Reptiles of United Arab Emirates    Bibliography of the reptiles of United Arab Emirates Taxonomic history of the reptiles of United Arab Emirates Introduced species

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Citation: Midtgaard, Rune. RepFocus - A Survey of the Reptiles of the World. www.repfocus.dk (Compilation date: 14th April, 2023).

Checklist General remarks: The list includes only formally described species. Newly discovered, but still unnamed species are excluded. However, unnamed species or species unassigned to species level are included, if they are the only representatives of their genus in the region. Species counts exclude extinct and introduced species. Doubtful records, typically older voucher specimens of uncertain origin, have been excluded if considered questionable, as are records based on sources considered unreliable or controversial. Note: Your comments to the checklist (omissions, errors, etc.) are welcomed, preferably if they are provided with reliable references. If possible, please attach a pdf file of the relevant reference to an email addressed to:

	Worm Lizards (Amphisbaenia)
	= endemicEX = extinct or assumed extinct Species References  Family Trogonophidae (Short-headed Worm Lizards) Genus Diplometopon (Arabian Speckled Worm Lizard)   Diplometopon zarudnyi Arabian Speckled Worm Lizard Arnold 1986 Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Leviton, Anderson, Adler & Minton 1992 Soorae, Al-Quarqaz & Gardner 2010

	Lizards (Sauria)
	= endemicEX = extinct or assumed extinct Species References  Family Agamidae (Agamas and Dragons) Genus Phrynocephalus (Toad-headed Agamas)   Phrynocephalus longicaudatus Long-tailed Toad-headed Agama Arnold 1984 [as maculatus] Baha el Din (in Osborne) 1996 [as maculatus] Drew, Al-Dhaheri, Barcelo & Tourenq 2005 [as maculatus] Gardner 2013 [as maculatus] Soorae, Al-Quarqaz & Gardner 2010 [as maculatus]   Phrynocephalus macropeltis Dhahran Toad-headed Agama Arnold 1984 [as arabicus] Arnold 1986 [as arabicus] Baha el Din (in Osborne) 1996 [as arabicus] Drew, Al-Dhaheri, Barcelo & Tourenq 2005 [as arabicus] Gardner 2013 [as arabicus] Leviton & Anderson 1967 [as arabicus] Melnikov, Melnikova, Nazarov, Rajabizadeh, Al-Johany, Amr & Ananjeva 2014 Genus Pseudotrapelus (Afro-Arabian Desert Agamas)   Pseudotrapelus jensvindumi East Arabian Desert Agama Arnold & Gallagher 1977 [as sinaitus] Arnold 1984 [as sinaitus] Baha el Din (in Osborne) 1996 [as sinaitus] Drew, Al-Dhaheri, Barcelo & Tourenq 2005 [as sinaitus] Gardner 2013 [as sinaitus] Soorae, Al-Quarqaz & Gardner 2010 [as sinaitus] Tamar, Scholz, Crochet, Geniez, Meiri, Schmitz, Wilms & Carranza 2016 Genus Trapelus (Afro-Asian Ground Agamas)   Trapelus flavimaculatus Yellow-spotted Agama Arnold 1984 Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Rastegar-Pouyani 1998 Soorae, Al-Quarqaz & Gardner 2010 Genus Uromastyx (Western Spiny-tailed Agamas)   Uromastyx aegyptia Egyptian Spiny-tailed Agama Arnold 1984 Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Soorae, Al-Quarqaz & Gardner 2010 Wilms & Boehme 2000 Wilms & Boehme 2007 Wilms 2001 Wilms, Boehme, Wagner, Lutzmann & Schmitz 2009  Family Gekkonidae (Typical Geckos) Genus Bunopus (South West Asian Tuberculated Geckos)   Bunopus tuberculatus Southern Tuberculated Gecko Arnold 1984 Baha el Din (in Osborne) 1996 Bourquin 2011 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Soorae, Al-Quarqaz & Gardner 2010 Szczerbak & Golubev 1996 Genus Cyrtopodion (Palearctic Bent-toed Geckos)   Cyrtopodion scabrum Keeled Rock Gecko Arnold 1984 Baha el Din (in Osborne) 1996 Bourquin 2011 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Soorae, Al-Quarqaz & Gardner 2010 Szczerbak & Golubev 1996 Genus Hemidactylus (Half-toed Geckos)   Hemidactylus flaviviridis Yellow-bellied House Gecko Arnold 1984 Baha el Din (in Osborne) 1996 Bourquin 2011 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Lajmi, Bansal, Giri & Karanth 2019 Lajmi, Giri & Karanth 2016 Soorae, Al-Quarqaz & Gardner 2010   Hemidactylus persicus Persian Half-toed Gecko Bourquin 2011 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2009 Gardner 2013 Soorae, Al-Quarqaz & Gardner 2010   Hemidactylus robustus Red Sea Half-toed Gecko Arnold & Gallagher 1977 [as turcicus] Arnold 1984 [as parkeri] Arnold 1986 [as turcicus] Baha el Din (in Osborne) 1996 [as turcicus] Bourquin 2011 Carranza & Arnold 2012 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 [as turcicus] Gardner 2013 Soorae, Al-Quarqaz & Gardner 2010 Genus Pseudoceramodactylus (Gulf Sand Gecko)   Pseudoceramodactylus khobarensis Gulf Sand Gecko Arnold 1980 Arnold 1984 Arnold 1986 Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Sindaco & Jeremchenko 2008 Soorae, Al-Quarqaz & Gardner 2010 Genus Stenodactylus (Afro-Asian Thin-toed Geckos)   Stenodactylus doriae Middle Eastern Thin-toed Gecko Arnold 1980 Arnold 1984 Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Soorae, Al-Quarqaz & Gardner 2010   Stenodactylus leptocosymbotes Eastern Arabian Thin-toed Gecko Arnold 1980 Arnold 1984 Arnold 1986 Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Leviton, Anderson, Adler & Minton 1992 Sindaco & Jeremchenko 2008 Soorae, Al-Quarqaz & Gardner 2010   Stenodactylus slevini Dharan Thin-toed Gecko Arnold 1980 Arnold 1984 Arnold 1986 Baha el Din (in Osborne) 1996 Bourquin 2011 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Soorae, Al-Quarqaz & Gardner 2010 Genus Trachydactylus (Arabian Tuberculated Geckos)   Trachydactylus hajarensis Hajar Tuberculated Gecko Arnold & Gallagher 1977 [as Bunopus spatalurus] Arnold 1984 [as Bunopus spatalurus] Drew, Al-Dhaheri, Barcelo & Tourenq 2005 [as Bunopus spatalurus] Gardner 2013 Leviton, Anderson, Adler & Minton 1992 Pous, Machado, Metallinou, Cervenka, Kratochvil, Paschou, Mazuch, Smid, Simo-Riudalbas, Sanuy & Carranza 2016 Soorae, Al-Quarqaz & Gardner 2010 Genus Trigonodactylus (Triangle-scaled Geckos)   Trigonodactylus arabicus Arabian Triangle-scaled Gecko Arnold 1980 Arnold 1984 Arnold 1986 Baha el Din (in Osborne) 1996 Bourquin 2011 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Leviton, Anderson, Adler & Minton 1992 Metallinou & Carranza 2013 Nazarov, Melnikov, Radjabizadeh & Poyarkov 2018 Sindaco & Jeremchenko 2008 Soorae, Al-Quarqaz & Gardner 2010  Family Lacertidae (Typical Lizards) Genus Acanthodactylus (Fringe-toed Lizards)   Acanthodactylus blanfordii Makran Fringe-toed Lizard Gardner 2005 Gardner 2013 Soorae, Al-Quarqaz & Gardner 2010   Acanthodactylus boskianus Afro-Arabian Fringe-toed Lizard Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Salvador 1982 Soorae, Al-Quarqaz & Gardner 2010   Acanthodactylus gongrorhynchatus Saudi Fringe-toed Lizard Arnold 1986 Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Leviton, Anderson, Adler & Minton 1992 Salvador 1982 Sindaco & Jeremchenko 2008 Soorae, Al-Quarqaz & Gardner 2010   Acanthodactylus haasi Eastern Saudi Fringe-toed Lizard Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Hellyer & Brown 1992 Leviton & Anderson 1967 Sindaco & Jeremchenko 2008 Soorae, Al-Quarqaz & Gardner 2010   Acanthodactylus opheodurus Snake-tailed Fringe-toed Lizard Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Hellyer & Brown 1992 Sindaco & Jeremchenko 2008 Soorae, Al-Quarqaz & Gardner 2010 Tiedemann 1991   Acanthodactylus schmidti Middle East Fringe-toed Lizard Arnold 1984 Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Leviton, Anderson, Adler & Minton 1992 Salvador 1982 Soorae, Al-Quarqaz & Gardner 2010 Genus Mesalina (Afro-Asian Desert Lizards)   Mesalina adramitana Hadramaut Desert Lizard Arnold 1984 Arnold 1986 Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Leviton, Anderson, Adler & Minton 1992 Sindaco & Jeremchenko 2008 Soorae, Al-Quarqaz & Gardner 2010 Yousefkhani, Marin & Rastegar-Pouyani 2015   Mesalina brevirostris Short-nosed Desert Lizard Arnold 1984 Arnold 1986 Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton, Anderson, Adler & Minton 1992 Smid, Moravec, Gvozdik, Stundl, Frynta, Lymberakis, Kapli, Wilms, Schmitz, Shobrak, Yousefkhani, Rastegar-Pouyani, Castilla, Els & Mayer 2017 Soorae, Al-Quarqaz & Gardner 2010 Genus Omanosaura (Oman Lizards)   Omanosaura cyanura Arabian Blue-tailed Lizard Arnold & Gallagher 1977 Arnold 1972 Arnold 1986 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Sindaco & Jeremchenko 2008 Soorae, Al-Quarqaz & Gardner 2010   Omanosaura jayakari Eastern Arabian Lizard Arnold & Gallagher 1977 Arnold 1972 Arnold 1986 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Sindaco & Jeremchenko 2008 Soorae, Al-Quarqaz & Gardner 2010  Family Phyllodactylidae (Leaf-toed Geckos) Genus Asaccus (Middle East Leaf-toed Geckos)   * Asaccus caudivolvolus Emirati Leaf-toed Gecko Arnold & Gardner 1994 Carranza, Simo-Riudalbas, Jayasinghe, Wilms & Els 2016 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Sindaco & Jeremchenko 2008 Soorae, Al-Quarqaz & Gardner 2010   Asaccus gallagheri Masafi Leaf-toed Gecko Arnold & Gardner 1994 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Pierson 2014 Sindaco & Jeremchenko 2008 Soorae, Al-Quarqaz & Gardner 2010   Asaccus gardneri Musandam Leaf-toed Gecko Carranza, Simo-Riudalbas, Jayasinghe, Wilms & Els 2016   Asaccus margaritae Wadi al Helo Leaf-toed Gecko Carranza, Simo-Riudalbas, Jayasinghe, Wilms & Els 2016 Genus Ptyodactylus (Fan-toed Geckos)   Ptyodactylus orlovi Eastern Arabian Fan-toed Gecko Els, Carranza, Gardner & Soorae 2019   Ptyodactylus ruusaljibalicus Ruus al Jibal Fan-toed Gecko Simo-Riudalbas, Metallinou, Pous, Els, Jayasinghe, Pentek-Zakar, Wilms, Al-Saadi & Carranza 2017  Family Scincidae (Skinks) Genus Ablepharus (Eurasian Snake-eyed Skinks)   Ablepharus pannonicus Southwest Asian Snake-eyed Skink Arnold & Gallagher 1977 Arnold 1972 Arnold 1986 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Grossmann, Kowalski & Schillert 2013 Soorae, Al-Quarqaz & Gardner 2010 Genus Chalcides (Cylindrical Skinks)   Chalcides ocellatus Mediterranean Ocellated Skink Arnold 1984 Arnold 1986 Baha el Din (in Osborne) 1996 Bourquin 2011 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton, Anderson, Adler & Minton 1992 Soorae, Al-Quarqaz & Gardner 2010 Genus Heremites (Western Palearctic Grass Skinks)   Heremites septemtaeniatus Seven-striped Grass Skink Gardner 2013 Moravec, Franzen & Boehme 2006 Soorae & Al Hameiri 2005 Soorae, Al-Quarqaz & Gardner 2010 Genus Scincus (Sand Fish)   Scincus conirostris Persian Sand Fish Arnold & Leviton 1977 Arnold 1984 Arnold 1986 Bourquin 2011 Carranza, Arnold, Geniez, Roca & Mateo 2008 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton, Anderson, Adler & Minton 1992 Smid, Uvizl, Shobrak, Salim, AlGethami, Algethami, Alanazi, Alsubaie, Busais & Carranza 2021 Soorae, Al-Quarqaz & Gardner 2010   Scincus mitranus Eastern Arabian Sand Fish Arnold & Leviton 1977 Arnold 1984 Arnold 1986 Baha el Din (in Osborne) 1996 Carranza, Arnold, Geniez, Roca & Mateo 2008 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Sindaco & Jeremchenko 2008 Smid, Uvizl, Shobrak, Salim, AlGethami, Algethami, Alanazi, Alsubaie, Busais & Carranza 2021 Soorae, Al-Quarqaz & Gardner 2010 Genus Trachylepis (African-American Grass and Forest Skinks)   Trachylepis tessellata Tessellated Skink Arnold & Gallagher 1977 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Sindaco & Jeremchenko 2008 Soorae, Al-Quarqaz & Gardner 2010  Family Sphaerodactylidae (Least Geckos, etc.) Genus Pristurus (Semaphore Geckos)   Pristurus carteri Masirah Semaphore Gecko Gardner 2009 Gardner 2013 Soorae, Quarqaz & Gardner 2010   Pristurus celerrimus Bar-tailed Semaphore Gecko Arnold & Gallagher 1977 Arnold 1984 Arnold 1993 Drew, Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Soorae, Quarqaz & Gardner 2010   Pristurus minimus Lesser Semaphore gecko Arnold 2009 Baha el Din (in Osborne) 1996 Drew, Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Hellyer & Brown 1992 Sindaco & Jeremchenko 2008 Soorae, Quarqaz & Gardner 2010 Tamar, Mitsi, Simo-Riudalbas, Tejero-Cicuendez, Al-Sariri & Carranza 2019   Pristurus rupestris Rock Semaphore Gecko Arnold & Gallagher 1977 Arnold 1984 Arnold 1993 Arnold 2009 Baha el Din (in Osborne) 1996 Bourquin 2011 Drew, Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Soorae, Quarqaz & Gardner 2010 Genus Teratoscincus (Plate-tailed Geckos)   Teratoscincus keyserlingii Southwestern Plate-tailed Gecko Arnold 1984 [as scincus] Arnold 1986 [as scincus] Baha el Din (in Osborne) 1996 [as scincus] Drew, Dhaheri, Barcelo & Tourenq 2005 [as scincus] Gardner 2013 Leviton, Anderson, Adler & Minton 1992 [as scincus] Macey, Ananjeva, Wang & Papenfuss 1997 [as scincus] Sindaco & Jeremchenko 2008 Soorae, Quarqaz & Gardner 2010 Szczerbak & Golubev 1996 Zwanzig & Grossmann 2015  Family Varanidae (Monitor Lizards) Genus Varanus (Monitor Lizards)   Varanus griseus Desert Monitor Arnold 1984 Baha el Din (in Osborne) 1996 Drew, Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Soorae, Quarqaz & Gardner 2010

	Snakes (Serpentes)
	= endemicEX = extinct or assumed extinct Species References  Family Boidae (Typical Boas) Genus Eryx (Sand Boas)   Eryx jayakari Arabian Sand Boa Baha el Din (in Osborne) 1996 Bourquin 2011 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Gasperetti 1988 Leviton & Anderson 1967 Soorae, Al-Quarqaz & Gardner 2010  Family Colubridae (Typical Snakes or Colubrids) Genus Lytorhynchus (Awl-headed Snakes)   Lytorhynchus diadema Crowned Awl-headed Snake Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Leviton & Anderson 1967 Soorae, Al-Quarqaz & Gardner 2010 Genus Platyceps (Flat-headed Whip Snakes)   Platyceps karelini Spotted Desert Whip Snake Baha el Din (in Osborne) 1996 [as ventromaculatus] Bourquin 2011 [as ventromaculatus] Brown 1991 [as Coluber ventromaculatus] Drew, Al-Dhaheri, Barcelo & Tourenq 2005 [as ventromaculatus] Gardner 2013 [as ventromaculatus] Schatti 2006 [as "cf. ventromaculatus"] Schatti, Kucharzewski, Masroor & Rastegar-Pouyani 2012 Soorae, Al-Quarqaz & Gardner 2010 [as ventromaculatus] Tiedemann 1991 [as ventromaculatus]   Platyceps rhodorachis Wadi Whip Snake Arnold & Gallagher 1977 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Gasperetti 1988 Sindaco, Venchi & Grieco 2013 Soorae, Al-Quarqaz & Gardner 2010 Genus Spalerosophis (Diadem Snakes)   Spalerosophis diadema Common Diadem Snake Arnold & Gallagher 1977 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Soorae, Al-Quarqaz & Gardner 2010 Genus Telescopus (Afro-Asian Cat Snakes)   Telescopus dhara Arabian Cat Snake Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Els, Al-Johany, Amr, Baha-el-Din, Disi, Boehme, Geniez & Werner 2021 Gardner 2009 Gardner 2013 Gardner, Tovey & Els 2009 Soorae, Al-Quarqaz & Gardner 2010  Family Elapidae [part 2: marine species] (Sea Snakes) Genus Hydrophis (Typical Sea Snakes)   Hydrophis curtus Short Sea Snake Baldwin & Gardner 2005 Gardner 2013 Soorae, Al-Quarqaz & Gardner 2010 Soorae, Das & Al-Mazrouei 2006   Hydrophis cyanocinctus Blue-banded Sea Snake Baldwin & Gardner 2005 Gardner 2013 Gasperetti 1988 Leviton & Anderson 1967 Soorae, Al-Quarqaz & Gardner 2010   Hydrophis lapemoides Persian Gulf Sea Snake Baldwin & Gardner 2005 Gardner 2013 Gasperetti 1988 Rasmussen 1993 Soorae, Al-Quarqaz & Gardner 2010 Soorae, Das & Al-Mazrouei 2006   Hydrophis ornatus Reef Sea Snake Baldwin & Gardner 2005 Gardner 2013 Soorae, Al-Quarqaz & Gardner 2010   Hydrophis platurus Yellow-bellied Sea Snake Baldwin & Gardner 2005 David & Ineich 1999 Gardner 2013 Gasperetti 1988 Leviton & Anderson 1967 Soorae, Al-Quarqaz & Gardner 2010 Soorae, Das & Al-Mazrouei 2006   Hydrophis schistosus Beaked Sea Snake Baldwin & Gardner 2005 Gardner 2013 Soorae, Al-Quarqaz & Gardner 2010   Hydrophis spiralis Narrow-banded Sea Snake Baldwin & Gardner 2005 Gardner 2013 Soorae, Al-Quarqaz & Gardner 2010   Hydrophis viperinus Viperine Sea Snake Baldwin & Gardner 2005 Gardner 2013 Gasperetti 1988 Soorae, Al-Quarqaz & Gardner 2010 Genus Microcephalophis (Small-headed Sea Snakes)   Microcephalophis gracilis Graceful Small-headed Sea Snake Baldwin & Gardner 2005 Gardner 2013 Soorae, Al-Quarqaz & Gardner 2010  Family Lamprophiidae (House Snakes, etc.) Genus Malpolon (Montpellier Snakes)   Malpolon moilensis Moila Snake Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Gasperetti 1988 Leviton & Anderson 1967 Soorae, Al-Quarqaz & Gardner 2010 Genus Psammophis (Old World Sand Snakes and Grass Snakes)   Psammophis schokari Arabian Sand Snake Baha el Din (in Osborne) 1996 Bourquin 2011 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Gasperetti 1988 Leviton & Anderson 1967 Marx 1988 Soorae, Al-Quarqaz & Gardner 2010  Family Leptotyphlopidae (Thread Snakes) Genus Myriopholis (Many-scaled Thread Snakes)   Myriopholis macrorhyncha Large-snouted Thread Snake Baha el Din (in Osborne) 1996 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Gasperetti 1988 Hahn 1978 Soorae, Al-Quarqaz & Gardner 2010  Family Viperidae (Vipers) Genus Cerastes (Horned Vipers)   Cerastes gasperettii Middle Eastern Horned Viper Baha el Din (in Osborne) 1996 David & Vogel 2010 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Gasperetti 1988 Leviton & Anderson 1967 McDiarmid, Campbell & Toure 1999 Soorae, Al-Quarqaz & Gardner 2010 Werner, Verdier, Rosenman & Sivan 1991 Genus Echis (Carpet Vipers)   Echis carinatus Common Carpet Viper Baha el Din (in Osborne) 1996 David & Vogel 2010 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Gasperetti 1988 Leviton, Anderson, Adler & Minton 1992 Pook, Joger, Stumpel & Wuster 2009 Sindaco, Venchi & Grieco 2013 Soorae, Al-Quarqaz & Gardner 2010   Echis omanensis Oman Carpet Viper Babocsay 2004 David & Vogel 2010 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2004 Gardner 2013 Pook, Joger, Stumpel & Wuster 2009 Sindaco, Venchi & Grieco 2013 Soorae, Al-Quarqaz & Gardner 2010 Genus Pseudocerastes (False Horned Vipers)   Pseudocerastes persicus Persian Horned Viper Cunningham 2002 Drew, Al-Dhaheri, Barcelo & Tourenq 2005 Gardner 2013 Soorae, Al-Quarqaz & Gardner 2010 Thomas 2019

	Turtles (Testudines)
	= endemicEX = extinct or assumed extinct Species References  Family Cheloniidae (Hard-shelled Sea Turtles) Genus Caretta (Loggerhead Sea Turtle)   Caretta caretta Loggerhead Sea Turtle Baldwin & Gardner 2005 Gardner 2013 Gasperetti, Stimson, Miller, Ross & Gasperetti 1993 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Soorae, Al-Quarqaz & Gardner 2010 Genus Chelonia (Green Sea Turtle)   Chelonia mydas Green Sea Turtle Al-Suweidi, Wilson, Healy & Vanneyre 2012 Baldwin & Gardner 2005 Gardner 2013 Gasperetti, Stimson, Miller, Ross & Gasperetti 1993 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Soorae, Al-Quarqaz & Gardner 2010 Genus Eretmochelys (Hawksbill Sea Turtle)   Eretmochelys imbricata Hawksbill Sea Turtle Baldwin & Gardner 2005 Gardner 2013 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Soorae, Al-Quarqaz & Gardner 2010 Genus Lepidochelys (Ridley Sea Turtles)   Lepidochelys olivacea Olive Ridley Sea Turtle Gardner 2013 King & Burke 1989 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Soorae, Al-Quarqaz & Gardner 2010 Yaghmour & Rodriguez-Zarate 2021  Family Dermochelyidae (Leatherback Sea Turtle) Genus Dermochelys (Leatherback Sea Turtle)   Dermochelys coriacea Leatherback Sea Turtle Baldwin & Gardner 2005 Gardner 2013 Gasperetti, Stimson, Miller, Ross & Gasperetti 1993 Rhodin, Iverson, Bour, Fritz, Georges, Shaffer & Dijk 2021 Soorae, Al-Quarqaz & Gardner 2010