Reptiles of Northern Cape [South Africa] Related checklists:  South Africa Bibliography of the reptiles of Northern Cape Taxonomic history of the reptiles of Northern Cape Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Amphisbaenidae (Typical Worm Lizards) Genus Dalophia (Blunt-tailed Worm Lizards)   Dalophia pistillum Zambezi Blunt-tailed Worm Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Pietersen & Measey 2010 Branch 1998 Genus Monopeltis (Spade-snouted Worm Lizards)   Monopeltis capensis Cape Spade-snouted Worm Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1997 Broadley, Gans & Visser 1976 [as capensis "Group A"] Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011   Monopeltis infuscata Dusky Spade-snouted Worm Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Pietersen & Measey 2010 Branch 1998 Broadley 1997 Broadley, Gans & Visser 1976 [as capensis "Group B"] Gans 2005   Monopeltis leonhardi Kalahari Spade-snouted Worm Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley, Gans & Visser 1976 Fitzsimons & Brain 1958   Monopeltis mauricei Botswana Spade-snouted Worm Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Pietersen & Measey 2010 Broadley, Gans & Visser 1976 Gans 2005   Monopeltis sphenorhynchus Slender Spade-snouted Worm Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Pietersen & Measey 2010 Fitzsimons & Brain 1958 Genus Zygaspis (Purple Round-headed Worm Lizards)   Zygaspis quadrifrons Kalahari Round-headed Worm Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley & Broadley 1997 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Agama (Typical Agamas)   Agama aculeata Kalahari Spiny Agama Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Baard, Villiers & Villiers 1988 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Fitzsimons & Brain 1958   Agama anchietae Western Rock Agama Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1989 Branch 1998 Baard, Villiers & Villiers 1988 Fitzsimons & Brain 1958   Agama atra Southern Rock Agama Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Baard, Villiers & Villiers 1988 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011   Agama hispida Southern Spiny Agama Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Cornelius, Marais & Cunningham 2012   Agama knobeli Namibian Black Agama Branch 1998  Family Chamaeleonidae (Chameleons) Genus Bradypodion (Southern African Dwarf Chameleons)   Bradypodion gutturale Little Karroo Dwarf Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014   Bradypodion occidentale Namaqua Dwarf Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch, Tolley & Tilbury 2006 Glaw 2015 Tilbury 2010   Bradypodion ventrale Eastern Cape Dwarf Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Tilbury 2010 Visser 1979 Genus Chamaeleo (Typical Chameleons)   Chamaeleo dilepis Flap-necked Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Tilbury 2010   Chamaeleo namaquensis Namaqua Chameleon Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1989 Tilbury 2010  Family Cordylidae (Girdled Lizards) Genus Cordylus (Girdled Lizards)   * Cordylus cloetei Fraserburg Girdled Lizard Branch 1998 Mouton & Wyk 1994   * Cordylus imkeae Rooiberg Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Mouton & Wyk 1994   Cordylus macropholis Large-scaled Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998   Cordylus mclachlani Ceres Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014   Cordylus minor Dwarf Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Genus Hemicordylus (Western Crag Lizards)   Hemicordylus capensis Graceful Crag Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Genus Karusasaurus (Karoo Girdled Lizards)   Karusasaurus polyzonus Southern Karoo Girdled Lizard Bauer & Branch 2001 Branch 1981 Branch 1998 Baard, Villiers & Villiers 1988 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Genus Namazonurus (Namaqua Girdled Lizards)   * Namazonurus lawrenci Richtersveld Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1989 Branch 1998 Visser 1979   Namazonurus peersi Garies Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Genus Ouroborus (Armadillo Girdled Lizard)   Ouroborus cataphractus Armadillo Girdled Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Genus Platysaurus (Flat Lizards)   Platysaurus attenboroughi Karasburg Flat Lizard Whiting, Branch, Pepper & Keogh 2015   * Platysaurus broadleyi Augrabies Flat Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Whiting 1997 Branch 1998 Whiting, Branch, Pepper & Keogh 2015   * Platysaurus capensis Cape Flat Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Broadley 1978 Visser 1979 Whiting, Branch, Pepper & Keogh 2015 Genus Pseudocordylus (Eastern Crag Lizards)   Pseudocordylus melanotus Common Crag Lizard Branch 1981   Pseudocordylus microlepidotus Cape Crag Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998  Family Gekkonidae (Typical Geckos) Genus Afroedura (Flat Geckos)   Afroedura karroica Karoo Flat Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Jacobsen, Kuhn, Jackman & Bauer 2014   * Afroedura namaquensis Namaqua Flat Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Fitzsimons 1938 Huyssteen & Petford 2021 Jacobsen, Kuhn, Jackman & Bauer 2014 Visser 1979 Genus Chondrodactylus (Giant Ground Gecko and Thick-toed Geckos)   Chondrodactylus angulifer Giant Ground Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1989 Baard, Villiers & Villiers 1988 Fitzsimons & Brain 1958 Heinz, Brennan, Jackman & Bauer 2021   Chondrodactylus bibronii Tubercled Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Bauer & Lamb 2005 Branch 1981 Branch 1998 Baard, Villiers & Villiers 1988 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Heinz, Brennan, Jackman & Bauer 2021   Chondrodactylus laevigatus Karas Thick-toed Gecko Heinz, Brennan, Jackman & Bauer 2021 Marques, Ceriaco, Blackburn & Bauer 2018 Genus Goggia (South African Dwarf Leaf-toed Geckos)   Goggia gemmula Richtersveld Dwarf Leaf-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Bauer, Good & Branch 1997 Branch 1998 Good, Bauer & Branch 1996 Heinicke, Turk & Bauer 2017   Goggia hexapora Cedarberg Dwarf Leaf-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Good & Branch 1997 Branch, Bauer & Good 1995 Heinicke, Turk & Bauer 2017   Goggia lineata Striped Dwarf Leaf-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Branch, Bauer & Good 1995 Good, Bauer & Branch 1996 Heinicke, Turk & Bauer 2017   Goggia matzikamaensis Matzikama Dwarf Leaf-toed Gecko Heinicke, Turk & Bauer 2017   * Goggia rupicola Namaqua Dwarf Leaf-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Good & Branch 1997 Branch 1981 Branch 1998 Branch, Bauer & Good 1995 Good, Bauer & Branch 1996 Heinicke, Turk & Bauer 2017 Genus Lygodactylus (African-Neotropical Dwarf Geckos)   Lygodactylus bradfieldi Namibian Dwarf Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Jacobsen 2011 Genus Pachydactylus (Typical Thick-toed Geckos)   * Pachydactylus amoenus Kamaggas Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Heinicke, Jackman & Branch 2011 Branch 1981 Wermuth 1965   Pachydactylus atorquatus Augrabies Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 [as goodi] Bauer, Barts & Hulbert 2006 Bauer, Lamb & Branch 2006 [as goodi]   Pachydactylus austeni Port Nolloth Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Lamb 2005 Branch 1981 Branch 1998   Pachydactylus barnardi Garies Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Bauer & Lamb 2002 Bauer & Lamb 2005 Branch 1981 Branch 1998 Lamb & Bauer 2000   Pachydactylus capensis Common Cape Thick-toed Gecko Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Lamb 2002 Bauer & Lamb 2005 Branch 1998 Baard, Villiers & Villiers 1988 Fitzsimons & Brain 1958   Pachydactylus carinatus Richtersveld Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Lamb & Branch 2006   Pachydactylus formosus Karoo Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Lamb & Bauer 2000   Pachydactylus geitje Ocellated Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014   Pachydactylus haackei Great Namaqualand Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Bauer & Lamb 2005 Branch 1998 Branch, Bauer & Good 1996   Pachydactylus kladaroderma Thin-skinned Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014   Pachydactylus labialis Western Cape Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Bauer & Lamb 2002 Bauer & Lamb 2005 Branch 1981 Branch 1998   Pachydactylus latirostris Broad-nosed Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Heinicke, Jackman & Branch 2011 Branch 1981 Branch 1998 Baard, Villiers & Villiers 1988   * Pachydactylus macrolepis Large-scaled Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Heinicke, Jackman & Branch 2011 Branch 1981   Pachydactylus maculatus Spotted Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014   Pachydactylus mariquensis Marico Thick-toed Gecko Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Bauer, Heinicke, Jackman & Branch 2011 Branch 1998   Pachydactylus monicae Sendelingsdrif Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Lamb & Branch 2006   Pachydactylus montanus Karas Mountains Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 [as P. cf. serval] Bauer, Lamb & Branch 2006 Branch 1981 [as serval onscepensis]   Pachydactylus namaquensis Namaqua Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Lamb 2002 Bauer & Lamb 2005 Branch 1981 Branch 1998 Branch, Bauer & Good 1996   Pachydactylus oculatus Golden Spotted Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Lamb 2005 Branch 1981   Pachydactylus punctatus Speckled Thick-toed Gecko Bauer & Branch 1995 Bauer & Branch 2001 Branch 1981 [as amoenoides]   Pachydactylus purcelli Karoo Spotted Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Lamb & Branch 2006 Branch 1989 Branch 1998 Baard, Villiers & Villiers 1988   Pachydactylus rangei Namib Web-footed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998   Pachydactylus rugosus Rough Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Bauer & Lamb 2002 Bauer & Lamb 2005 Branch 1981 Branch 1989 Branch 1998 Fitzsimons & Brain 1958 Lamb & Bauer 2000   Pachydactylus visseri Ai-Ais Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer, Lamb & Branch 2006   Pachydactylus wahlbergii Kalahari Ground Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Fitzsimons & Brain 1958   Pachydactylus weberi Klipfontein Thick-toed Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Bauer & Lamb 2002 Bauer & Lamb 2005 Bauer, Lamb & Branch 2006 Branch 1981 Genus Ptenopus (African Barking Geckos)   Ptenopus garrulus Common Barking Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Fitzsimons & Brain 1958 Genus Rhoptropella (Namaqua Day Gecko)   * Rhoptropella ocellata Namaqua Day Gecko Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Hallmann, Kruger & Trautmann 1997 Visser 1979  Family Gerrhosauridae (Plated Lizards) Genus Cordylosaurus (Dwarf Plated Lizard)   Cordylosaurus subtessellatus Dwarf Plated Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Bauer & Branch 2001 Branch 1981 Branch 1998 Genus Gerrhosaurus (Typical Plated Lizards)   Gerrhosaurus flavigularis Yellow-throated Plated Lizard Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011   Gerrhosaurus typicus Namaqua Plated Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Tolley, Edwards, Davids, Silva & Branch 2013 Branch 1981 Branch 1998 Genus Tetradactylus (Plated Snake Lizards)   Tetradactylus tetradactylus Four-toed Plated Snake Lizard Bates 2011 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014  Family Lacertidae (Typical Lizards) Genus Heliobolus (Bushveld Lizards)   Heliobolus lugubris Southern African Bushveld Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Fitzsimons & Brain 1958 Marques, Ceriaco, Heinicke, Chehouri, Conradie, Tolley & Bauer 2022 Genus Meroles (Southwest African Desert Lizards)   Meroles ctenodactylus Giant Desert Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Lamb & Bauer 2003   Meroles cuneirostris Wedge-snouted Desert Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Bischoff 1991 Branch 1981 Branch 1998 Lamb & Bauer 2003   Meroles knoxii Round-snouted Desert Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Lamb & Bauer 2003 Plessis & Mouton 2011   Meroles squamulosus Rough-scaled Desert Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011   Meroles suborbitalis Spotted Desert Lizard Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1989 Branch 1998 Baard, Villiers & Villiers 1988 Fitzsimons & Brain 1958 Lamb & Bauer 2003 Genus Nucras (Sandveld Lizards)   Nucras holubi Southern Sandveld Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011   Nucras intertexta Spotted Sandveld Lizard Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Broadley 1972 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Fitzsimons & Brain 1958   Nucras livida Karoo Sandveld Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Conradie, Busschau & Jordaan 2020 Toit & Albas 2003   Nucras tessellata Western Sandveld Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Broadley 1972 Baard, Villiers & Villiers 1988 Fitzsimons & Brain 1958 Oosthuizen 2018 [as N. livida] Plessis & Mouton 2011 Genus Pedioplanis (African Sand Lizards)   Pedioplanis burchelli Southern Sand Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014   Pedioplanis inornata Plain Sand Lizard Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 [as undata] Branch 1989 Branch 1998 Baard, Villiers & Villiers 1988 [as undata]   Pedioplanis laticeps Cape Sand Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Baard, Villiers & Villiers 1988 Kirchhof, Penner, Roedel & Muller 2017 Plessis & Mouton 2011   Pedioplanis lineoocellata Spotted Sand Lizard Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Baard, Villiers & Villiers 1988 Fitzsimons & Brain 1958   Pedioplanis namaquensis Namaqua Sand Lizard Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Baard, Villiers & Villiers 1988 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Fitzsimons & Brain 1958   Pedioplanis pulchella Venus Sand Lizard Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998  Family Scincidae (Skinks) Genus Acontias (Greater Legless Skinks)   Acontias gariepensis Gariep Blind Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Broadley 1968 Lamb, Biswas & Bauer 2010   Acontias kgalagadi Kgalagadi Legless Skink Bates 1991 [as Typhlosaurus lineatus] Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 [as Typhlosaurus lineatus] Branch 1998 [as Typhlosaurus l. lineatus] Broadley 1968 [as Typhlosaurus lineatus] Conradie & Bourquin 2013 Fitzsimons & Brain 1958 [as Typhlosaurus lineatus] Lamb, Biswas & Bauer 2010   Acontias lineatus Striped Dwarf Legless Skink Bates 1991 Bates 1996 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Broadley & Greer 1969 Lamb, Biswas & Bauer 2010   Acontias litoralis Coastal Legless Skink Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Broadley & Greer 1969 Lamb, Biswas & Bauer 2010 Mashinini 2010   * Acontias namaquensis Namaqua Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch & Maritz 2010 Branch 1981 Branch 1998 Broadley & Greer 1969 Lamb, Biswas & Bauer 2010   Acontias occidentalis Western Black Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 [as g. gracilicauda] Branch 1981 [as g. gracilicauda] Broadley & Greer 1969 [as g. gracilicauda]   Acontias tristis Northern Cape Striped Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Broadley & Greer 1969 Lamb, Biswas & Bauer 2010 Genus Mochlus (African Supple Skinks)   Mochlus sundevallii Common Savanna Supple Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Genus Panaspis (African Snake-eyed Skinks)   Panaspis wahlbergii Savanna Snake-eyed Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Medina, Bauer, Branch, Schmitz, Conradie, Nagy, Hibbitts, Ernst, Portik, Nielsen, Colston, Kusamba, Behangana, Roedel & Greenbaum 2016 Genus Scelotes (Dwarf Burrowing Skinks)   Scelotes caffer Cape Three-toed Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1989   Scelotes capensis Namibian Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998   Scelotes sexlineatus Striped Dwarf Burrowing Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Heideman, Mulcahy, Sites, Hendricks & Daniels 2011 Visser 1979 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis capensis Cape Skink Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Broadley 2000   Trachylepis homalocephala Red-sided Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Portik, Bauer & Jackman 2010   Trachylepis occidentalis Western Three-striped Skink Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Baard, Villiers & Villiers 1988 Fitzsimons & Brain 1958   Trachylepis punctatissima Montane Speckled Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981   Trachylepis punctulata Eastern Variegated Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Broadley 1975 Broadley 2000 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Portik & Bauer 2012   Trachylepis sparsa Karasburg Tree Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1981 Branch 1998 Broadley 1977 Broadley 2000 Baard, Villiers & Villiers 1988   Trachylepis spilogaster Namibian Tree Skink Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Broadley 1977 Broadley 2000 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011   Trachylepis sulcata Koppie Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Broadley 2000 Baard, Villiers & Villiers 1988 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Portik, Bauer & Jackman 2010 Portik, Bauer & Jackman 2011   Trachylepis varia Savanna Variable Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Fitzsimons & Brain 1958   Trachylepis variegata Western Variegated Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Broadley 1975 Baard, Villiers & Villiers 1988 Portik & Bauer 2012 Portik, Bauer & Jackman 2010 Genus Typhlosaurus (Blind Legless Skinks)   * Typhlosaurus lomiae Northern Cape Blind Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Huyssteen & Petford 2021   Typhlosaurus meyeri Sperrgebiet Blind Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1981 Branch 1998 Broadley 1968   * Typhlosaurus vermis Pink Blind Legless Skink Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bates, Heideman, Wilson, Hendricks, Don & Moses 1998 Branch 1981 Branch 1998  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus albigularis White-throated Monitor Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bayless, Switak & Patterson 2002 Branch 1981 Branch 1989 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Switak 2007   Varanus niloticus Nile Monitor Bauer & Branch 2001 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Colubridae (Typical Snakes or Colubrids) Genus Crotaphopeltis (Herald Snakes)   Crotaphopeltis hotamboeia Common Herald Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Visser 1979 Genus Dasypeltis (African Egg Eaters)   Dasypeltis scabra Common African Egg Eater Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Broadley 1983 Genus Dispholidus (Boomslangs)   Dispholidus viridis Green Boomslang Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 [as D. typus] Genus Philothamnus (African Green Snakes)   Philothamnus semivariegatus Variegated Bush Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983 Haacke 1985 Loehr 2005 McLachlan 1984 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus beetzii Namib Tiger Snake Bates 1991 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1989 Broadley 1983 Visser 1979 Visser 1981   Telescopus semiannulatus Southern Tiger Snake Bates 1996 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Broadley 1983 Fitzsimons & Brain 1958  Family Elapidae [part 1: terrestrial species] (Cobras, Mambas, etc.) Genus Aspidelaps (Shield-nosed Cobras)   Aspidelaps lubricus African Coral Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Broadley 1983 Dobiey & Vogel 2007 Schultz & Massyn 2008 Visser 1979 Wallach, Williams & Boundy 2014   Aspidelaps scutatus Shield-nosed Coral Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Genus Elapsoidea (African Garter Snakes)   Elapsoidea sundevallii Southern African Garter Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1971 Broadley 1983 Fitzsimons & Brain 1958 Golay, Smith, Broadley, Dixon, McCarthy, Rage, Schatti & Toriba 1993 Genus Hemachatus (Ring-necked Spitting Cobra)   Hemachatus haemachates Ring-necked Spitting Cobra Dobiey & Vogel 2007 Genus Naja (Typical Cobras)   Naja nigricincta Western Barred Spitting Cobra Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Broadley 1983 Dobiey & Vogel 2007 Wallach, Williams & Boundy 2014 Wuster, Crookes, Ineich, Mane, Pook, Trape & Broadley 2007   Naja nivea Yellow Cobra Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Broadley & Wuster 2004 Broadley 1983 Baard, Villiers & Villiers 1988 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Dobiey & Vogel 2007 Fitzsimons & Brain 1958 Goldberg 2010  Family Lamprophiidae (House Snakes, etc.) Genus Alopecion (Spotted House Snake)   Alopecion guttatum Spotted House Snake Broadley 1983 Visser 1979 Genus Amblyodipsas (Purple-glossed Snakes and Quill-snouted Snakes)   Amblyodipsas bicolor Bicoloured Quill-snouted Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1971 Broadley 1983 Genus Atractaspis (Stiletto Snakes)   Atractaspis bibronii Southern Stiletto Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1989 Broadley 1983 Broadley 1991 Fitzsimons & Brain 1958   Atractaspis duerdeni Beaked Stiletto Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Genus Boaedon (Afro-Arabian House Snakes)   Boaedon capensis Brown House Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Broadley 1983 [as fuliginosus] Fitzsimons & Brain 1958 [as lineatus]   Boaedon mentalis Namibe House Snake Hallermann, Ceriaco, Schmitz, Ernst, Conradie, Verburgt, Marques & Bauer 2020 Genus Dipsina (Dwarf Beaked Snake)   Dipsina multimaculata Dwarf Beaked Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1989 Broadley 1983 Fitzsimons & Brain 1958 Genus Homoroselaps (African Harlequin Snakes)   Homoroselaps lacteus Spotted Harlequin Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983 Genus Lamprophis (SubSaharan House Snakes)   Lamprophis aurora Aurora House Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983   Lamprophis fiskii Cape House Snake Barts, Schneider, Boone, Marais, Branch & Haacke 2012 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Marais 1981 Genus Lycodonomorphus (Southern African Water Snakes)   Lycodonomorphus rufulus Common Brown Water Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Genus Lycophidion (African Wolf Snakes)   Lycophidion capense Cape Wolf Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1996 Genus Prosymna (Shovel-snout Snakes)   Prosymna bivittata Two-striped Shovel-snout Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Broadley 1980 Broadley 1983   Prosymna frontalis Southwestern Shovel-snout Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Broadley 1980 Broadley 1983 Visser 1981   Prosymna sundevallii Southern African Shovel-snout Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1980 Broadley 1983 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Visser 1981 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis crucifer Cross-marked Grass Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1989 Branch 1998 Broadley 1983 Broadley 2002 Fitzsimons & Brain 1958   Psammophis namibensis Namib Sand Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Broadley 1983 Broadley 2002   Psammophis notostictus Karoo Sand Snake Bates 2010 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Broadley 1983 Baard, Villiers & Villiers 1988 Fitzsimons & Brain 1958   Psammophis trigrammus Western Sand Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Broadley 1983 Broadley 2002   Psammophis trinasalis Kalahari Sand Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1977 Broadley 1983 Broadley 2002 Baard, Villiers & Villiers 1988 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Fitzsimons & Brain 1958 Genus Psammophylax (Skaapstekers)   Psammophylax rhombeatus Spotted Skaapsteker Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Broadley 1976 Broadley 1983 Keates, Conradie, Greenbaum & Edwards 2019   Psammophylax tritaeniatus Southern Striped Skaapsteker Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley 1983 Genus Pseudaspis (African Mole Snake)   Pseudaspis cana African Mole Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Broadley 1983  Family Leptotyphlopidae (Thread Snakes) Genus Leptotyphlops (Typical Thread Snakes)   Leptotyphlops scutifrons Zambezi Thread Snake Adalsteinsson, Branch, Trape, Vitt & Hedges 2009 Broadley & Broadley 1999 Broadley 1983 Fitzsimons & Brain 1958 Genus Namibiana (Southwest African Thread Snakes)   Namibiana gracilior Slender Thread Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Broadley & Broadley 1999 Broadley 1983   Namibiana occidentalis Western Namaqua Thread Snake Adalsteinsson, Branch, Trape, Vitt & Hedges 2009 Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Broadley & Broadley 1999 Broadley 1983 McDiarmid, Campbell & Toure 1999  Family Pythonidae (Pythons) Genus Python (Typical Pythons)   Python natalensis Southern Rock Python Branch 1998  Family Typhlopidae (Typical Blind Snakes) Genus Afrotyphlops (African Giant Blind Snakes)   Afrotyphlops bibronii Southern African Blind Snake Broadley & Wallach 2009 Broadley 1983 Genus Rhinotyphlops (African Beaked Blind Snakes)   Rhinotyphlops lalandei Cape Blind Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Broadley & Wallach 2009 Broadley 1983 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011   Rhinotyphlops schinzi Black-spotted Beaked Blind Snake Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1989 Branch 1998 Broadley & Wallach 2009 Broadley 1983 Fitzsimons & Brain 1958  Family Viperidae (Vipers) Genus Bitis (African Adders)   Bitis arietans Common Puff Adder Bauer & Branch 2001 Branch 1998 Broadley 1983 Fitzsimons & Brain 1958 Girard 1999   Bitis caudalis Horned Adder Bauer & Branch 2001 Broadley 1983 Dobiey & Vogel 2007 Fitzsimons & Brain 1958 McDiarmid, Campbell & Toure 1999 Visser 1979   Bitis cornuta Many-horned Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Branch 1999 Broadley 1983 Dobiey & Vogel 2007   Bitis rubida South African Red Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983 [as inornata]   Bitis schneideri Namaqua Dwarf Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Broadley 1983 Dobiey & Vogel 2007 Maritz & Alexander 2011   Bitis xeropaga Desert Mountain Adder Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Broadley 1983 Dobiey & Vogel 2007 McDiarmid, Campbell & Toure 1999

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Pelomedusidae (African Side-necked Turtles) Genus Pelomedusa (African Helmeted Turtles)   Pelomedusa galeata South African Helmeted Turtle Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 [as subrufa] Petzold, Vargas-Ramirez, Kehlmaier, Vamberger, Branch, Preez, Hofmeyr, Meyer, Schleicher, Siroky & Fritz 2014  Family Testudinidae (Tortoises) Genus Chersina (Angulate Tortoise)   Chersina angulata Angulate Tortoise Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Hofmeyr 2009 Genus Chersobius (Five-toed Dwarf Tortoises)   Chersobius boulengeri Karroo Padloper Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998   Chersobius signatus Speckled Padloper Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Boycott 1986 Branch 1998 Daniels, Hofmeyr, Henen & Baard 2010 Genus Homopus (Four-toed Dwarf Tortoises)   Homopus areolatus Common Padloper Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Fritz & Havas 2006   Homopus femoralis Greater Padloper Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Branch 1998 Genus Psammobates (South African Star Tortoises)   Psammobates oculiferus African Serrated Star Tortoise Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011 Wermuth & Mertens 1977   Psammobates tentorius African Tent Tortoise Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Bauer & Branch 2001 Branch 1998 Baard, Villiers & Villiers 1988 Fritz & Havas 2006 Genus Stigmochelys (Leopard Tortoise)   Stigmochelys pardalis Leopard Tortoise Bates, Branch, Bauer, Burger, Marais, Alexander & Villiers [eds.] 2014 Conradie, Doucette-Riise, Vanhooydonck, Engelbrecht, Measey & Tolley 2011