Reptiles of Western Sahara  [disputed territory: claimed by Morocco and the Sahrawi Arab Democratic Republic]   Bibliography of the reptiles of Western Sahara Taxonomic history of the reptiles of Western Sahara Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Crocodilians (Crocodylia)
	= endemicEX = extinct or assumed extinct Species References  Family Crocodylidae (Crocodiles) Genus Crocodylus (Typical Crocodiles)   EX Crocodylus suchus West African Crocodile Geniez, Mateo, Geniez & Pether 2004 Lescure 2014

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama bibronii Atlas Agama Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012 Genus Trapelus (Afro-Asian Ground Agamas)   Trapelus boehmei Moroccan Plain Agama Berre 1989 [as mutabilis] Geniez, Mateo, Geniez & Pether 2004 [as mutabilis] Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco & Jeremchenko 2008 [as mutabilis] Wagner, Melville, Wilms & Schmitz 2011 [as mutabilis (s.s.)] Genus Uromastyx (Western Spiny-tailed Agamas)   Uromastyx dispar South Saharan Spiny-tailed Agama Bons & Geniez 1996 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Trape, Trape & Chirio 2012 Wilms & Boehme 2001 Wilms 2001 Wilms, Boehme, Wagner, Lutzmann & Schmitz 2009   * Uromastyx occidentalis Giant Spiny-tailed Agama Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Mateo, Geniez, Lopez-Jurado & Bons 1998 Trape, Trape & Chirio 2012 Wilms 2001  Family Chamaeleonidae (Chameleons) Genus Chamaeleo (Typical Chameleons)   Chamaeleo chamaeleon Mediterranean Chameleon Geniez, Mateo, Geniez & Pether 2004 Glaw 2015 Klaver & Boehme 1997 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Trape, Trape & Chirio 2012  Family Gekkonidae (Typical Geckos) Genus Hemidactylus (Half-toed Geckos)   Hemidactylus angulatus West African House Gecko Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Genus Stenodactylus (Afro-Asian Thin-toed Geckos)   Stenodactylus mauritanicus Mauritanian Thin-toed Gecko Arnold 1980 [as sthenodactylus] Geniez, Mateo, Geniez & Pether 2004 Loveridge 1947 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco & Jeremchenko 2008 [as sthenodactylus]   Stenodactylus petrii North African Thin-toed Gecko Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Trape, Trape & Chirio 2012   Stenodactylus sthenodactylus Elegant Thin-toed Gecko Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Genus Tropiocolotes (Afro-Asian Desert Dwarf Geckos)   Tropiocolotes algericus Algerian Dwarf Gecko Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Ribeiro Junior, Tamar, Maza, Flecks, Wagner, Shacham, Calvo, Geniez, Crochet, Koch & Meiri 2022 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012 Wilms, Shobrak & Wagner 2010   Tropiocolotes tripolitanus Tripoli Dwarf Gecko Berre 1989 Geniez, Mateo, Geniez & Pether 2004 Loveridge 1947 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Minton, Anderson & Anderson 1970 Ribeiro Junior, Tamar, Maza, Flecks, Wagner, Shacham, Calvo, Geniez, Crochet, Koch & Meiri 2022 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Szczerbak & Golubev 1996 Trape, Trape & Chirio 2012 Wermuth 1965  Family Lacertidae (Typical Lizards) Genus Acanthodactylus (Fringe-toed Lizards)   Acanthodactylus aureus Golden Fringe-toed Lizard Arnold 1983 Brito, Rebelo, Crochet & Geniez 2008 Crochet, Geniez & Ineich 2003 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Salvador 1982 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Acanthodactylus boskianus Afro-Arabian Fringe-toed Lizard Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Salvador 1982 Trape, Trape & Chirio 2012   Acanthodactylus busacki South Moroccan Fringe-toed Lizard Brito, Rebelo, Crochet & Geniez 2008 Crochet, Leblois & Renoult 2015 [as "cf. busacki"] Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Salvador 1982 Sindaco & Jeremchenko 2008 Tamar, Geniez, Brito & Crochet 2017 Trape, Trape & Chirio 2012   Acanthodactylus dumerilii Lesser Fringe-toed Lizard Crochet, Geniez & Ineich 2003 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Salvador 1982 [as dumerili X exiguus] Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008 Genus Mesalina (Afro-Asian Desert Lizards)   Mesalina adrarensis Mauritanian Desert Lizard Pizzigalli, Crochet, Geniez, Martinez-Freiria, Velo-Anton & Brito 2021   Mesalina guttulata Common Small-spotted Desert Lizard Berre 1989 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019   Mesalina pasteuri Sahara Desert Lizard Brito, Rebelo, Crochet & Geniez 2008 Geniez & Geniez 1995 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco & Jeremchenko 2008   Mesalina rubropunctata Red-spotted Desert Lizard Berre 1989 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Mesalina simoni Moroccan Small-spotted Desert Lizard Pizzigalli, Crochet, Geniez, Martinez-Freiria, Velo-Anton & Brito 2021 Genus Timon (Ocellated Lizards)   Timon tangitanus Moroccan Ocellated Lizard Berre 1989 [as lepidus] Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco & Jeremchenko 2008  Family Phyllodactylidae (Leaf-toed Geckos) Genus Tarentola (Wall Geckos)   Tarentola annularis Egyptian White-spotted Gecko Berre 1989 Geniez, Mateo, Geniez & Pether 2004 Joger 1984 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Schleich, Kastle & Kabisch 1996 Sindaco & Jeremchenko 2008   Tarentola boehmei Moroccan Wall Gecko Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019   Tarentola chazaliae Helmeted Gecko Berre 1989 Geniez, Mateo, Geniez & Pether 2004 Loveridge 1947 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Wermuth 1965   Tarentola hoggarensis Hoggar Wall Gecko Berre 1989 Geniez, Mateo, Geniez & Pether 2004 Joger 1984 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Tarentola mauritanica Moorish Wall Gecko Geniez, Escatllar, Crochet, Mateo & Bons 1999 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Tarentola panousei Dra Wall Gecko Koppetsch & Boehme 2022   Tarentola parvicarinata Small-keeled Gecko Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Trape 2012  Family Scincidae (Skinks) Genus Chalcides (Cylindrical Skinks)   Chalcides delislei Central Saharan Three-fingered Skink Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco & Jeremchenko 2008 Trape, Trape & Chirio 2012   Chalcides ocellatus Mediterranean Ocellated Skink Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019   Chalcides polylepis Many-scaled Cylindrical Skink Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019   Chalcides sphenopsiformis Senegal Sand Skink Carranza, Arnold, Geniez, Roca & Mateo 2008 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco & Jeremchenko 2008 Genus Scincus (Sand Fish)   Scincus albifasciatus White-banded Sand Fish Arnold & Leviton 1977 Berre 1989 [as scincus] Geniez, Mateo, Geniez & Pether 2004 Hartmann 1989 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco & Jeremchenko 2008  Family Sphaerodactylidae (Least Geckos, etc.) Genus Quedenfeldtia (Moroccan Day Geckos)   Quedenfeldtia moerens Southern Moroccan Day Gecko Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco & Jeremchenko 2008 Genus Saurodactylus (Lizard-toed Geckos)   Saurodactylus harrisii Guelmime Lizard-toed Gecko Berre 1989 [as S. mauritanicus] Geniez, Mateo, Geniez & Pether 2004 [as S. brosseti] Javanmardi, Vogler & Joger 2019 Trape, Trape & Chirio 2012 [as S. brosseti]  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus griseus Desert Monitor Bayless 2002 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Mertens 1963

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Dasypeltis (African Egg Eaters)   Dasypeltis sahelensis Sahel Egg Eater Geniez, Mateo, Geniez & Pether 2004 [as scabra] Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco, Venchi & Grieco 2013 Venchi & Sindaco 2007 [as scabra] Genus Hemorrhois (Palearctic Whip Snakes)   Hemorrhois algirus Algerian Whip Snake Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Papenfuss 1969 Sindaco, Venchi & Grieco 2013 Villiers 1975 Genus Lytorhynchus (Awl-headed Snakes)   Lytorhynchus diadema Crowned Awl-headed Snake Berre 1989 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Mediani, Fahd, Chevalier & Brito 2015 Genus Macroprotodon (False Smooth Snakes)   Macroprotodon cucullatus Eastern False Smooth Snake Busack & McCoy 1990 [as brevis] Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 [as brevis] Wade 1988 Genus Spalerosophis (Diadem Snakes)   Spalerosophis diadema Common Diadem Snake Berre 1989 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco, Venchi & Grieco 2013 Venchi & Sindaco 2007   Spalerosophis dolichospilus Mograbin Diadem Snake Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus tripolitanus North African Cat Snake Crochet, Rasmussen, Wilms, Geniez, Trape & Boehme 2008 Geniez, Mateo, Geniez & Pether 2004 [as guidimakaensis] Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco, Venchi & Grieco 2013 Venchi & Sindaco 2007 Wilms, Wagner & Spawls 2021  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Naja (Typical Cobras)   Naja haje Egyptian Cobra Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco, Venchi & Grieco 2013 Spawls & Branch 1995 Trape, Chirio, Broadley & Wuster 2009  Family Lamprophiidae (House Snakes, etc.) Genus Boaedon (Afro-Arabian House Snakes)   Boaedon fuliginosus Common African House Snake Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco, Venchi & Grieco 2013 Genus Malpolon (Montpellier Snakes)   Malpolon moilensis Moila Snake Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019   Malpolon monspessulanus Western Montpellier Snake Berre 1989 Geniez, Cluchier & Haan 2006 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco, Venchi & Grieco 2013 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis schokari Arabian Sand Snake Berre 1989 Brito, Rebelo, Crochet & Geniez 2008 Geniez, Mateo, Geniez & Pether 2004 Kramer & Schnurrenberger 1963 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019  Family Leptotyphlopidae (Thread Snakes) Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis algeriensis Algerian Thread Snake Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco, Venchi & Grieco 2013 Venchi & Sindaco 2007  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Crochet, Leblois & Renoult 2015 Geniez & Geniez 1995 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Sindaco, Venchi & Grieco 2013 Genus Cerastes (Horned Vipers)   Cerastes cerastes Sahara Horned Viper Berre 1989 Crochet, Leblois & Renoult 2015 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 McDiarmid, Campbell & Toure 1999 Sindaco, Venchi & Grieco 2013 Spawls & Branch 1995   Cerastes vipera Sahara Sand Viper Crochet, Leblois & Renoult 2015 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 McDiarmid, Campbell & Toure 1999 Sindaco, Venchi & Grieco 2013 Spawls & Branch 1995 Villiers 1975 Genus Daboia (Daboias)   Daboia mauritanica Moorish Viper Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 Genus Echis (Carpet Vipers)   Echis leucogaster White-bellied Carpet Viper Berre 1989 Geniez, Mateo, Geniez & Pether 2004 Martinez del Marmol, Harris, Geniez, Pous & Salvi 2019 McDiarmid, Campbell & Toure 1999 Spawls & Branch 1995

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Brongersma in Bjorndal (ed.) 1982 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Brongersma in Bjorndal (ed.) 1982 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle King & Burke 1989  Family Testudinidae (Tortoises) Genus Centrochelys (African Spurred Tortoise)   EX Centrochelys sulcata African Spurred Tortoise Geniez, Mateo, Geniez & Pether 2004