 Smith 1943 [as albolabris]

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Chelidae (Austro-American Side-necked Turtles) Genus Chelodina (Indo-Australian Snake-necked or Long-necked Turtles)   Chelodina mccordi Rote Island Snake-necked Turtle Georges & Thomson 2010 Kaiser, Carvalho, Ceballos, Freed, Heacox, Lester, Richards, Trainor, Sanchez & O'Shea 2011 Kuchling, Rhodin, Ibarrondo & Trainor 2007 McCord, Joseph-Ouni & Hagen 2007 O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle O'Shea, Sanchez, Kathriner, Mecke, Carvalho, Ribeiro, Soares, Araujo & Kaiser 2015 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Natator (Flatback Sea Turtle)   Natator depressus Flatback Sea Turtle Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021  Family Geoemydidae (Eastern Pond Turtles) Genus Cuora (Asian Box Turtles)   Cuora amboinensis Southeast Asian Box Turtle Iverson 1992 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Schoppe & Das 2011